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FLORA 
MALESIANA 



UNDER THE AUSPICES OF 

THE BOTANIC GARDENS OF INDONESIA, BOGOR, 

AND THE RIJKSHERBARIUM, LEYDEN 



ERICACEAE (H. Sleumer, Leyden) 

Trees or shrubs, very rarely herbs or fleshy saprophytes. Leaves spiral, some- 
times opposite or pseudowhorled, simple, entire, crenate or serrate, mostly ever- 
green and i coriaceous (Malesia), exstipulate (stipule-like perulae of axillary 
buds occur in Diplycosia and Vacciniump.p.). Flowers bisexual (rarely functionally 
unisexual; or the plant dioecious in extra-Mai.), characteristically regular, (4-)5 
(rarely 6-7)-merous. Inflorescences terminal or axillary, entirely covered by perulae 
in bud, mostly in racemes, these sometimes arranged to panicles or condensed 
to umbels, or reduced to few-flowered fascicles, or even to a solitary flower. Sepals 
(reduced in Monotropastrum and Wirtgenia) very rarely free, generally connate 
below to a calyx tube, the latter free or ± adnate to the ovary, persistent, whether 
or not accrescent in fruit, lobes imbricate or open in bud. Corolla campanulate 
to funnel-shaped, urceolate or cylindric, sometimes slightly zygomorphous, 
caducous, lobed to various degree, lobes imbricate (sometimes 4; contorted), 
rarely valvate in bud. Stamens usually 10 (rarely 5, 8, or up to 20), obdiploste- 
monous. rarely haplostemonous, inserted at the outer margin of the disk between 
its lobes, or sHghtly attached to the base of the corolla; filaments free (Malesia); 
anthers dorsifixed to almost basifixed, the 2 cells (thecae) not rarely extending 
into free or connate tubules, these muticous or sometimes (bi)aristate distally by 
the prolonged back-wall, opening by terminal or introrse, very rarely extrorse 
pores or slits, not rarely with projecting dorsal appendages or spurs; pollen in 
tetrads, simple in Monotropoideae. Gynoecium syncarpous, 5- or pseudo-10-, 
rarely 2-4- or 6-7-celled. Disk hypogynous or epigynous, often fleshy and nectar- 
iferous, entire or mostly 5-10-lobed. Ovary 1, superior, half-inferior or inferior, 
generally with as many cells as carpels; placentation central, with 1 or 2 lamellas 
per cell, each bearing mostly numerous, rarely 1, anatropous or obliquely amphi- 
tropous, 1-tegumented ovulus. Style 1; stigma obtuse, capitate or peltate, whether 
or not 5-7-lobed. Fruit a 5(-7)-valved, septicidal or (sometimes lately or irregularly) 
loculicidal capsule, which may be ± included by the accrescent, i fleshy calyx, 
or a rather dry to fleshy berry (Malesia). Seeds usually numerous, small, whether 
or not winged or tailed at one or both ends; testa thin, often reticulate; embryo 
cylindric, small, with copious endosperm. 

Distribution. About 125 genera with approximately 3500 spp., predominantly woody, 
distributed all over the world. 

In Malesia 12 genera, with a total of about 740 spp. Five genera are endemic or nearly so, 
viz Wirii^enia and Pernettyopsis in the Malay Peninsula, Costera in Sumatra, Borneo, the 
Philippines, Talaud, and Morotai, Dimorphani/iera in the Philippines, Moluccas, New Guinea 
and adjacent islands, and Diplycosia which occurs throughout Malesia with 1 sp. in S. Annam. 

Two other genera, viz Monotropastrum and Agapetes, find their main area of distribution in 
SE. Asia and Malesia, the latter extending to Queensland, New Caledonia, and Fiji. 

Of the remaining genera, two are restricted to the northern hemisphere but have one station 
in northwest Malesia, viz Pyrula (Europe, Asia, N. and Central America) which occurs in 
N. Sumatra, and Lyonia (SE. Asia, N. and Central America) which occurs in the Malay Penin- 
sula. 

The other three are still wider distributed. The colossal genus Rhododendron is most 
abundantly developed in Asia, with some dozens of species in N. America and Europe, and 
extends towards the southeast as far as N. Queensland (1 sp.), the Solomon Is. and New Britain 
(3 spp.), with many species in Malesia. 



470 Flora Malesiana [ser. I, vol. 6* 

Vaccinium occurs in both temperate and tropical America, in East Africa (Madagascar 
inclusive), the Pacific, and all over Asia and Malesia, but not in Australia. 

The area of Gaultheria extends over the Americas, SE. Asia, Malesia, SE. Australia (incl. 
Tasmania), and New Zealand. 

The non-endemic Ericaceae found in Malesia show a close alliance with those of SE. Asia, 
and the same can be said of the few Ericaceae found in N. Queensland, the Solomon Is., the 
New Hebrides, Samoa, and Fiji. Australian-antarctic relations within the Malesian Ericaceae 
are probably expressed only by Gaultheria mundiila (eastern part of New Guinea), which seems 
to be nearest to some species of the genus found in SE. Australia (incl. Tasmania), New Zealand, 
and possibly in temperate S. America. 

More than half of the Malesian species occur in New Guinea and are endemic there, two 
excepted. 

Ecology. Malesian Ericaceae occur predominantly in the lower and upper montane forest, 
both terrestrial and epiphytic. Quite a number is found above the timber line, mainly dwarf shurbs 
in subalpine grassland or even in true alpine vegetation, ascending in Borneo (Mt Kinabalu) 
and in the Main Range of New Guinea to 4000-4400 m. A few live at low altitudes, as some 
species of Vaccinium on coastal sands, species of Costera, or some species of Rhododendron 
on rocks near the sea or even as epiphytes on mangrove trees. 

Two saprophytic genera, viz Monotropastrum and Wirtgenia, are echlorophyllose, humicolous 
herbs, growing in shady places in the lower montane forest, apparently very rare (or overlooked 
due to their ephemeral appearance?), and only known from a few places in Sumatra and the 
Malay Peninsula. 

As to climate, Ericaceae in general prefer the wet montane to subalpine subtropical and 
tropical, and the (mostly rather dry) warm-temperate zone; they extend into the arctic and 
subantarctic zone. They are predominantly found on acid, i.e. sandy, humous or peaty soil. 
A few are reported from limestone, but it is not clear in how far they grow there on humus- 
covered soil. For example, Stresemann found dense brushwood Rhododendron under Casuarina 
on steep limestone ridges in the Ora Mts in W. Ceram, together with Podocarpus, Myrtaceae, 
and Nepenthes, May 1911. Brass found in New Guinea several Rhododendrons on limestone rock. 

Malesian Ericaceae are mostly of proportionally small size, but some may under favourable 
conditions grow into trees up to c. 15 m or are high lianas (Dimorphanthera). By their helio- 
philous preference they occur most abundant as epiphytes in the forest and are not infrequently 
gregarious or co-dominant terrestrially on steep slopes, volcanic terrain, ridge-crests, glades and 
other sterile places in the mountains as 'cricoid brushwood'. 

In Sumatra Frey-Wyssling found on the volcano Kerintji above the forest border at 3000 m 
a brushwood consisting largely of Gleichenia and Ericaceae ( Vaccinium miquelii. Rhododendron 
retusum and Gaultheria leucocarpa) (Trop. Natuur 22, 1933, 1-10). From the Gajo Lands 
(Atjeh) VAN Steenis reported the important role of Ericaceae in the brushwood and glades of 
the high non-volcanic mountains (Tijd. Kon. Ned. Aardr. Gen. 55, 1938, 751, f. 12). On the 
volcano Dempo in Bencoolen Forbes found Vaccinium miquelii dominating the zone above 
2500 m where it may be a tree with a diameter of 40 cm (Wanderings, 1885, 209). 

On the Javanese volcanoes, especially on the wetter western peaks, Vaccinium varingiaefolium 
and Rhododendron retusum are gregarious and essential co-dominants in the brushwood of 
slopes, crests and peaks, and are even obvious from a far distance when the new shoots expose 
their pink or red flush. Other co-dominants of this gnarled forest, either mossy or not, are 
Rapanea, Myrica javanica, and Leptospermum flavescens. 

On the mountains of the Malay Peninsula and Borneo Ericaceae are common, but they 
attain their greatest and most profuse development in North Borneo on Mt Kinabalu where 
again they are important co-dominants in the upper zones. 

From the Philippines, Celebes, and the Moluccas no special great importance as constituents 
of vegetation are known, but in the New Guinean mountains they are as gregarious at high 
altitude as on Mt Kinabalu, e.g. on Mt Arfak. From the Balim Valley Brass reported that 
Rhododendron brassii composed nearly pure, greyish stands best developed on forest edges 
and in glades between low-tree thickets, seldom extending far beyond the territory of conifers 
(J. Am. Arb. 22, 1941, 321). He reported from the same area, at the mouth of the Bele Valley, 
at 2100 m, a distinctive shrub community c. 3-4 m high, of Vaccinium acrobracteatum on a.patch 
of loose sand, associated with Gahnia, Myrtaceae, Rhododendron inconspicuum and R. herzogii. 



Jan. 1966] Ericaceae (Sleumer) 471 

On Mt Wilhelmina he found Vaccinium dominans in dense, 5-6 m high, often wind-clipped 
forest, co-dominant with Rapanea from 3500-3800 m {I.e. 327). On Mt Albert Edward on the 
Wharton Range, from 3200 m upwards, forests were dominated by Podocarpus papuanus and 
Libocedrus papiiana, and at higher altitude by a Vaccinium with the conifers rising in open stand 
above the canopy; this forest type was considerably interrupted by grassy glades, hollows and 
lake basins {I.e. 336). The Rev. Cruttwell found Rhododendron christianae locally forming 
pure stands well recognizable at flowering time from the air. 

By their heliophile ecology and their ability to stand sterile, acid, or poor soil conditions 
Ericaceae are not rarely found as pioneers or in secondary vegetation in open or fired places, 
on landslides, in clearings made by man following burning, logging, gold digging, or making 
roads, on volcanic sands and rock, on lapilli, near craters, on lavastreams, and rockwalls. 

In almost every crater or solfatara field in Sumatra, Java, and the Lesser Sunda Is. one can 
observe that in approaching such terrain the percentage of the Ericaceous component increases, 
and they are among the lot which comes closest to the sulphur vents, notably Gaultherias, 
Rhododendron javanicum, R. retusum, Vaccinium laurifolium, and V. varingiaefoUum, often 
associated with Dianella javanica, Gahnia javanica, Pleopeltis feci, and some Carex. The ad- 
vanced pioneers often attain queerly clipped or prostrate shape, and wind forms (fig. 101-104) 
as described by van Steenis from Mt Papandajan (Trop. Natuur 24, 1935, 141-145, f. 2-5) 
and from Mt Idjen {ibid. 29, 1940, 182, f. 9). On the solfatara fields of Samosir I. in Lake Toba, 
N. Sumatra, Rhododendrons are conspicuous. Vaccinium varingiaefoUum is also able to with- 
stand a partial burial by acid volcanic mud and lapilli, as observed on summit of Mt Tjeremai 
above Cheribon, N. Central Java. 

To a certain extent Ericaceae may also be fire-resistant as described by Junghuhn from Mt 
Lawu, where he found a pyrogenous grassland thinly set with Vaccinium varingiaefoUum. On 
the west slope of the Latimodjong Range, SW. Central Celebes, near Rante Lemo, the large- 
flowered, Z-A m high Rhododendron vanvuurenii forms pure stands in pyrogenous grassland, at 
c. 1000 m; also near Rante Pao. Lane-Poole reported Vaccinium striicaule to be able to stand fire 
in the grasslands of Mt Saru waged, associated with tree ferns and Rhododendron; this Vaccinium 
originally grows below Dacrydium and Libocedrus, and after these have been burned it stays in 
the open grassland between 3300 and 4000 m (For. Res. Papua, 1925, 179-180). 

The root-system of Malesian Ericaceae is, particularly in crater fields and near solfataras, 
very horizontal, and roots may attain large dimensions in such places. This is of course common 
to almost all plants growing on sterile rocky, impervious or badly drained soils; it may also be 
ascribed in part as a reaction to the heat of the deeper soil in crater fields. Such roots have also 
a distinctly corky bark, and von Faber holds that this would be a protection to heat (Die 
Kraterpflanzen Javas, 1927, 71-75, f. 7-9). 

The stem-base and upper part of the root of Vaccinium is often swollen and superficially 
resembles a lignotubcr. It is obviously a storage organ and is already found in epiphytic seedlings 
(von Faber, I.e.); such seedlings have also a proportionally very large root-system. 

Dispersal. Not much is known about the dispersal of Malesian Ericaceae besides what 
DocTERS VAN Leeuwen has observed on Mt Pangrango in W. Java. The very small seeds of 
Rhododendron are presumably dispersed by wind (see what is said sub Rhododendron below). 
The fruits of genera as Diplycosia, Gaulfheria, Vaccinium, Costera, Agapetes, and Dimor- 
phaniera are fleshy, and, so far as known, eaten by birds, rodents and small mammals. The 
seeds arc small, and besides in Rhododendron, have a hard testa; they may be disposed in 
a viable state with the excrements of animals within a certain distance. Docters van Leeuwen 
made observations on bird dispersal of berries of Vaccinium and GauUheria on Mt Pangrango 
in W. Java (Vcrh. Kon. Ak. Wet. A'dam sect. 2, 31, 1933, 198-212, and in Flora 118/119, 1925, 
86). His son, Hans, examined stomach contents of birds on the Central Javanese volcanoes 
Sumbing and Sindoro and had similar results, the chief birds being species of the genera Zostera 
and Merula (Treubia 10, 1929, 443, 446). 

Galls. No Zoocecidia specific for a distinct genus or species arc hitherto known from Male- 
sian Ericaceae {cf Doc-iers van Lj-.euwi.n, The Zoocecidia of the Netherlands East Indies, 
1926. 439, and Pangrango, 1933. 198 .seq.). 

Anomalous flowers, due to a fungus or insects, are not rarely found in Vaccinium (see there). 

On the leaves of both Vaccinium and Rhododendron spots occur due to the parasitic Exobasi- 



472 Flora Malesiana [ser. I, vol. 6* 

dium, described by Raciborski. In Java this does not lead to the formation of galls; however, 
very large Exobasidium galls are found on Rhododendron atjehense in N. Sumatra. Fig. 48. 

The roots of Ericaceae possess endotrophic mycorrhiza which, according to von Faber, 
are spread by seed; they cause gall-like thickened malformations on the roots (Die Krater- 
pflanzen Javas, 1927, 71-75, f. 7-9). 

Wood anatomy. Den Berger, Determinatietabel houtsoorten van Malesie, Veenman, 
Wageningen (1949) 31, 35 (hand lens). Metcalfe & Chalk, Anat. Die. 1 (1950) 823, 831; 
Moll & Janssonius 4 (1920) 269; Record, Trop. Woods 32 (1932) 11. Primitive features 
present: conspicuous many-barred scalariform perforation plates in Rhododendron and in 
some species of Vaccinium, pits to ray cells opposite, ground tissue composed of fibre tracheids, 
wood parenchyma practically absent. Record I.e.: a relationship to the Theaceae seems to be 
strongly indicated. - C.A.R.-G. 

Phytochemistry. Ericaceae were rather intensively investigated since Plugge showed in 
1882 and 1883 that many members contain a toxic constituent, called andromedotoxin by him. 
From the chemical point of view the family may be characterized as follows at present: 

(i) Polyphenolic substances and tannins are accumulated by all members investigated. 
Leucoanthocyanins, flavonols (including derivatives of myricetin), catechins and a number of 
phenolic /3-glucosides like arbutin {Arbutus, Arctostaphylos, Pyrola, Vaccinium), monotropitoside 
{i.e. the former gaultherin, a glucoside of methyl salicylate; in Gaultheria and Monotropa), 
betuloside (= rhododendrin; in Rhododendron) and lyonoside (in Lyonia) represent the poly- 
phenolic group of plant constituents. The tannins are galli- and ellagitannins in some members 
and condensed tannins in others; many species contain mixtures of both types of tannins. 

(ii) Triterpenes, in many instances predominantly free ursolic acid, are deposited in large 
amounts in the very often thick cuticule of the leaves. 

(iii) The well known toxic constituents of the family are diterpenes. Andromedotoxin (= 
rhodotoxin = grayanotoxin-I), desacetylandromedotoxin (= grayanotoxin-III) and desacetyl- 
anhydroandromedotoxin (= grayanotoxin-II) are the best known representatives of these 
principles. They have been demonstrated to be present in members of the genera Kalmia, 
Rhododendron, Andromeda, Chamaedaphne, Leucothoe, Lyonia, Pernettya and Pieris, but have 
not been detected in Pyroleae and in Vaccinioideae. 

The tendency to accumulate still other constituents seems to be more restricted in the family. 
Many Ericaceae bear, besides ordinary trichomes, abundantly glandular hairs. The latter seem 
to produce often essential oils in quantity. For species of Cassiope, Ledum and Rhododendron 
investigations of the essential oils were performed. The herbaceous members, reunited by many 
authors in the separate family Pyrolaceae, produce besides arbutin a number of closely related 
glucosides (homoarbutin, pyrolatin, renifolin). From the latter the naphtoquinone chimaphilin 
probably arises after enzymatic splitting of the glucoside and oxydation of the aglykon, process- 
es which take place when the plants are dried. Another y?-glucoside present in all Pyrolaceae 
(including Monotropa) is monotropeoside, which belongs to the so-called pseudoindicans or 
iridoid plant constituents. It seems highly probably that some members of arborescent Ericaceae 
{e.g. Arbutus) contain such compounds too. 

In concluding we may state that chemistry agrees very well with the inclusion of Pyrolaceae 
in Ericaceae. With regard to possible relationships of Ericaceae nothing of convincing nature 
can be said. In many respects chemistry of Ericaceae reminds of amentiferous families {e.g. 
Betulaceae, Fagaceae) on the one hand and Saxifragaceae, Hamamelidaceae, and Rosaceae 
on the other hand. In the light of present-day knowledge biochemical resemblance between 
Ericaceae and Theaceae seems to be less striking than the resemblance between Ericaceae and, 
for instance, Saxifragaceae sensu lato. For references I refer to my 'Chemotaxonomie der 
Pflanzen' vol. 4 (1965). — R. Hegnauer. 

Uses. Apart from ornamental purpose but few useful plants are found among Malesian 
Ericaceae. The hard timber, due to the small size, is only occasionally used for implements. 
Fruits are reported to be partly sweet or at least palatable in all genera of the subfamily Vacci- 
nioideae, but no regular use by men is known except of Vaccinium myrtoides in the Philippines. 
The leaves of several species of Gaultheria have a strong smell and contain wintergreen-oil 
(methyl-salicilate); they are used externally against rheumatism, or internally as medicinal tea. 



Jan. 1966] Ericaceae (Sleumer) 473 

In New Guinea the big, scented flowers of Rlwdodendron and the ones of Dimorphanthera 
are locally used for decoration. 

Wild Rhododendron species from Malesia are cultivated in Europe, N. America, Hawaii, 
and Australia; their hardiness, even of "alpine" species, in temperate zones is questionable. 
Seeds or cuttings of wild species of Rhododendron for horticultural purposes should be taken and 
airmailed: the pods should be almost ripe, not yet open, or just apically opening. 

Taxonomy. The family Ericaceae is taken here in a wide sense, including Pyrolaceae, 
Monotropaceae, and Vacciniaceae, which are kept as proper families by various authors, though 
their close connection within the Ericales is generally adniitted. Of the 5 subfamilies the Rhodo- 
dendroideae, Arbutoideae, Monotropoideae, and Vaccinioideae are represented in Malesia; the 
Ericoideae are confined to Europe and Africa. 

No precursory work on Malesian Ericaceae as a whole has been done. J. J. Smith devoted 
many papers to Malesian Ericaceae. Local revisions have been made by J. J. Smith and Miss 
J. Amshoff for Java, by H. F. Copeland for the Philippines, by J. J. Smith and R. Schlechter 
for parts of New Guinea, and the genera Agapetes, Dimorphanthera, Diplycosia, Gaultheria, and 
Vaccinium for the whole of New Guinea were previously treated by the author. 

In connection with the present work precursory revisions and/or supplements for the whole 
of Malesia (partly including the adjacent regions, as far as this seemed to be necessary) have 
been published by the author on Agapetes (Nova Guinea, Bot. n. 1, 1960, 1-7; Blumea 12, 1964, 
344), Costera (Blumea 11, 1961, 3-4), Dimorphanthera (Nova Guinea, Bot. n. 7, 1961, 73-102 
Blumea 12, 1963, 137-142; ibid. 12, 1964, 344-345), Diplycosia (Reinwardtia 4, 1957, 119-161 
Blumea 11, 1961, 4-8; ibid. 12, 1963, 116-123; ibid. 12, 1964, 342-343), Gaultheria (Reinwardtia 
4, 1957, 163-188; Blumea 11, 1961, 1-3; ibid. 12, 1963, 114-116; ibid. 12, 1964, 342), Modo- 
dendron (Reinwardtia 5, 1960, 45-231; Blumea 11, 1961, 113-130; ibid. 12, 1963, 90-114; 
ibid. 12, 1964, 339-341), and Vaccinium (Blumea 11, 1961, 9-112; ibid. 12, 1963, 123-137; 
ibid. 12, 1964, 343-344). 

The treatment given here is tentative in many respects, especially for Borneo and New Guinea, 
which islands are hardly more than superficially known in many parts of their mountains which 
are so rich in Ericaceae. Up till now every larger recent collection from these islands has yielded 
new species and localities, mainly in the genera Rlwdodendron and Vaccinium. Therefore, the 
range of even the more common Bomean and New Guinean Ericaceae is still imperfectly 
known in detail. 

Generic delimitation. In general there is no difficulty in the delimitation of the genera in 
Malesia, but in the Vaccinioideae generic distinction is unclear. Though Costera and Dimor- 
phanthera are distinct 'Sippen' by their discrete area of distribution, the characters by which they 
are distinguished from Vaccinium are of no great importance within the subfamily. The distinc- 
tion between Agapetes and Vaccinium is even less distinct, and without a clear geographical 
background, it is merely based on convenience. As a closer insight in and reconsideration of 
generic limits would involve a complete revision of the subfamily I have refrained from taking 
premature steps on a local basis. 

Phytography. The descriptions of most species treated in this work are based on herbarium 
specimens; the size of fleshy organs, as corollas and soft fruits is therefore given in the dry 
state; ± 20 % (sometimes up to 40%) must be added to get the size in the fresh state. The author 
has studied Ericaceae in the field during a short stay in the Philippines (Luzon: Mountain 
Province), in the Malay Peninsula (Cameron Highlands) and in North Borneo (Mt Kinabalu), 
and a prolonged one in New Guinea (Vogelkop Peninsula, Cycloop Mts, Eastern Highlands, 
Bululo-Wau area) which enabled him to take measurements of such fleshy organs of numerous 
species on the spot; these have been added to the descriptions for the use of field collectors. 

Notes for collectors. In general both flowers and fruit are necessary for determining the genus; 
fruiting material alone is generally insufficient for this purpose, and in Vaccinioideae it is as 
worthless as sterile material. 

For identification of species within the genera the presence of flowers is a conditio sine qua 
non. Collectors should never collect sterile material in this family. 



474 Flora Malesiana [ser. I, vol. 6^ 

KEY TO THE GENERA 

1. Ovary superior or nearly so, i.e. the calyx free at anthesis (if accrescent only seemingly connate with 
the fruit). Fruit mostly a capsule, rarely berry-like or a true berry. 

2. Fruit a septicidal capsule (Rhododendroideae-Rhododendreae) 1. Rhododendron 

2. Fruit a loculicidal capsule or a berry. 

3. Echlorophyllose, saprophytic, humiculous herbs. Leaves scale-like, i imbricate along the fleshy 

stem. Below the petals a whorl of leaves, here called 'sepals'. Roots fleshy. Pollen simple (Mono- 

tropoideae). 

4. 'Sepals' 5. Anthers horseshoe-shaped, the cells linear, confluent only at the apex. Ovary fusiform, 

gradually narrowed to the rather long cylindric style 2. Wirtgenia 

4. 'Sepals' 0-2. Anthers subreniform, the cells subglobular, connected lengthwise. Ovary ovoid 
or pear-shaped, narrowed to a very short and thick style 3. Monotropastrum 

3. Shrubs or trees with green leaves. Pollen in tetrads (Arbutoideae). 

5. Calyx small and remaining so, not accrescent nor becoming succulent in fruit. Fruit a capsule. 
6. Low suffruticose plants with creeping stem and ~ basal leaves. Raceme solitary, terminal 

(Pyroleae) 4. Pyrola 

6. Erect shrubs or trees with many, normally lateral racemes (Andromedeae) ... 5. Lyonia 
5. Calyx large from the beginning or ± accrescent, surrounding the ripe fruit, mostly becoming 

± succulent in fruit. Fruit a capsule or a berry (Gaultherieae). 

7. Fruit a berry with scanty pulp. Calyx lobes large from the beginning, as long as or longer than 
the corolla, — spreading, neither accrescent nor becoming fleshy 6. Pemettyopsis 

7. Fruit a capsule, sometimes rather thin-walled and tardily splitting. Calyx lobes small initially 

and shorter than the corolla, ± accrescent and succulent in fruit, finally ± incl. the capsule. 
8. Anther-cells not elongate into tubules, muticous, or rarely (bijaristate by the elongate back- 
wall of each cell 7. Gaultheria 

8. Anther-cells passing into short or mostly elongate apical tubules, these connate to or free 

from each other and never aristate 8. Diplycosia 

1. Ovary inferior or nearly so, i.e. the calyx at anthesis at least partly connate with the ovary. Fruit a 

(sometimes rather dry) berry, crowned by the persistent calyx lobes and the disk (Vaccinioideae). 

9. Stamens isomorphous, equal or slighty unequal in length; tubules not expanded apically. 

10. Corolla generally small to medium-sized (0.4-1.5, rarely up to 2, occasionally {Vaccinium pilosilo- 

bum) up to 3 cm) and membranous, rarely somewhat fleshy. 

1 1 . Flowers solitary or in few-flowered fascicles which are axillary or not rarely extra-axillary. 

Pedicels rather long and filiform, not articulate with the calyx (ovary) 9. Costera 

11. Flowers in axillary racemes, sometimes reduced to a solitary flower. Pedicels mostly relatively 

short and/or stoutish, rarely slender, manifestly articulate with the calyx (ovary). 10. Vaccinium 

10. Corolla generally large and fleshy ((0.8-) 1.7-4.5 cm). (Pedicelsmostlyarticulate, rarely continuous 

with the calyx) 11. Agapetes 

9. Stamens manifestly dimorphous and very unequal in length; tubules expanded, i.e. conspicuously 
ear-shaped 12. Dimorphanthera 

1. RHODODENDRON 

LiNNE, Sp. PI. 1 (1753) 392; Sleum. Bot. Jahrb. 74 (1949) 511; Reinwardtia 5 
(1960) 47; Blumea 11 (1961) 113; Jahrb. Rhodod. Ges. Bremen (1963) 7; Blumea 
12 (1963) 90; ibid. 12 (1964) 339. — Fig. 1—53, 104. 

Evergreen or deciduous (in Mai. 1 sp.) shrubs or small trees, terrestrial or 
epiphytic, the subsequent new shoots marked by distinct nodes. Stem often thicken- 
ed at the base; roots not rarely large with soft tips in epiphytic specimens. Leaves 
petioled, rarely sessile, spirally arranged or opposite, or mostly in 3-8-merous 
pseudowhorls, ~ coriaceous, entire or crenulate, whether or not clothed with 
scales and/or hairs. Flowers in a mostly terminal, sometimes lateral, it condensed 
umbelliform raceme which is occasionally reduced to a single flower. Flower buds 
with numerous perulae varying in shape and size from the outer to the inner ones, 
early or late caducous. Bracteoles 2 at the base of each pedicel. Calyx cup- or 
disk-shaped, sometimes merely rim-like, ± deeply 5(-7)-partite to subentire, often 
oblique, persistent. Corolla tubular or campanulate to funnel-shaped, sometimes 
salver-shaped, regular or somewhat zygomorphous, 5(-7)-lobed to various degree; 



Jan. 1966] 



Ericaceae (Sleumer) 



475 



lobes imbricate in bud, erect to expanded, sometimes even horizontally spreading 
at anthesis. Stamens 5-10(-14), inserted at the base of the corolla, often unequal; 
filaments linear to filiform; anthers dorsifixed, in general introrse, very rarely 
extrorse, whether or not appendiculate at the base, opening by round terminal or 
— oblique and more elliptic sublateral pores; pollen agglutinate. Disk mostly 
thick and — prominent, 5-10(-14)-lobed. Ovar;' superior, 5(-7)-locular, placentas 
bifid, bearing numerous ovules; style mostly as long as, rarely shorter than the 
stamens; stigma capitate to turbinate, sometimes almost peltate, crenate to 
deeply lobed. Capsule dehiscent from the top, firstly by whether or not throwing 
off" a soft-celled outer stratum, then dividing its valves into a star-shaped figure, 
valves thin to very thick, straight or twisting in later stages, the central column 
persistent. Seeds very numerous, minute, whether or not bearing appendages or 
thread-like tails at both ends; testa thin, reticulate; albumen fleshy. 

D i s t r . Approximately 850 spp. in all, of which c. 525 occur in Asia (mainly in tropical and subtropical 
E. and SE. Asia), c. 25 spp. in N. America, 9 spp. in Europe, 1 sp. in Australia (N. Queensland), in 
Malesia c. 280 spp., about half of them in New Guinea incl. New Britain and the Solomon Is. (Bougain- 
ville to Malaita), not extending into the proper Pacific flora. Fig. 1. 




Fig. 1. Distribution of Rhododendron in Malesia and Australia. The number above the hyphen indicates 
the number of endemic species, that below the hyphen the number of non-endemic species in each island 

or district. 



The bulk of the Malesian species belong to the lepidotc seel. Vireya, which outside Malesia has 2 spp. 
in Indochina (/?. iriumphans YfRsrN & Cheval. and R. chevalieri Dop in S. Annam), and 1 sp. in Australia 
(R. lochae F. v. M. in NE. Queensland). Only 5 elepidotc spp. arc native in Malesia, all closely related 
to contincntui, Asia-centred groups, extending into the F'hilippincs (Luzon), the Malay Peninsula, and 
Sumatra. 

Ecol. Most species occur as terrestrial substagc shrubs or as epiphytes in the mountain forest, especi- 
ally in the moss forest, and only occasionally (and then apparently always as epiphytes) in lowland 
forest, even in the mangrove (R. hrookeanum and R. lonni/lorum). Terrestrial in general in the fringes of 
the upper forest belt and adjacent savannahs, they may grow there occasionally as epiphytes on Podo- 
carpus and tree-ferns. They become dwarfs in the alpine /one in grassland and on rocks, reaching altitudes 
of about 4<X)() m both in Borneo (Mt Kinabalu) and in New Cniinea. In most epiphytic species flowering 
seems to be rather irregular, so that flowers may be found during the whole year, other taiul predominant- 
ly the terrestrial) species have a more pronounced flowering time. 



476 Flora Malesiana [ser. 1, vol. 6* 

Dispersal. Each capsule contains very numerous light seeds, each in most of the Malesian species 
provided with a narrow wing or tail at both ends. These seeds are apparently easily carried by wind, 
especially from epiphytic specimens living in the crowns of high forest trees. The fact, that landslides, 
or artificially cleared ground (abandoned cultures, air strips), or sluiced areas of gold mines in Borneo 
and New Guinea are often quickly grown with Rhododendron species from the neighbouring forests, 
points to wind as the main factor in natural dispersal. 

Ridley (Dispersal, 1930, 127) thinks, that these seeds are attached by their tails to the branches 
of the trees on which they grow, but there is no proof of that. 

Pollination. The only observations on pollination in Malesian Rhododendron refer to R. retusum 
from Sumatra and Java. According to Docters van Leeuwen "the anthers in the adult bud lie close 
under the hood formed by the corolla lobes; they are all open, and the pollen hangs in threads from the 
anthers. In this stage the stigma lies at a distance of about 2 mm below the anthers, and is not yet recep- 
tive. The flowers are therefore distinctly protandrous. Not much is changed on the first flowering day; 
during the second and the third flowering-day the style elongates, and as the now receptive stigma 
brushes past the anthers, it will come in contact with the pollen-masses hanging down from the latter. 
Finally the stigma projects 1 or 2 mm beyond the corolla". R. retusum has been seen visited bij a honey- 
bird (Aethopyga eximea) and bumble-bees (Docters van Leeuwen, Verh. Kon. Ak. Wet. A'dam sect. 2, 
31, 1933, 204). 

During his stay in New Guinea (Kubor Range, Western Highlands), Mr W. Vink has observed many 
times at 3600 m altitude that Rhododendron flowers were visited by the sooty honey sucker bird Melidectes 
fuscus fuscus (de Vis), taking honey from the red, tubular corollas. 

The salver-shaped, partly sweet-scented flowers of the species belonging to subsect. Siphonovireya 
and subsect. Solenovireya suggest, that Sphingidae might play a role in visiting and pollinating flowers 
of these groups at night. 

Hybridisation. From experiments with several Malesian species in the past century, it is known 
that not only crossing between two species, but also multiple crossing between hybrids is rather easily 
obtained. It can therefore be expected, that hybridisation occurs under natural conditions. Several 
presumable hybrids have been recognized in herbarium materials, of which one parent must have been 
R. malayanum of the subsect. Malayovireya which is so distinct in the peculiar type of its scales from all 
other subsections of the genus in Malesia. In other cases it was R. jasminiflorum of the subsect. Soleno- 
vireya with its outstanding salver-shaped corolla, which has entered as one parent in hybrids with species 
of other subsections. Less or almost not recognizable in dry material are hybrids between species of the 
same subsection or even of the same series, which, however, might be disclosed on the spot among the 
parental species by a combination of minor characters as colour of the leaves or flowers, which are nor- 
mally lost in the drying process, or from the habit, which cannot be judged from plants cut into small 
herbarium samples. It is for that reason, that in this treatment probably several natural hybrids have 
been described from dried material as proper species, which in future might be recognized as local 
hybrids in the field. 

During my field work in the Arfak Mts I have observed both single specimens and small populations 
of apparently hybrid origin among or not far from the presumed parents, such as 248. R. laetum (corolla 
pure yellow) x 56. R. konori (corolla white to pinkish), 255. R. zoelleri (corolla tube yellow, lobes orange) 
X 57. R. phaeopeplum (corolla pure white), 56. R. konori (corolla large, white to pinkish) x 62. R. 
asperum (corolla much smaller, white or generally pink), 62. R. asperum (the pink form) x 248. R. laetum 
(corolla yellow), and 268. R. culminicolum var. angiense (corolla pinkish red x 68. R. phaeochristum 
(corolla deep red to carmine). These hybrids were striking at first glance by the intermediate colour of 
their flowers, white x yellow or orange resulting in pale yellow or pink corollas, pink x yellow in salmon 
shades of various degree. The pods found on such hybrids were intermediate in size and density of in- 
dument, and regularly full of seeds. Hybrids between species oi subsect. Euvireya (268. R. culminicolum 



Fig. 3. Main types of scales on the undersurface of leaves and used in the key to the subsections of 
Rhododendron sect. Vireya, all X 20. a. Rather small sessile and moderately stellate-incised scales on a 
dry leaf of R. inconspicuum. b. Rather large subsessile (subdendroid) and more deeply stellate scales on 
a fresh leaf of R. dianthosmum. c. Stalked (dendroid) scales on a fresh leaf of R. leucogigas. d. Tomentum 
(felt) of densely set dendroid and deeply stellate-lacerate scales on a fresh leaf of R. beyerinckianum. 
e. Dendroid and deeply stellate scales, each scale on top of an epidermal tubercle, on leaf margin of a 
fresh leaf of R. bryophilum. f. Epidermal tubercles (papillae) on old dry leaf of R. konori, the scales 
already gone. g. Scales typical for the species of subsect. Malayovireya on a dry leaf of R. malayanum. 
h. Disk-shaped and sessile scales on a dry leaf of /?. retusum. i-i'. Disk-shaped and sessile scales on upper 
surface (/) and undersurface (/') of a dry leaf of /?. gaultheriifolium, also showing the crenulation of the 
leaf margin by impressed scales (a Sleumer &. Vink 4261, b-e cultivated at Proefstation voor de Boom- 
kwekerij (Experim. Station for the cultivation of trees and shrubs) at Boskoop, Holland, / Koster- 
MANS 2248, g Meijer s.n. from Mt Sago, Sumatra, h Van Ooststroom 13310, /-/' Sleumer 4156). 



Jan. 1966] 



Ericaceae (Sleumer) 



477 





1^ 










<>«»>. 

^y.: 



478 



Flora Malesiana 



[ser. I, vol. 6* 



var. angiense, 248. R. laetum, 255. R. zoelleri) with rather scattered flattish scales, with such of subsect. 
Phaeovireya (62. R. asperum, 56. R. konori, 68. R. phaeochristum) with dense, dendroid scales, showed 
an intermediate type and/or density of scales on flowers and leaves. The enormous variation in size of 
corolla and fruit around the Anggi Lakes of R. konori is probably due to introgression of local R. 
asperum into the very abundant R. konori. Combination of 5-merous species (/?. asperum, R. zoelleri) 
with normally 7-merous species (/?. konori, R. phaeopeplum) resulted in specimens with 6- or 5- and 
6-merous flowers. 

Such hybrids have in New Guinea exclusively been found in burned or otherwise devastated places 
within the surrounding Castanopsis-Quercus-Nothofagus forest, where the parental species grow rather 
scattered and often as epiphytes under natural conditions. Obviously species of the forest invade such open 
places covered by regrowths oi Pteridium, Gleichenia, grasses, sedges, Lycopodium, etc., they may become 
terrestrial, multiply considerably, can meet each other and thus get a better chance for crossing and 
back-crossing. 

Vern. Tjantigi, J, S; New Guinea: warade, Kapauku, tschop-schai, Manikiong, krobua, Kebar, rohn, 
Mendi. 

Uses. Although the wood is hard, no real use is known as apparently the stems are to small in dia- 
meter in general. The flowers are occasionally used for decoration purposes by the natives. 

Cultivation. The genus plays an important role in horticulture. In the first place, the most hardy 
species from temperate regions have been cultivated outdoors, but more and more less hardy and more 
showy ones have been introduced from the tropical and subtropical mountains of SE. Asia into those 
parts of Europe and America, where the winter is not too strong. W. Malesian Rhododendrons such as 
R. longiflorum, R. javanicum, R. malayanum, and R. jasminiflorum have been cultivated in European 
glasshouses since a hundred years (c/. Henslow, J. R. Hort. Soc. 13, 1891, 240-283). Apparently they 
were disappointing to breeders and amateurs and are today confined to Botanic Gardens. Holttum 
reported on his experience with the cultivation of R. longiflorum, R. jasminiflorum, and various hybrids 
at Singapore (M.A.H.A. Mag. 9, 1939, 34-37; ibid. 11, 1941, 93-95, with phot.). Seeds germinated and 
seedlings grew best on finely broken brick mixed with fern-roots (Asplenium nidus) or on chopped 
(tree-)fern-roots only, probably due to the favourable aeration in such a substratum. The first flowering 
began at c. 2\ years from seed. 

A new interest started when the first species of the upper montane part of New Guinea, R. beyerinck- 
ianum, with its deep blood-red flowers, came into bloom from a seven years old seedling in the Edinburgh 
Botanical Garden in 1956. Through the Rijksherbarium at Leyden, seeds of about 40 spp. of New 
Guinean Rhododendron have been distributed to various nurseries (Boskoop, Seattle) and Botanical 
Gardens (Edinburgh, Kew, San Francisco) since 1955, several of which (/?. macgregoriae, R. wrightianum, 
R. leptanthum, R. zoelleri) have already come into flower. Among others I introduced the big-flowered 
238. R. leucogigas from the Cycloop Mts in northern New Guinea in 1961 by cuttings, one of which 
has flowered in 1964 at Boskoop, cf. Schneider, Jahrb. Deutsch. Rhodod. Ges. Bremen (1965) 29-33, 
f . 1 (/?. zoelleri), f. 2-4 (/?. leucogigas). R. zoelleri has been pictured on a stamp of the former Dutch 
New Guinea. 




Fig. 2. Main types of scales on the undersurface of leaves and used in the key to the subsections of 
Rhododendron sect. Vireya, all X 70. a. Scale variously lobed, with dark chestnut coloured centre and 
of two different sizes, exclusively found in subsect. Malayovireya (R. malayanum). b. Scale deeply stellate- 
incised or -lacerate and stalked ("dendroid") to various degree (R. konori); on top of an epidermal 
tubercle exclusively found in subsect. Phaeovireya. c. Scale moderately substellately angled or dented 
and sessile {R. javanicum). d. Scale entire or almost so and sessile, exclusively found in subsect. Pseudo- 
vireya {R. quadrasianum var. cuneifolium). (a Meijer s.n., Mt Sago, Sumatra, b. Kostermans 2248, 
c JuNGHUHN s.n., Java, d Sinclair 9084). 



Jan. 1966] Ericaceae (Sleumer) 479 

Notes. When not stated otherwise all measurements given in the descriptions are taken from dry 
specimens; to get the full life size of fleshy organs as corollas, fruits, etc., generally ± 20 % should be 
added. The colours mentioned are those taken from the collectors labels or observed by me in the field. 

For the determination of the lepidote species the detail examination of the various types of scales 
on which the diff"erent subsections are mainly based, and the use of a lens are indispensable. See fig. 2-3. 

KEY TO THE SUBGENERA, SECTIONS AND SUBSECTIONS 

1. Plants lepidote, i.e. covered with scales (= epidermal trichomes) at least on the undersurface of the 
young leaves, mostly also on the tips of the branchlets and in the floral region. Seeds manifestly 

appendaged by - long tails or wings at both ends 1. Subg. Rhododendron sect. Vireya 

2. Scales "disk-shaped', i.e. their marginal zone entire or almost so (at most slightly-regularly or mostly 

irregularly-crenulate) and narrow in relation to the generally darker, often thick or swollen centre. 

(Scales on the branchlets and/or the pedicels sometimes on top of thick persistent, epidermal, 

wart-like protuberances, never so on the blades). — Fig. 2d. 

3. Corolla generally short-tubular, rarely funnel-shaped or (tubular-)campanulate; lobes erect or 

spreading. Spp. 1-25 1. Subsect. Pseudovireya 

3. Corolla trumpet-like or salver-shaped, tube elongate and ± narrow, straight or slightly curved; 
lobes equalling J or less of the total length of the corolla, spreading ~ horizontally. Spp. 26-32. 

2. Subsect. Siphonovireya 
2. Scales 'star-shaped', i.e. their marginal zone distinctly — regularly or irregularly — lobed, dented, 
incised or lacerate to various degree on the branchlets and the foliage (sometimes less deeply dented 
or even subentire and more disk-like on the corolla and/or the ovary), narrow to wide in relation 
to the mostly darker, generally but slightly or not thickened centre. Centre either flat to slightly 
deepened (scale 'sessile'), or distinctly protracted downwards into a shorter or longer, slender 
foot, stem or column (scale 'dendroid', i.e. stalked). — Fig. 2a-c. 

4. Scales in general markedly 'dendroid', and each on top of a distinct and permanent, bluntish or 
subapiculate, tubercle; upper and/or undersurface of the leaves ir rough to the touch after the 
scales are gone. (Marginal zone of the scales usually wide and deeply, often narrowly incised; 
extreme forms similar to stellate hairs, which, however, are absent). Spp. 33-74. — Fig. 2b. 

3. Subsect. Phaeovireya 

4. Scales usually 'sessile', rarely 'subdendroid' (sometimes so in subsect. Solenovireya, rarely in 

subsect. Euvireya), and each either on top of a low and rather inconspicuous, apparently not 

permanent epidermal tubercle, or mostly on no tubercles at all; leaves quite smooth at least at full 

age on both faces. (Marginal zone of the scales narrow to wide; centre often i sunk into the 

epidermis of the leaves). 

5. Centre of the scales rather large and very dark (sooty or dark chestnut or blackish brown). 

Marginal zone of the scales relatively wide, r: deeply and broadly obtusely several-lobed, becoming 

silvery with age. Scales of 2 obviously different sizes, i.e. very many smaller and few much larger 

ones irregularly mixed, all very dense, touching or at least in part overlapping each other. (Scales 

of a similar type, but smaller, less distinctly unequal and spaced for their greater, overlapping 

for their smaller number: hybrids of the species of this subsection with species of other subsections). 

Spp. 75-88. — Fig. 2a 4. Subsect. Malayovireya 

5. Centre of the scales mostly rather small and variously coloured (though never as dark as in 

subsect. Malayovireya). Marginal zone of the scales wider or narrower, mostly but moderately 

(up to - halfway) lobed or dented. Scales all equal in size, very dense to very spaced. — Fig. 2c. 

6. Scales very dense, touching or slightly overlapping each other, generally rather large, still forming 

a coherent layer on the undersurface of r. mature leaves which persists for a rather long time 

(the pits which remain, when the proper scales have finally gone, dense to very dense, at least 

denser than in subsect. Euvireya). Spp. 89-102 5. Subsect. Albovireya 

6. Scales lax to subdense, rarely dense, always distinctly spaced on the undersurface of ' mature 

leaves, generally rather small and often early caducous (or the marginal zone early dissolute). 

7. Corolla trumpet-like or salver-shaped, the straight or somewhat curved tube elongate and 

narrow, lobes relatively short, equalling { of less of the total length of the corolla and spreading 

r horizontally. .S>/7. 103-136 6. 6'M/).swr Solcnovircya 

7. Corolla cither tubular, campanulale or funnel-shaped, tube generally wider and shortci than 
in subsect. Solenovireya, lobes equalling \ or more of the total length of the corolla, erect or 

funnel-shaped, rarely horizontally expanded. 5/jp. 137-276 1. Subsect. V.uy'ircya 

I. Plants cicpidotc. i.e. without any scales. Seeds whether or not provided with short, irregular, mostly 

one-sided appendages, but no proper tails at both ends. 
8. Flower buds (or inflorescences) strictly terminal (solitary). 
9. Flowers and leafy branchlets (new foliate shoots) from difl'ercnt buds, the first from terminal 
(strictly floral) buds, the latter from lateral buds. i.e. from the axils of the leaves of the past season. 
Hairs, if bristle-like, not flattened. 



480 Flora Malesiana [ser. I, vol. 6* 

10. Leaves persistent. 5p;j. 277-279. . . . 2. Subg. hymenanthes ^ecr. Hymenanthes 5«/65ecr. Irrorata 

10. Leaves deciduous. Sp. 280 3. Subg. Pentanthera sect. Pentanthera 

9. Flowers and leafy branchlets (new foliate shoots) from the same terminal bud, the latter from the 
axils of the lowest, scale-like peruiae of the recent season. Hairs often bristle-like and flattened. 

Leaves persistent or deciduous. Spp. 281-286 4. Subg. Tsutsutsi sect. Tsutsutsi 

8. Flower buds (or inflorescences), aUhough seemingly 'terminal', really lateral (mostly several to- 
gether), coming from the axil(s) of the uppermost leaf (or leaves). Leaves persistent. Spp. 287-288. 

5. Subg. Azaleastrum sect. Choniastrum 

1. Subgenus Rhododendron 

Rhododendron sect. Enrhododendron Endl. Gen. (1839) 159 p.p.; DC. Prod. 7 (1839) 721 p.p.; 
MiQ. Fl. Ind. Bat. 2 (1859) 1057. — Rhododendron subg. Lepidorrhodion Koehne, Deutsch. 
Dendrol. (1893) 449; Sleum. Bot. Jahrb. 74 (1949) 530. 

Plants lepidote, i.e. covered with scales (epidermal trichomes) at least on the undersurface 
of the young leaves, generally also on the branchlets and in the floral region. 

1. Section Vireya 

(Bl.) Copel./. Philip. J. Sc. 40 (1929) 136, 151; Sleum. Bot. Jahrb. 74 (1949) 536; Reinwardtia 
5 (1960) 50. — Vireya Bl. Bijdr. (1826) 854; G. Don, Gen. Syst. 3 (1834) 846; Steud. Norn. 
Bot. ed. 2, 2 (1841) 774 {'Viereyd'). — Rhododendron subg. Vireya (Bl.) Clarke in Hook./. 
Fl. Br. Ind. 3 (1882) 462; Copel./. Am. Midi. Nat. 30 (1943) 604. 
Seeds manifestly appendaged by ± long tails or wings at both ends. 

Distr. E. Asia, Australia (N. Queensland), and Malesia. 



1. Subsection Pseudovireya 

(Clarke) Sleum. Bot. Jahrb. 74 (1949) 537; Blumea Suppl. 4 (1958) 47; Reinwardtia 5 (1960) 
50. — Rhododendron subg. Pseudovireya Clarke in Hook. / Fl, Br. Ind. 3 (1882) 464. — 
Rhododendron sect. Zygomorphanthe Schltr, Bot. Jahrb. 55 (1918) \A5 p.p. — Rhododendron 
sect. Discovireya Sleum. Bot. Jahrb. 74 (1949) 539. 

Scales disk-shaped, i.e. their marginal zone entire or almost so (at most slightly — ^regularly 
or mostly irregularly — crenulate) and narrow in relation to the generally darker, often thick or 
swollen centre, on the branchlets and/or the pedicels sometimes on top of thick, persistent, 
wart-like stalks. Corolla mostly i shortly tubular, rarely funnel-shaped or (tubular-)campanu- 
late, lobes erect to spreading. 

Distr. E. Asia, Malesia. Fig. 4. 




Fig. 4. Distribution of Rhododendron subsect. Pseudovireya in Malesia and Asia. The number above the 
hyphen indicates the number of endemic species, that below the hyphen the number of non-endemic 

species in each island or district. 



Jan. 1966] Ericaceae (Sleumer) 481 

KEY TO THE SPECIES 

1. Branchlets covered at the tips with scales on stalks (r. 0.5 mm), densely verruculose in the older 
parts by the remaining stalks when the scales are gone. 

2. Calyx long-ciliate, i.e. laxly or ~ densely set with rather long whitish hairs at the margin. 
3. Leaves obovate to oblong-obovate, arranged in distinct pseudowhorls, (1.5-)2.5-4(-6.5) by 
(0.8-)1.2-1.8(-3) cm. (Umbels (2-)4-10-flowered) 1. R. retusum 

3. Leaves lanceolate or oblong-lanceolate, spirally arranged, very dense together, 6-12 by 2-3 mm. 

2. R. taxoides 

2. Calyx very laxly and shortly ciliate, or not ciliate. 

4. Flowers solitary or more rarely in twos. 

5. Corolla 1-1.3 (rarely up to 1.5) cm long in all. Leaves 4-9 mm long. 
6. Leaves entire or nearly so, the margin hardly or not thickened and but slightly revolute. 
7. Leaves elliptic or elliptic- or oblong-obovate, obtuse or subacute at apex, laxly lepidote under- 
neath. Pedicels very slender, lepidote and shortly patently hairy 3. R. pulleanum var. puUeanum 
7. Leaves broad-obovate to nearly rounded, often retuse, ± densely lepidote beneath. Pedicels 

rather stoutish, densely lepidote, very sparsely or not hairy 4. R. nummatum 

6. Leaves well visibly crenulate by impressed scales at the ± thickened margin, ovate or elliptic- 
ovate, obtusely acuminate 5. R. gaultheriifolium 

5. Corolla (1.5-)1. 7-2.4 cm long in all (when 1.5 cm long, then the leaves at least in part longer 
than 10 mm). Leaves 6-20 mm long. 
8. Leaves elliptic, or ovate- or oblong-elliptic, gradually subacuminate towards the apex, subobtuse. 

6. R. oreites 
8. Leaves narrowly to broadly obovate, or suborbicular, obtuse or retuse at apex. 
9. Leaves ± elongate-obovate, 6-9(-10) by (2-)2.5-4 (rarely in part up to 6) mm 7. R. erosipetalum 
9. Leaves obovate to broadly obovate or suborbicular, or, if oblong- or elUptic-obovate, the 
smallest at least 5 mm wide in the same specimen. 

10. Leavesobovate-suborbicular, 7-lOby 5-8mm 8. R. hameliiflorum 

10. Leaves obovate or broadly obovate, or oblong- or elliptic-obovate. 

11. Leaves 6-10 by 4-6 mm 3. R. pulleanum var. maiusculum 

11. Leaves (8-) 10-25 by (5-)6-12(-18) mm. 
12. Anthers 1-1.2(-1.5) nam long. Capsules 1-1.5(-1.8) cm long .... 9. R. lindaueanum 

12. Anthers 2 mm long. Capsules 7-8 mm long 10. R. spathulatum 

4. Umbels 3-5-flowered. 
13. Leaves minutely, though well visibly crenulate, ovate to elliptic- or oblong-ovate or elliptic, 
rarely obovate-elliptic, slightly or not revolute at the margin. 
14. Leaves 0.7-1.8 by 0.5-1.2 cm; scales very dense, partly touching each other on the under- 

surface of the leaves; nerves ± inconspicuous beneath 11. R. saruwagedicum 

14. Leaves (1.5-)2-3.5 by (1.2-)1.5-2(-2.2) cm; scales less dense, all distant from each other on 

the undersurface of the leaves; nerves ± conspicuous beneath 12. R. invasorium 

13. Leaves quite entire, obovate-spathulate or broadly oblanceolate, much revolute at the margin; 

nerves mostly distinctly prominent beneath 13. R. perakense 

1 . Branchlets covered at the tips with scales on very low stalks, or sessile, inconspicuously verruculose 
or mostly smooth when the scales are gone. 
15. Corolla distinctly funnel-shaped or campanulate. 
16. Leaves reticulate beneath. Branchlets dirty brown-velutinous at the tips . . 14. R. vanderbiltianum 
16. Leaves not reticulate beneath. Branchlets papillose-puberulous, or not hairy at all at the tips. 
17. Leaves oblong-obovate or narrowly elliptic, sometimes almost lanceolate, 0.5-1.1 cm wide. 

IS. R. seimundii 
17. Leaves elliptic, or elliptic-suborbicular, or (broadly) obovate, (1.5-)1.8-2.3(-3) cm wide. 

16. R. scortechinil 
15. Corolla tubular or nearly so, at most slightly widened towards the limb. 
18. Umbels (4-)6-12-flowercd (with longish subpatent hairs at the pedicels when 4- or 5-flowcrcd). 
19. Leaves 0.8-1.8 cm by 2-3 mm 17. R. adinophyllum 

19. Leaves 2.5-3 by 1.2-1.9 cm 18. R. cyrtophyllum 

18. Flowers solitary or in twos (very rarely up to 4 in an umbel, but then the pedicels not longish 

subpatcnt-hairy). 

20. Calyx lobes distinct, clongalc-subovatc-triangular, obtuse, : 2 mm . . .19. R. crlcoidcs 
20. Calyx lobes broadly ovatc-roundcd, obtuse, up to I mm, or generally rather inconspicuous. 

21. Leaves lanceolate, subacute at apex and base, 4-6 by 1.5-2.5 mm. 

20. R. nanophyton var. nanophyton 
21. Leaves ovate, or oblong, or elliptic, or (elongate-)obovatc, i obtuse at apex and/or ba.se. 
22. Corolla 2.5-3.2 cm long in all. 
23. Ovary densely and exclusively lepidote. 



482 



Flora Malesiana 



[ser. I, vol 6*. 



24. Leaves ovate to elliptic-ovate or elliptic, 5-8(-12) by 3-5(-8) mm . . 21. R. schizostigma 

24. Leaves obovate, 1.9-4.3 by 0.7-1.9 cm 22. R, meliphagidum 

23. Ovary densely hairy and lepidote (scales almost covered by the hairs) . . . . 23. R. vinkii 
22. Corolla 1-1.8 (rarely up to 2.5) cm long in all. 
25. Leaves obovate-oblong, 2.5-4(-6) by 1.5-2(-2.5) mm. 20. R. nanophyton var. petrophilum 
25. Leaves longer. 
26. Branchlets besides the scales bearing no hairs at all. Corolla lobes distinctly erose-denticulate 

at apex 24. R. ciliilobum 

26. Branchlets besides the scales generally short-pubescent or patent-puberulous at the tips 
(and may be on the petioles). Corolla lobes entire 25. R. quadrasianum 



1. Rhododendron retusum [Reinw. Verb. Bat. Gen. 
9 (1823) 32, nom. nud.\ (Bl.) Benn. in Benn. & Br. 
PI. Jav. Rar. (1838) 86 in text, 88, t. 20; DC. Prod. 
7 (1839) 724; Hassk. Cat. Hort. Bog. (1844) 161; 
MoR. Syst. Verz. (1846) 42; Planch. F1. Serres 5 
(1849) 477-480 h; Zoll. Syst. Verz. 2 (1854) 137; 
HouTTE, Fl. Serres 10 (1854-55) t. 1044; Hook. 
Bot. Mag. (1855) t. 4859; Lemaire, III. Hort. 2 
(1855) t. 70; Miq. Fl. Ind. Bat. 2 (1859) 1058; 
ibid. Suppl. 1 (1 860) 25 1 , 586 (J. angustifolia); Ann. 
Mus. Bot. Lugd.-Bat. 1 (1863) 44; Bl. Fl. Jav. Pl. 
ined. (1863-83) t. 7 & 7B, f. 2, A & B; Boerl. in 
Veth, Midden Sumatra 4, 2 (1884) 22; Koord. 
Junghuhn Gedenkb. (1910) 185; Exk. Fl. Java 
3 (1912) 5, f. 2; Koord. -Schum. Syst. Verz. 1 
(1912) fam. 233, p. 106; J.J.S. in K. & V. Bijdr. 13 
(1914) 110; RiDL. J. Fed. Mai. St. Mus. 8 (1917) 
58; MiLLAis, Rhodod. (1917) 235; Koord. Fl. 
Tjib. 3 (1918) 4; Hochr. Candollea 2 (1925) 491, 
incl. var. macranthum Hochr. I.e. A9T, Sp. Moore, 
J. Bot. 63 (1925) Suppl. 57; DocTERS VAN Leeuwen, 
Trop. Natuur 16 (1927) 190, f. 33; Pangrango 
(1933) 204, t. 29A; J.J.S. in Merr. Contr. Am. 
Arb. 8 (1934) 122; Steen. Trop. Natuur 24 (1935) 
143; Amsh. in Back. Bekn. Fl. Java (em. ed.) 7 
(1948) fam. 162, p. 3; Sleum. Reinwardtia 5 
(1960) 54; Steen. Endeavour 21 (1962) 189, t. 6; 
Sleum. Blumea 12 (1963) 90. — Vireya retusa Bl. 
Bijdr. (1826) 856; G. Don, Gen. Syst. 3 (1834) 
848. — Azalea retusa (Bl.) O. Ktze, Rev. Gen. 
PI. 2 (1891) 386, 387. — Fig. 104. 

KEY TO THE VARIETIES 

1 . Corolla strictly tubular, 2-2. 5 (-3) cm long in all, 

lepidote and hairy outside. 
2. Style lepidote at the base, bearing no hairs 

at all 1. var. retusum 

2. Style laxly patent-hairy at the base or in the 

lower half 2. var. trichostylum 

1. Corolla tubular-campanulate, c. 1.4 cm long in 
all, exclusively lepidote outside. 

3. var. epilosum 

1. var. retusum. 

Shrub or small tree, (0.3-)2-4(-7) m, in general 
terrestrial, erect or occasionally creeping as pio- 
neer; stem or trunk nodose, dark brown, irregular- 
ly branched; branches grey-corticated. Branchlets 
terete, younger parts red, densely clad with stalked 
roundish rather soon caducous scales, the remain- 
ing stalks in form of thickish warts, which make 
the upper 3-4 internodes brownish-blackish 
verruculose; internodes 2-6 cm. Leaves in 4-7- 



merous pseudowhoris at the upper nodes, obovate 
to oblong-obovate, apex rounded or mostly retuse, 
base cuneate into the petiole, subcoriaceous to 
coriaceous, dark green and shining above, paler 
to almost whitish green and — dull beneath, 
laxly to subdensely lepidote on both faces initially, 
early glabrescent above only (scales disk-like, 
marginal zone narrow and i entire, centre rather 
large and thick, manifestly impressed), margin ± 
distinctly recurved, (1.5-)2.5^(-6.5) by (0.8-)1.2- 
1.8 (-3) cm, midrib impressed above, strongly 
raised beneath, nerves 4-6(-8) pairs, lower ones i 
ascendent, higher ones irregularly spreading, all 
slightly impressed above in fully mature leaves 
only, otherwise but faintly visible, ± obsolete 
beneath, reticulation 0; petiole stoutish, grooved 
above, lepidote or finally rugose, 1-5 mm. Umbels 
(2-)4-10-flowered; flowers erect in the early bud 
stage, pendulous afterwards. Outer perulae subu- 
late to ovate-acuminate, inner ones gradually in- 
creasing in size and broadly ovate to spathulate, 
shortly acute, ih lepidote outside, inner ones 
sericeous in their lower part, all ± densely and 
longish white-ciliate, up to 1.3by0.8cm.Bracteoles 
linear to filiform, up to 6 mm. Pedicels rather 
slender, densely ± patently hairy (to nearly villous), 
lepidote only at the apex, (0.5-)1.2-1.5(-2) cm. 
Calyx small, oblique, shortly 5-lobed or -dentate, 
c. 4 mm 0, densely lepidote outside, lobes c. 1 mm 
long ciliate. Corolla tubular, light red to intense 
scarlet, or flame- or orange-red, 2-2.5 (rarely up to 
3) cm long in all, tube rather straight-cylindric, 
base slightly 5-saccate or dilated in form of a 
ring, 4-6 mm at the very base, dilated upwards 
to 8 mm, laxly to rather densely yellow-lepidote 
and short whitish hairy outside, glabrous inside, 
lobes erecto-patent or somewhat spreading, broad- 
ly oblong to suborbicular, 4-6 by 3-5 mm. 
Stamens 10, alternately slightly unequal, mostly — 
exserted; filaments linear, somewhat widened 
below, glabrous; anthers broad-oblong, truncate 
at the top, c. 1.5 by 1 mm. Disk glabrous. Ovary 
oblong-conical, 5-angular or -grooved, densely 
lepidote, not hairy at all, c. 4 imn, rather abruptly 
narrowed to the style, the latter filiform, lepidote 
at the base only, otherwise glabrous, accrescent 
during anthesis, finally slightly exserted; stigma 
capitate, shortly 5-lobed. Capsule erect, oblong- 
fusiform, lepidote, 1-1.5(-1.8) by 0.4-0.5 cm, on 
a hairy pedicel 1.8-2.4 cm. Seeds oblong, c. 2 mm, 
winged in two opposite directions, wings membra- 
nous, obcuneate, irregularly incised distally. 

Distr. Malesia: Sumatra, Java (especially in 
W. Java; in central Java only known from Mt 



Jan. 1966] 



Ericaceae (Sleumer) 



483 



Prahu (Dieng) and Mt Merapi; in E. Java from 
Mt Ardjuno). 

Ecol. In subalpine forests or shrubberies, in 
mossy Vaccinium forest, as well in a soil rich in 
humus (also in the moss-cushions of old fallen 
trees) as on sandy, stony or rocky grounds, often 
near craters and solfataras, there forming very 
low and small-leaved, windridden forms, the root- 
system of which extends near the surface of the 
ground, the stems able to root; locally not rare, 
(1350-) 1500-3400 m. Fl. Jan.-Dec. 

Pollination. After Docters van Leeuwen 
"the flowers are distinctly protandrous. During 
anthesis the style elongates, and the then receptive 
stigma brushes past the anthers, coming in con- 
tact with the pollen-masses, which hang down 
from the anthers in threads. In the basal annular 
part of the corolla nectar is secreted. The flowers 
are visited by small honey birds {Aethopyga) and 
bumble-bees, which apparently take part in the 
pollination". 

Vern. Bunga tjanki, Sumatra (Kerintji), 
tjantigi, J, S, tjantigi beureum, S. 

2. var. trichostyliun Sleum. Reinwardtia 5 (1 960) 55. 
Leaves 1.5-2.5 by 0.7-1.4 cm. Style laxly patent- 
hairy at the base or in the lower half. 

Distr. Malesia: N. Sumatra (Atjeh: Goh Lem- 
buh, Putjuk Angasan, Bur ni Telong), on forested 
slopes. 1350-2700 m. 

3. var. epUosum J. J. S. Bull. Jard. Bot. Btzg III, 13 

(1935) 446; Sleum. Reinwardtia 5 (1960) 55. 
Corolla tubular-campanulate, c. 1.4 cm long in 

all, besides the scales not hairy at all outside. 
Distr. Malesia: Central Sumatra (Padang High- 
lands: Ml Gombak), once found in low forest at 
2330 m. 

2, Rhododendron taxoides J. J. S. Nova Guinea 18 

(1936) 92, t. 1 8, 2; Sleum. Reinwardtia 5 (1960) 55. 
Shrub, 0.4-1.5 m. Branchlets very densely set 

with leaves in their upper and with very numerous 
scars in the lower defoliate part respectively, and 
rather densely with caducous scales on short stalks, 
which remain in form of small, rather dense warts. 
Leaves scattered, lanceolate to oblong-lanceolate, 
apex obtusely acuminate, base shortly broad- 
attenuate, obtuse, stiffly coriaceous, shining green 
above and light green with brown scales beneath, 
glabrous above at full maturity, beneath rather 
laxly Icpidote, more densely so on the midrib 
(scales sessile, rounded or nearly so, entire to 
irregularly undulate, marginal zone rather narrow, 
centre large, thick, dark and impressed), recurved 
at the whole margin, not rarely r convex in dry 
specimens, edge somewhat thickened and crenulate 
by impressed scales, 6-12 by 2-3 mm, midrib 
very slightly grooved towards the base above, 
thick and obtusely raised beneath, nerves ! 
indistinct; petiole rather thick, Icpidote, 1 .5-2 mm. 
Flowers mostly in twos, sometimes solitary. 
Pcrulae narrow-ovate to obovatc, apiculatc, outer 
ones laxly Icpidote dorsally, inner ones f dense- 
ly sericeous, ciliate, up to 6 by 3 mm. Bracteoles 



linear, pubescent, up to 5 mm. Pedicels slender, 
subdensely covered with shortly stalked scales and 
short hairs, 8-9 mm. Calyx oblique, small, disk- 
shaped, wavy, c. 3 mm o, shortly obtusely 5-lobed, 
lepidote, margin laxly set with white hairs (c. 
0.5 mm). Corolla dark red, 1.4-1.5 cm long in all, 
tubular, somewhat oblique and curved, rather 
densely lepidote outside except the upper part of 
the lobes, which bears some sparse hairs, glabrous 
inside, tube 1-1.2 cm long, c. 2.5 mm at base, 
widened up to 5 mm below the lobes, the latter ± 
erect, suborbicular to ovate, subtruncate-obtuse, 
minutely irregularly crenulate, ciliolate, c. 3 by 
2-3 mm. Stamens 10, unequal, the longest nearly 
equalling the corolla; filaments filiform, glabrous; 
anthers subelliptic, c. 1 mm. Disk glabrous. Ovary 
obliquely conical, 5-ribbed, c. 2.5 by 1.5 mm, 
densely lepidote, gradually tapering to the glabrous 
and c. 1 cm long style; stigma thick-obconical, 
lobulate. Capsule obliquely fusiform, sparsely 
lepidote, c. 9 by 3 mm. 

Distr. Malesia: W. New Guinea (Mt Door- 
man), on open slope or summit on serpentine 
rocks, 3250-3520 m. Fl. Oct. 

3. Rhododendron pulleanum Koord. Nova Guinea 
8 (1912) 879, t. 154, 4; J. J. S. ibid. 12 (1914) 130; 
ibid. 18 (1936) 90; Sleum. Reinwardtia 5 (I960) 
56. — R. minimifolium Wernh. Trans. Linn. Soc. 
II, Bot. 9 (1916)98/7./?. 

var. pulleanum. 

Terrestrial or epiphytic shrub, up to 2 m, much 
branched. Branchlets terete, covered in the upper- 
most part with stalked scales, lower parts glabres- 
cent though still densely verruculose by the remain- 
ing stalks, old parts smooth and grey-corticate. 
Leaves — scattered or subopposite, rather dense 
along the branchlets, elliptic or elliptic- or oblong- 
obovate, apex subacute or mostly obtuse, base 
— cuneate, coriaceous, deep green above, paler 
beneath, glabrous above, laxly lepidote beneath 
(scales brownish reddish, sessile, round, rather 
large, marginal zone narrow, entire or nearly so, 
centre thick, slightly or not impressed), edge ± 
revolute, or whole blade ± convex above in dry 
specimens, 4-7 (-9) by 2.5-5 mm, midrib ir. obscure 
above, slightly prominent beneath, nerves obsolete 
above, faintly visible beneath; petiole rather slen- 
der, 1-1.5 mm. Flowers solitary. Pcrulae ovate to 
oblong-ovate, subacute, unequal, up to 7 by 2-4 
mm, glabrous, whitish ciliate. Bracteoles linear 
to narrow-spathulate, very shortly hairy, 5-7 mm. 
Pedicels very slender, nearly filiform, densely 
lepidote and finely patently hairy, c. 7 mm. 
Caly.x oblique, lepidote, shortly 5-dcnted, 3 of the 
teeth short-triangular (1 mm), 2 other larger 
(1.5-3 mm), • spreading. Corolla tubular, red, 
1.3-1.5 cm in all, rather densely lepidote outside, 
glabrous inside, tube cylindric, straight. 10-12 mm 
long. 3-4 mm at base, 5-6 mm below the lobes, 
the latter erect, obovatc-oblong, 2-3 (rarely up to 
4) by 2-2.5 mm. Stamens 10, not or slightly 
exserted; filaments linear, filiform, glabrous; 
anthers oblong-obovate, truncate, c. 1 mm. Disk 



484 



Flora Malesiana 



[ser. I, vol. 6* 



glabrous. Ovary ovoid-conical, entirely covered 
with scales, c. 3 mm, rather gradually tapering 
to the style, the latter thickish, lepidote at the 
very base, glabrous otherwise, attaining c. 1 cm 
in length; stigma subclavate, moderately thickened, 
indistinctly 10-lobed. 

Distr. Malesia: W. New Guinea (Nassau Mts). 

Ecol. In mossy forest, or mostly in dense low 
scrub on exposed summits or in open dwarfish 
moss grown Vacciniiim forest, 1420-2600(-3450?) 
m. Fl. Jan.-Dec. 

var. maiusculum Sleum. Reinwardtia 5 (1960) 56; 
Blumea 11 (1961) 113. 

Leaves 6-10 by 4-6 mm. Corolla 1.7-2 cm long 
in all. Pedicels densely lepidote and entirely 
epilose, otherwise ± as var. pulleanum. 

Distr. Malesia: Central New Guinea (on open 
sandy crest of ridge at 2300 m near Lake Habbema 
and in alpine shrub vegetation on Mt Antares at 
3300-3400 m, said to be common in both places). 
Fl. July, Oct. 

Note. This variety approaches R. hameliiflorum 
in many respects. 

4. Rhododendron nummatum J. J. S. Nova Guinea 
1 8 (1 936) 9 1 , t. 1 7, 2; Sleum. Reinwardtia 5 (1 960) 
56; Blumea 11 (1961) 113; ibid. 12 (1963) 90; ibid. 
12 (1964) 339. 

Stiflf, erect, terrestrial or epiphytic shrub, 0.3-0.5 
(rarely up to 1.5) m, much and closely branched. 
Branchlets slender, terete, tips densely clad with 
stalked scales, lower parts verruculose where the 
scales have gone, finally glabrescent and dark 
greyish blackish corticate. Leaves scattered, more 
dense towards the tips, broad-obovate to nearly 
rounded, apex rounded-obtuse, not rarely slightly 
retuse, base ± broadly attenuate or rounded, 
coriaceous, somewhat convex above, initially 
rather densely lepidote on both faces, glabrous 
above at maturity (scales round, rather large, 
sessile, marginal zone entire or nearly so, centre 
thick, slightly or not impressed), margin slightly 
or not revolute, lamina mostly flat or but slightly 
convex, 4-9(-13) by 3.5-8(-9) mm, midrib promi- 
nent beneath only, otherwise ± obsolete as are 
the 2 or 3 nerves; petiole slender, 1-1.5 mm. 
Flowers pendent, solitary, or rarely in twos. Fe- 
rulae narrow-ovate, subacute to obtuse, glabrous, 
ciliolate, up to 3 mm. Bracteoles linear-spathulate, 
short hairy distally, up to 4 mm. Pedicels rather 
stoutish, densely lepidote (the scales on very short 
stalks), very sparsely or not hairy, 5-6.5 mm. 
Calyx oblique, small, irregularly 5-lobed, lepidote 
dorsally, lobes triangular, obtuse, denticulate, up 
to 0.5 mm. Corolla pink or (dark) red, whether 
or not with yellow at the base, tubular, slightly 
zygomorphous, tube cylindric, straight, sparsely 
to subdensely lepidote outside, glabrous inside, 
c. 1 by 0.35 cm, lobes erect, ovate-orbicular, 
irregularly denticulate, laxly lepidote outside in 
their lower half, 1.2-1.8 by 1.5-2 mm. Stamens 10, 
± included; filaments linear, slightly dilated to- 
wards the base, glabrous; anthers 0.8-1 mm. Disk 
glabrous. Ovary cylindric, 5-ribbed, covered with 



rusty scales, c. 3 mm, attenuate to the rather thick- 
ish style, which is laxly lepidote in its lower half 
and attains 4-5 mm in length; stigma subclavate- 
capitate, indistinctly 10-lobed. Capsule narrow- 
oblongoid, 8-13 by 2.5-3 mm, distinctly 5-ribbed. 

Distr. Malesia: New Guinea (Main Range 
from Mt Doorman and the Hellwig Mts to Mt 
Victoria, also on Mt Saruwaged). 

Ecol. In sheltered ravine, forest borders, open 
mossy bush, rest clearings in ridge crest forest, 
stunted shrubbery on ridge top or open hill side or 
open clayey ground of old gold workings, on 
rocks, often together with mosses and lichens 
when terrestrial, 2060-3250 m. Fl.fr. Jan. -Oct. 

5. Rhododendron gaultheriifolium J. J. S. Nova 
Guinea 18 (1936) 90, t. 17, 1; Sleum. Reinwardtia 
5 (1960) 56; Blumea 11 (1961) 113; ibid. 12 (1963) 
90. 

var. gaultheriifolium. 

Slender shrub, 0.4— 2.5(-5) m, branches numer- 
ous, rather stout, erect, terete. Branchlets slender, 
tips covered with stalked scales, lower parts densely 
verruculose. Leaves scattered, more crowded to- 
wards the ends of the branchlets, ovate, rarely 
elliptic-ovate, shortly obtusely acuminate, base 
rounded to subtruncate, rigidly coriaceous, light 
or yellowish green, paler below, subdensely 
lepidote on both faces (scales ± round, sessile, 
entire, centre dark, rather thick and ± impressed 
especially on the undersurface), margin ± thicken- 
ed and revolute, irregularly crenulate by the im- 
pressed scales, (5-)6-9 (rarely and but partly in 
the same specimen up to 12) by 4-7 mm, midrib 
slightly impressed above, a little raised beneath, 
nerves indistinct; petiole lepidote, ± 2 mm. 
Flowers ± nodding, solitary or in twos. Outer 
perulae subulate and ± densely lepidote, inner 
ones ovate-acuminate, apiculate, puberulous and 
lepidote dorsally in the upper part, ± puberulous 
inside, densely ciliate, up to 7 by 4 mm. Brac- 
teoles linear, glabrous, up to 7 mm. Pedicels 
slender, densely lepidote and ± laxly minutely 
pubescent, 5-8 (-9 mm). Calyx oblique, disk- 
shaped, densely lepidote outside, obtusely 5-lobed, 
occasionally one or a few lobes elongate-tri- 
angular, acute, up to 1 mm. Corolla deep red or 
sordid to brownish red, sometimes greenish yellow 
or orange, tubular, somewhat zygomorphous, 
11-13 mm long, densely yellow-lepidote outside 
except a wide margin of the lobes, glabrous inside, 
lobes besides the scales laxly hairy outside in their 
upper half and distinctly ciliate, tube straight, 
somewhat widened upwards, 8-10 mm long, 2.5-3 
mm at the base, c. 5 mm below the lobes, the 
latter erect or a little spreading, ovate-quadrangu- 
lar, rounded at the apex and irregularly crenulate- 
denticulate, c. 3 by 2.5 imn. Stamens 10, slightly 
unequal; filaments red, linear at the base, filiform 
upwards, glabrous, 9-10 mm; anthers c. 1 mm. 
Disk glabrous. Ovary red, conical, densely lepidote, 
c. 2.5 mm, gradually attenuate to the style, the 
latter thick, lepidote at base, otherwise glabrous, 
c. 7 mm; stigma shortly club-shaped, irregularly 



Jan. 1966] 



Ericaceae (Sleumer) 



485 



lobed. Capsule erect, narrow-conical, longitudi- 
nally 5-ribbed. densely lepidote, 8-9 by 3.5-4 mm, 
on stoutish pedicel 1-1.5 cm. 

Distr. Malesia: New Guinea (Oranje Mts, Mt 
Doorman, and in the Eastern and Western High- 
lands). 

Ecol. In shrubberies bordering the (sub)alpine 
forest, occupying patches of wet sandy or peaty 
soil, or on the more sterile peaty ridges, also in 
grassland above the timber line, between summit 
rocks, locally common, occasionally dominant, 
2900-4150 m. Fl. Feb.-Oct. 

vur. expositum Sleum. Reinwardtia 5 (I960) 56; 
Blumea 11 (1961) 113. 

Shrub of erect ascending or prostrate habit, 
0.05-1 m. Leaves very coriaceous and shining, 
(2-)3-4(-5) by 3-^ mm. Flowers dark red or 
orange red. Capsule 6-7 by 2.5-3 mm. The alpine 
form with smaller leaves and capsules. 

Distr. Malesia: W. New Guinea (Mt Wilhel- 
mina and Mt Antares). 

Ecol. Xerophytic shrub on sterile limestone 
slopes, on exposed sandstone crests or in alpine 
rocky grassland, locally common, 3300-3800 m. 
Fl. fr. July, Sept. 

6. Rhododendron oreites Sleum. Reinwardtia 5 
(1960) 57. 

rar. oreites. 

Erect, slender, terrestrial, sparsely branched 
shrub, 2-2.5 m. Branchlets terete, 1.5-2 mm 0, 
tips densely set with stalked scales, glabrescent 
and verruculose-rugose in the lower parts. Leaves 
scattered, crowded especially at the upper part 
of the new shoots, oblong-elliptic, gradually sub- 
acuminate towards the apex, subobtuse, base 
broadly attenuate into the petiole or subtruncate- 
rounded, coriaceous, subdensely persistently lepi- 
dote on both faces (scales round, the narrow mar- 
ginal zone entire or subentire, centre rather large, 
thick, somewhat impressed), revolute at the lower 
part of the edge, which is subcrenulate by the 
impressed scales, 8-12 by 4—6 (rarely up to 8) 
mm, midrib slightly impressed above, prominent 
beneath, nerves very slightly or not visible at all; 
petiole rather slender, lepidote, c. 2 mm. Flowers 
solitary. Outer perulae triangular, inner ones ovate 
to oblong, apiculatc, firmly membranous, lepidote 
dorsally at the apex, otherwise glabrous, rather 
long and densely ciliate, up to 5 by 2.5 mm, often 
surrounding the base of the pedicel at anthcsis or 
still in fruit. Bracteoles filiform, laxly hairy, r. 6 
mm. Pedicels slender, with dense scales on very 
short stalks, not hairy at all, c. I cm at anthcsis, 
up to 1.5 cm in later stages. Calyx oblique, 
cupular, c. 2.5 mm 0, shortly 5-lobed, lobes 
subacute or obtuse, apprcsscd to the corolla, 
c. 1 mm. Corolla tubular, slightly /ygomorphic, 
red, 1.6-1.7 cm long, r. 3 mm « at the base, widened 
to r. 6 mm below the lobes, densely lepidote out- 
side, glabrous inside, the margin of the lobes 
excepted, lobes nearly rounded, the marginal part 
glabrous or very laxly and shortly hairy outside. 



glabrous inside, edgecrenulateorerose-denticulate, 
not ciliate, 3—4 mm o. Stamens 10, as long as the 
corolla, slightly unequal; filaments glabrous, 
linear below, filiform distally; anthers brown, 
obovate-oblong, c. 1 mm. Disk glabrous. Ovary 
shortly ovoid-cylindrical, ± densely covered with 
very slightly stalked or sessile scales, c. 4 mm by 
1.5 mm, gradually tapering to the rather slender 
and glabrous style, which equals the corolla at 
anthesis, but exceeds it in later stages. Capsule 
cylindric, (0.8-)l-1.2 by c. 0.35 cm, deeply 5- 
grooved, rather densely lepidote, valves not or 
slightly twisted. 

Distr. Malesia: W. New Guinea (Mt Wilhel- 
mina). 

Ecol. In subalpine forest or forest borders, or in 
shrubberies, occupying patches of wet sandy or 
peaty soil, common, 2900-3650 m. Fl. fr. Sept.- 
Oct. 

var. chlorops Sleum. Reinwardtia 5 (1960) 58. 

Erect, sparsely branched shrub, 1-2.5 m. Leaves 
elliptic or ovate-elliptic, 1 1-1 5 by 5-8 mm. Corolla 
greenish yellow. 

Distr. Malesia: W. New Guinea (Lake Habbe- 
ma), once collected, said to be plentiful in a 
shrubby opening in the forest at 3000 m. Fl. fr. 
Oct. 

7. Rhododendron erosipetalum J. J. S. Nova Guinea 
18 (1936) 91, t. 18, 1; Kaneh. & Hatus. Hot. Mag. 
Tokyo 56 (1942) 479; Sleum. Reinwardtia 5 
(1960) 58; Jahrb. Rhodod. Ges. Bremen (1963) 
f. 30; Blumea 12 (1963)90. 

Erect and spreading shrublet or shrub, (0. 1-) 
0.5-2 m, epiphytic or terrestrial. Branchlets slender, 
tips densely clad with stalked scales, lower parts 
verruculose where the proper scales have gone and 
the stalks remained. Leaves J: scattered, often 
somewhat crowded in the upper part of the branch- 
lets, narrow-obovate, apex obtusely attenuate or 
mostly rounded, not rarely slightly retuse, gradu- 
ally cuneate at the base, coriaceous, dark green 
above, paler beneath, glabrous above at full age, 
rather laxly lepidote beneath (scales round, their 
marginal zone narrow, entire, the centre darker, 
thickish, slightly impressed), somewhat revolute 
at margin, 6-IO(-l3) by (2-)2..5-3.5(-4, rarely in 
part up to 6) mm, midrib slightly or obscurely 
impressed above, raised beneath, nerves obsolete; 
petiole 1-1.5 mm. Flowers solitary. Perulae ovate, 
apiculatc, subsericeous and laxly lepidote dorsally, 
shortly white-ciliate, up to 4 by 3 mm. Bracteoles 
filiform, up to 5 mm. Pedicels slender, laxly to 
subdensely covered with scales and minute, 
whitish, i patent hairs, 8-10 mm, accrescent to 
15 mm in final stages. Caly.x small, disk-like, 
5-angular or shortly obtusely 5-lobed, lepidote. 
Corolla tubular, somewhat curved and a little 
compressed laterally, deep pink to dark red, all 
over I laxly lepidote, moreover minutely hairy 
at the upper part of the lobes outside, glabrous 
inside. (1.5 -)l. 8 19 cm long in all, 3-4 mm o at 
base, gradually widened to 5-6 mm o below the 
lobes, the latter unequal, obliquely ovate or 



486 



Flora Malesiana 



[ser. I, vol. 6* 



ovate-oblong, i rounded at the apex, though 
minutely denticulate-erose there, 3-4 by 3 mm. 
Stamens 10, unequal, some slightly exserted from 
the corolla; filaments narrow-linear, glabrous; 
anthers c. 1.2 mm. Disk glabrous. Ovary obliquely 
conical, longitudinally 5-ribbed, densely lepidote, 
c. 3 mm, tapering to the style, the latter glabrous, 
somewhat thickened upwards to the club-shaped 
irregularly lobed stigma, c. 1.3 cm. Capsule sub- 
cylindric, deeply 5-grooved, (0.8-)l-1.5 by c. 0.3 
cm. Seeds shortly tailed at both ends, 0.15-2 mm. 

Distr. Malesia: W. New Guinea (Vogelkop 
Peninsula: Arfak Mts, Tohkiri, Tamrau and 
Nettoti Range). 

Ecol. Rarely on burnt open summit, mostly on 
edge of (moss) forest or summit scrub on peaty 
soil, 1400-2650 m. Fl.fr. Jan. -Dec. 

Vern. Mambau, Kebar. 

8. Rhododendron hameliiflorum Wernh. Trans. 
Linn. Soc. H, Bot. 9 (1916) 98; Sleum. Reinwardtia 
5 (1960) 58. 

Very low, perhaps creeping shrublet. Branchlets 
slender, tips densely covered with stalked scales, 
or verruculose in the lower parts where the scales 
have gone. Leaves scattered or subopposite, 
broadly obovate to suborbicular, apex rounded 
or minutely retuse, base broadly attenuate or 
mostly rounded, coriaceous, rather laxly lepidote 
initially (scales round, entire, flattish, the large 
dark centre slightly impressed), early glabrescent 
above, persistently lepidote beneath, entire, 
margin not or very slightly recurved, 7-10 by 
5-8 mm, midrib impressed above, prominent 
beneath, nerves i obsolete; petiole 1.5-2 mm. 
Flowers solitary or rarely in twos. Ferulae oblong- 
obovate, sparsely pubescent dorsally, densely 
white-ciliate. Pedicels slender, densely lepidote, 
5-6 mm. Calyx small, obscurely 5-lobed. Corolla 
tubular, 1.7-2 cm by 3-4 mm, subdensely lepidote 
all over outside, glabrous inside, lobes obovate- 
suborbicular, not erose, c. 4 by 3 mm. Stamens 10, 
nearly equalling the corolla in length; filaments 
linear, glabrous; anthers 1 mm. Disk glabrous. 
Ovary subconical, c. 3 mm, densely lepidote, tape- 
ring to the style, which is laxly lepidote in its lower 
\, glabrous elsewhere, and equals or slightly 
exceeds the corolla in length; stigma club-shaped. 
Distr. Malesia: W. New Guinea (MtCarstensz), 
between 2350 and 3350 m, once collected. 

9. Rhododendron lindaueanum Koord. Nova 
Guinea 8 (1912) 878; J. J. S. ibid. 12 (1914) 129, 
incl. var. latifolium J. J. S. I.e. 130, t. 29A; ibid. 12 
(1917) 498; ibid. 18 (1936) 89; Sleum. Rein- 
wardtia 5 (1960) 58, incl. var. psilacrum Sleum. et 
var. cyclopicum Sleum. /. c. 59; Blumea 12 (1960) 
90. — R. retusum [non (Bl.) Benn.] Wernh. 
Trans. Linn. Soc. H, Bot. 9 (1916) 94. — R. 
fuchsioides Schltr, Bot. Jahrb. 55 (1918) 147. 

var. lindaueanum. 

Low shrub,0.25-2(-3)m, terrestrial or epiphytic; 
branches stoutish. Branchlets terete, slender, tips 
densely covered with stalked scales, lower parts 
verrucose by the remaining stalks. Leaves scattered, 



mostly zb densely together at the upper 2 or 3 
nodes, sometimes 2-4 arranged in a kind of whorl, 
obovate to spathulate or sometimes oblong- or 
elliptic-obovate, apex rounded, not rarely slightly 
retuse, base cuncate, coriaceous or subcoriaceous, 
dark green above, greyish green below, a little 
shining and laxly lepidote or completely glabres- 
cent at maturity above, persistently laxly lepidote 
beneath (scales red-brown, disk-shaped, marginal 
zone narrow, entire, centre thick, not or slightly 
impressed), margin r markedly revolute, lamina 
not rarely l convex, (0.8-) 1-2.5 by (0.5-)0.8-1.3 
(-1.8) cm, midrib t distinctly impressed above, 
prominent beneath, nerves 3-4(-5) pairs, lower 
1-2 ones ascending, upper ones spreading, 
anastomosing, faintly impressed above, somewhat 
raised beneath, occasionally hardly visible; petiole 
stoutish, lepidote or verruculose, (1.5-)2-2.5 mm. 
Flowers solitary. Ferulae unequal, outer ones 
narrow-ovate, subacute, lepidote outside, inner 
ones larger, broadly ovate to suborbicular, lepidote 
and/or greyish sericeous dorsally at least in the 
upper part, ciliate, up to 7 by 4 mm. Bracteoles 
linear to spathulate, hairy towards the top, up to 
7 mm. Pedicels slender, densely lepidote and 
whether or not laxly short hairy, 7-10 mm. Calyx 
minute, oblique, lepidote, obtusely 5-denticulate, 
whether or not shortly ciliate, lobes c. 0.5 mm, 
spreading or reflexed. Corolla tubular, slightly 
zygomorphic and laterally compressed, straight, 
somewhat pouched at the base, deep red to pink, 
laxly lepidote all over outside, glabrous inside, 
(1.7-)1. 8-2.4 cm long in all, tube 1.6-1.8(-2) cm, 
c. 2.5 mm at base, 4-5 mm at the throat, lobes 
ovate-oblong, obtuse, 3^(-5) by ± 2.5 mm, 
erect or slightly spreading, often provided with 
minute hairs near the apex outside and ciliate. 
Stamens 10, slightly or not exserted, unequal; fila- 
ments filiform, glabrous; anthers blackish purple, 
oblong-ovate, 1-1.2(-1.5) mm. Disk glabrous or 
rarely slightly hairy. Ovary narrowly ovoid- 
conical, densely lepidote, c. 2.5 by 1.5 mm, 
gradually tapering to the style, the latter 1-1.3 
cm at anthesis, accrescent and slightly exserted in 
later stages, glabrous, swollen distally to a club- 
shaped, shortly and irregularly 10-lobed stigma. 
Capsule subcylindrical, 1-1.5(-1.8) by 0.3-0.4 cm, 
densely lepidote, deeply 5-grooved. 

Distr. Malesia: New Guinea (Arfak Mts to Mt 
Saruwaged), many collections. 

Ecol. In forest, generally epiphytic, but also 
terrestrial in swampy, shrub-covered places, or in 
alpine scrub on exposed summits, on poor, often 
clayey or sandy soil, 1250-2500(-3200) m. Fl. 
Jan. -Dec. 

Vern. Kieta, kogle, Kapauku, tambies, Hattam. 

var. bantaengense J. J. S. in Fedde, Rep. 30 (1932) 
163; H. J. Lam, Blumea 5 (1945) 576, //; text; 
Sleum. Reinwardtia 5 (1960) 59. 

Differs from the typical variety by longer calyx 
lobes (c. 1 mm) which are ± appressed to the 
corolla. Corolla lobes not ciliate. 

Distr. Malesia: SW. Celebes (Mt Bantaeng or 
Peak of Bonthain), in brushwood at 2800-2890 m. 



Jan. 1966] 



Ericaceae (Sleumer) 



487 



10. Rhododendron spathulatiim Ridl. J. Str. Br. 
R. As. Soc. n. 61 (1912) 25; J. Fed. Mai. St. Mus.6 
(1915) 49; Fl. Mai. Pen. 2 (1923) 220; Sleum. 
Reinwardtia 5 (1960) 59. 

Shrub up to 1.5 m, epiphytic or on moss- 
covered ground, with slender branches. Branchlets 
dark brown, slender, the upper 3 or 4 internodes 
densely clothed with stalked scales, verruculose 
when the scales have gone, older parts smooth 
and soon greyish corticate; internodes 0.5-2 cm. 
Leaves in spaced 3-7-merous pseudowhorls at the 
upper nodes, obovate-spathulate, ape.x rounded, 
mostly slightly retuse, base cuneate, coriaceous, 
la.xly impressed-lepidote to glabrous and ± 
shining above, more densely and rather minutely 
lepidote and dull beneath (scales round, marginal 
zone narrow, entire or nearly so, centre rather 
dark and th'ck, manifestly impressed), entire or 
minutely crenulate, margin flat or slightly revolute, 
1-2 by 0.5-1 cm, midrib mostly impressed above 
in the lower half, prominent beneath, nerves 3-4 
pairs, faintly impressed above and raised beneath, 
or almost invisible; petiole rather slender, ± 2 
mm, lepidote. Flowers solitary, or rarely in twos. 
Ferulae ovate, subacute, lepidote dorsally up- 
wards, ciliate, up to 5 by 3 mm. Pedicels slender, 
densely lepidote, sometimes also minutely laxly 
hairy, 6-8 mm. Calyx oblique, very small, lepidote, 
obscurely obtusely 5-lobed. Corolla tubular, 
slightly dilated towards the mouth, --,: zygomor- 
phous and — laterally compressed, rather straight, 
dark red or salmon red, 2-2.2(-2.6) cm long in all, 
tube 1 .8-2.2 cm, c. 3 mm o at base, 5-6 mm below 
the lobes, laxly lepidote outside, glabrous inside, 
lobes obovate, somewhat spreading, manifestly 
ciliate, 4-5 by 2.5-3 mm. Stamens 10, unequal, 
manifestly exserted; filaments glabrous, linear; 
anthers oblong, 2 by 0.8 mm. Disk glabrous. 
Ovary broadly ellipsoid, densely lepidote, epilose, 
2.5 by 1.5 mm. gradually attenuate into the glabrous 
style which is finally long exserted; stigma capitate, 
indistinctly lobed. Capsule short-cylindric or 
-conical, 7-8 by 2.5-3 mm, densely lepidote. 

Distr. Malesia: Malay Peninsula (Perak: Mt 
Kerbau; Pahang: Mt Siku, Ulu Tclom). 

Ecol. Mossy forest, locally plentiful on Mt 
Kerbau, 1525-2135 m. 

II. Rhododendron saruwagedicum Foerster in 
Fcddc, Rep. 13 (1914) 222; Schltr, Bot. Jahrb. 55 
(1918) 148; Slei;m. Reinwardtia 5 (I960) 59, 
incl. var. alpinum; Blumca II (1961) 113; ihiJ. 12 
(1963) 91; ibid. 12 (1964) 339. — Fig. 5. 

Small tree or tree-like shrub, rarely up to 12 m 
high and with a trunk up to 12 cm 0. reduced to a 
small shrub hardly 0.4 m high at alpine altitudes, 
much branched. Branchlets terete, erect, 

ultimate parts densely clad with stalked rusty 
scales, verruculose in the lower parts where the 
scales have gone. Leaves scattered, laxly set. more 
congested distally, but not in distinct, spaced 
pseudowhorls. ovate to ovate-elliptic, sometimes 
oblong-ovatc, apex obtusely acuminate, base 
rounded to subtruncatc, coriaceous, ■ shining 
above, nearly rubiginosc and dull beneath. 




Fig. 5. Rhododendron saruwagedicum Foerster, 
small-leaved alpine form. a. Branchlet with flowers, 
X 2/3, b. stamen, X 2, c. ovary and style, x 2, 
d. young fruit, x 2, e. ripe fruit, x 2 {a-e Pullen 
223). 

densely lepidote above still at maturity, or at least 
the impressed centre of the scales well visible there, 
very densely covered with copper-coloured or 
rusty scales beneath (scales sessile, round, partly 
in touch with each other, marginal zone narrow 
and entire or nearly so, centre thickish, mani- 
festly impressed), minutely crenulate by the im- 
pressed scales along the subrevolute margin, (0.7-) 
1-1.8 by (0.5-)0.7-l(-l.2) cm, midrib impressed 
above, rather strongly raised beneath, nerves 3-4 
pairs, lower 1-2 curved-ascending, higher ones 
more spreading, all very slightly impressed above 
in fully mature leaves, slightly raised beneath, or 
sometimes hardly visible on both faces; petiole 
stoutish, lepidote, 1-2 mm. Flowers (3-)4-5 per 
umbel. Outer perulae elongate-triangular, densely 
lepidote. inner ones elliptic-ovate to obovate, r 
apiculate, sericeous and lepidote dorsally in the 
upper part, up to 8 by 6 mm. Bracteolcs linear to 
linear-spathulate, finely hairy, up to 6 mm. Pedicels 
rather stoutish, densely rubiginous-lepidotc, not 
hairy, 6-9 mm. Calyx minute, indistinctly obtusely 
5-lobed. Corolla tubular, slightly zygomorphous, 
straight or a little curved, rather densely yellow- 
lepidote outside, completely glabrous inside, dull 
red, 1 .2-1 .3(-l .5) cm long in all. c. 3 mm o at base, 
widened to c. 5 mm at the mouth, lobes 4 or 5, 
t erect, obovate to suborbicular, r. 3 by 2 3 mm, 
not ciliate. Stamens 8 or 10. subcqualling the 
corolla in length; filaments linear, glabrous; 
anthers • obovate, I mm. Disk glabrous, with 
thick, prominent lobes. Ovary narrow-ellipsoid, 
all over covered with scales, r. 4 by 1.5 mm. with 
4 or 5 longitudinal ribs; style abrupt with the 



488 



Flora Malesiana 



[ser. I, vol. 6* 



ovary, glabrous, 5-6 mm, slightly club-shaped 
distally; stigma shortly 5-lobed. Capsule sub- 
cylindric, densely lepidote, (11-)13-15 by 3.45-4.5 
mm. 

Distr. Malesia: New Guinea (Morobe Distr.: 
Mt Saruwaged; Eastern Highlands: Mt Michael; 
Western Highlands: Kubor Range). 

Ecol. Open bush edge or sheltered forest near 
timberline, or (with smaller leaves) in open (sub)- 
alpine grassland, 3050-3820 m. Fl. March-Oct. 

Note. The scales are at least in part rather 
intermediate between those found in subsect. 
Pseudovireya subsect. Euvireya, and subsect. Albo- 



12. Rhododendron invasorium Sleum. Reinwardtia 
5 (1960) 60; Blumea 11 (1961) \\7,;ibid. 12 (1963) 
91; Jahrb. Rhodod. Ges. Bremen (1963) f. 11. 

Erect, stiff shrub, 0.3-1 m. Branchlets densely 
set with round thickish stalked scales on the young- 
er, somewhat angular and redbrown parts (c. 2 
mm 0), subdensely verruculose in the older ones 
(3-4 mm 0); internodes (l-)2-6 cm. Leaves in 
3-4-merous pseudowhorls at the upper 2( — 3) 
nodes, 1 or 2 additional ones scattered in the 
upper third of the internodes, elliptic or more 
rarely obovate-elliptic, apex rounded, the apical 
gland rather indistinct, base very broadly attenuate 
or rounded, rarely very slightly cordate, coriaceous, 
sordid olivaceous to dark green and a little shining 
above, rather densely lepidote on both faces 
initially, glabrescent above with age, subdensely 
and persistently lepidote beneath (scales rather 
small, distant from each other, intensely rufescent, 
orbicular, entire, the rather large centre much 
impressed), margin cartilagineous, not revolute, 
regularly minutely crenulate, (1.5-)2-3.5 by (1.2-) 
1.5-2(-2.2) cm, midrib narrow and depressed 
above, as broad and prominent as the petiole in 
its lower part, gradually less so distally beneath, 
nerves 4-6 pairs, irregular, curved-anastomosing, 
impressed above but in mature leaves, prominent 
beneath, veins laxly reticulate, rather indistinct; 
petiole flattened, c. 3-4 by 1.5 mm. Umbels 3^- 
flowered. Flower buds obovoid, c. 1 by 0.5 cm. 
Outer perulae broadly ovate, inner ones oblong- 
obovate, obtuse, very densely lepidote at the upper 
part of the mediane dorsally, the margin shortly 
fimbriate, 4-8 by c. 4 mm. Pedicels stoutish, 
densely substipitate-lepidote, 1-1.5 cm. Calyx 
obliquely disk-shaped, densely lepidote dorsally, 
obscurely lobed, c. 3 mm 0. Corolla tubular, deep 
scarlet or magenta, a little zygomorphous, some- 
what dilated apically, 1.5-1.8 cm long in all, 
densely lepidote outside, glabrous inside, tube 
1-1.2 cm, 3-4 mm o at base, lobes erect, obovate, 
5-6 by c. 4 mm. Stamens 10, unequal, the longer 
ones attaining half the length of the corolla lobes; 
filaments linear below, filiform above, quite 
glabrous; anthers obovate, 1.8 by 0.6 mm. Disk 
glabrous. Ovary conical, abrupt with the style, 
very densely lepidote, c. 4 by 2 mm; style lepidote 
at the base, glabrous for the rest, 5-6 mm; stigma 
capitate, shortly 5-lobed. 

Distr. Malesia: New Guinea (Morobe Distr.: 



Wau-Edie Creek area). 

Ecol. Common on open gravelly or clayey 
ground of old gold workings, coloniser in sluiced 
area, at 1800-2000 m. Fl. May-Sept. 

13. Rhododendron perakense. K. & G. J. As. Soc. 
Beng. 74, ii (1905) 76; Ridl. Fl. Mai. Pen. 2 (1923) 
216; Sleum. Reinwardtia 5 (1960) 60. 

Small shrub, up to c. 1.8 m, whether or not 
epiphytic. Branchlets slender, terete, tips densely 
covered with stalked scales, otherwise densely 
verruculose at the upper 3-5 internodes, which 
are 0.5-4 cm long. Leaves in 3-5(-7)-merous 
pseudowhorls at the apical nodes, early caducous 
at the lower nodes, obovate-spathulate or broadly 
oblanceolate, apex rounded-obtuse, sometimes 
minutely retuse, base cuneate into the petiole, 
coriaceous, stiff, glabrous above at full maturity, 
laxly to subdensely and more persistently lepidote 
beneath (scales sessile, round, entire or undulate 
at the narrow marginal zone, centre impressed, 
rather large and thick), edge much recurved, 
1.3-1.8(-2) by (0.5-)0.6-l cm, midrib much im- 
pressed above, very stout and prominent beneath, 
nerves 2-3 pairs, spreading, — slightly immersed 
above, mostly well raised beneath, veins obscure; 
petiole rather slender, grooved above, lepidote, 
2-3 mm. Flowers 3-5 per umbel. Outer perulae 
subulate, lepidote dorsally, inner ones — ovate, 
obtuse, lepidote at the top outside, densely white- 
ciliate, up to 6 by 3 mm. Bracteoles not seen. 
Pedicels slender, densely clad with stalked scales, 
not hairy at all, 6-7 mm long at anthesis, up to 
10 mm in fruit. Calyx minute, shortly obtusely 
5-lobed, densely lepidote. Corolla white or bright 
yellow, tubular below, campanulate distally, laxly 
lepidote outside, glabrous inside, seen only in not 
yet fully developed state, at least 10 mm long, 
shortly 5-lobed. Stamens 10; filaments linear, 
somewhat dilated towards the base, glabrous; 
anthers c. 1 mm. Disk glabrous. Ovary conical, c. 
2 mm, densely lepidote; style glabrous; stigma sub- 
capitate. Capsule subcylindric, 9-10 by c. 2.5 mm, 
deeply 5-furrowed longitudinally, lepidote, valves 
a little twisted. Seeds very narrow, c. 3 mm incl. 
the longish tail at both ends. 

Distr. Malesia: Malay Peninsula (Cameron 
Highlands). 

Ecol. In mossy summit forest, locally common, 
at c. 1830 m. Fl.fr. April, Aug. 

14. Rhododendron vanderbUtianum Merr. Not. Nat. 
Ac. Nat. Sc. Philad. 47 (1940) 5; Sleum. Rein- 
wardtia 5 (1960) 61. — Fig. 6. 

Dwarf shrub, up to 1 m high. Branchlets — 
crooked, slender, tips shortly sordid-brownish 
velvety and very laxly lepidote, older parts 
glabrous. Leaves in 3-7-merous pseudowhorls 
at the upper 2 or 3 nodes (internodes 0.3-2 cm), 
ovate or oblong-ovate to elliptic, apex broadly 
acuminate, acutely apiculate, base broadly cuneate 
to rounded, coriaceous, shining on both faces 
especially above, subdensely impressed-lepidote to 
entirely glabrescent on the upper surface, the 
puberulous midrib excepted, — densely lepidote 



Jan. 1966] 



Ericaceae (Sleumer) 



489 




Fig. 6. Rhododendron vanderbiltianum Merr., Mt Goh Lcmbuh, MM) m, 

9039) (VAN Steems, 1937). 



N. Sumatra (Van Steems 



beneath (scales sessile, roundish, margin entire or 
nearly so, rarely crenulate, centre large in relation 
to the marginal zone, thick, dark, impressed), 
margin cartilagineous, crenulate, very slightly or 
not revolute. (l-)1.5-2.3 by (0.8-) 1-1.5(-1.7) 
cm, midrib flat or slightly prominent on both faces, 
nerves 4-5 pairs, • ascending, rather inconspicu- 
ous, veins and veinlcts obsolete above, but mostly 
slightly raised beneath, forming a fine dense 
reticulation: petiole flattened, puberulous, 1.5-2.5 
by I mm. Umbels 2-5-flowcred. Pcrulae ovate to 
obovate-spathulate, apiculate, glabrous or seri- 
ceous and laxly Icpidotc in the upper dorsal part, 
densely ciliatc to lanatc-fimbriatc, up to 1 by 0.6 
cm. Bracteolcs lincar-spathulatc, laxly hairy, 
nearly woolly at the very apex, up to 6 mm. 
Pedicels stoutish, densely lepidote, laxly to sub- 
dcnscly short-hairy, 5-9 mm. Calyx small, obtusely 
*i-dcntcd (I mm), densely lepidote, sometimes 
moreover very laxly hairy, not ciliatc. Corolla 
funnel-shaped, white to cream or pale lemon, 



sometimes bright yellow, 1.8-2.2 cm long in all, 
deeply 5-lobed, ^ laxly lepidote outside, the 
margin of the lobes excepted, whether or not 
slightly to subdensely hairy in longitudinal stripes 
along the faintly pronounced angles of the corolla 
tube, which bears a laxly hairy zone inside above 
the base but is glabrous otherwise inside, tube 
0.8-1 cm long. c. 3 mm o at base, 7-8 mm at the 
throat, lobes somewhat unequal, slightly spreading, 
subobovate-spathulate to suborbicular, c. 1.3 by 
0.8-1.3 cm. Stamens 10, unequal, included; fila- 
ments linear, glabrous at the base, patently hairy 
for 2-4 mm upwards, then again glabrous to the 
top; anthers broadly oblong, c. 2 mm. Disk 
glabrous. 6> van conical, the apex truncate, covered 
with overlapping scales, occasionally bearing also 
some short hairs at the apex, 2.5-3 by 1.5-2 mm; 
style abrupt, slender, glabrous, finally attaining 
nearly 2 cm; stigma short-obconical. rather small, 
5-lobed. C iipsulc (perhaps submature) c. 6 by 2.5 
mm. 



490 



Flora Malesiana 



[ser. I, vol. 6* 



Distr. Malesia: N. Sumatra (Atjeh, Gajolands: 
Mts Losir, Lembuh and Kemiri). 

Ecol. Open heath scrub or open heathy plains 
with Quercus, Finns and other species of Rlwdu- 
dendron, locally common, 2100-3300 m. Fl. Jan.- 
April. 

15. Rhododendron seimundii J. J. S. Card. Bull. 
S. S. 8 (1935) 262; Sleum. Reinwardtia 5 (1960) 61. 

Shrub. Branchlets slender, apical 2 or 3 inter- 
nodes 0.7-6 cm, patent-papillose-puberulous 
(lens!), tips moreover laxly clad with sessile 
scales. Leaves in 4-7(-10)-merous pseudowhorls 
at the upper l-2(-3) nodes, oblong-obovate to 
narrow-elliptic, sometimes nearly lanceolate, apex 
broadly attenuate, mostly obtuse and ending in a 
small somewhat protruding gland, base attenuate 
into the petiole, coriaceous, rugulose when dry, 
shining above, initially laxly lepidote, finally 
glabrous above, rather dull and laxly lepidote 
beneath (scales sessile, round, margin entire or 
nearly so, centre thick, darker, manifestly im- 
pressed), edge entire or minutely crenulate by the 
impressed scales, slightly revolute, 1.3-2.8 by 
0.5-1.1 cm, midrib sunk above, strongly raised 
beneath, nerves obsolete, sometimes faintly 
visible underneath; petiole rather slender, lepidote 
and puberulous, 2-3 (-4) mm. [/mfee/.y2-4-flowered. 
Ferulae ovate, obtuse, i densely short-pubescent 
dorsally, ± laxly lepidote, ciliate, up to 7 by 4 
mm. Bracteoles filiform, hairy, up to 6 mm. 
Pedicels slender, ± densely clothed with sessile 
scales, furthermore whether or not minutely hairy, 
9-12 mm. Calyx oblique, inequally obtusely 
5-lobed, lepidote, not ciliate. Corolla funnel- 
shaped, 1.6-1.9 cm long in all, 1.5-2.5 cm across 
the limb, laxly lepidote outside, margin of the 
lobes excepted, glabrous inside, tube 8-9 mm 
long, c. 2.5 mm at base, 4-6 mm below the lobes, 
the latter obovate, 9-10 by 6-7 mm. Stamens 10, 
very slightly or not exserted; filaments linear, 
glabrous; anthers oblong, c. 1 .5 mm. Disk glabrous. 
Ovary ovoid-conical, obtuse, 5-grooved longitudi- 
nally, densely lepidote, c. 2.5 by 2 mm, gradually 
attenuate to the style, which is glabrous and ± 1.6 
cm long; stigma capitate, 5-lobulate. Capsule 8 by 
2.5 mm. 

Distr. Malesia: Malay Peninsula (Pahang: Mt 
Tahan), 1675 m. Fl. Feb. -March. 

16. Rhododendron scortechinii K. & G. J. As. Soc. 
Beng. 74, ii (1905) 76; Ridl. Fl. Mai. Pen. 2 
(1923) 219; Sleum. Reinwardtia 5 (1960) 61. — 
R. orion Ridl. J. Fed. Mai. St. Mus. 5 (1914) 39; 
Fl. Mai. Pen. 2(1923)217. 

Epiphytic or terrestrial shrub, sometimes sub- 
arboreous. Branchlets terete, wrinkled and red- 
brown at the tips, -t_ smooth below, the ends rather 
densely covered with flat scales. Leaves in 3-5(-7)- 
merous pseudowhorls at the upper 2 or 3 nodes 
(internodes 1.5-5 cm), elliptic to elliptic-subor- 
bicular or (broadly) obovate, apex obtuse to 
rounded and with a small terminal gland, some- 
times a little notched, base narrowed to the petiole, 
upper surface light green, caducously lepidote, 



finally completely glabrous and shining, under- 
surface yellowish when fresh, paler brown when 
dry, densely lepidote (scales sessile, round, mar- 
ginal zone narrow, entire or nearly so, centre 
thick, dark, rather large and f: impressed), edge 
strongly recurved, (2.5-)3-5 by (l.5-)1.8-2.5(-3) 
cm, midrib sulcate above, thick and obtusely 
prominent beneath, nerves 4—6 pairs, spreading, 
anastomosing, slightly to deeply depressed above, 
distinct and normally much raised beneath, veins 
-i: obsolete; petiole grooved above, lepidote, 2-4 
(-5) by 1.5-2 mm. Umbels 4— 7-flowered. Perulae 
ovate to spathulate, obtuse, ± sericeous and 
lepidote in the upper dorsal part, ciliate, up to 7 
by 4 mm. Bracteoles linear, slightly hairy, up to 
5 mm. Pedicels 1.5-2 cm, rather stoutish, ± 
densely lepidote, not hairy at all. Calyx short- 
cupular, lepidote, with 5 ovate, obtuse, whether 
or not ciliate teeth (1 mm). Corolla yellow, 
sometimes paler or even white, or yellow at the 
lobes, orange at the tube, widely funnel-shaped, 
c. 1.8 cm long in all, 1.5-2 cm wide at the limb, 
± densely set with dark scales outside except the 
margin of the lobes, glabrous inside, tube 9-14 
mm long, 3-4 mm at base, 8-10 mm at the throat, 
lobes broadly obovate to suborbicular, 6-8 by 
5-7 mm. Stamens 10, unequal, the longest as long 
as the corolla or slightly exserted; filaments linear, 
dilated (0.5 mm) at the base, filiform upwards, 
glabrous; anthers oblong, c. 1.8 mm. ZJ/^/c glabrous. 
Ovary conical, densely covered with scales, c. 2.8 
by 1.8 mm, gradually tapering to the style, the 
latter glabrous, finally somewhat exserted; stigma 
turbinate-capitate, slightly 5-lobed. Capsule sub- 
fusiform, (1-)1.3 by 0.35 cm, shortly apiculate, 
densely lepidote, on pedicel up to 3.2 cm. 

Distr. Malesia: Malay Peninsula (Main Range 
from Perak to Selangor). 

Ecol. Summit vegetation, 1465-2135 m. Fl.fr. 
Jan. -April. 

17. Rhododendron adinophyllum Merr. Not. Nat. 
Ac. Nat. Sc. Phiiad. 47 (1940) 3; Sleum. Rein- 
wardtia 5 (1960) 61. — Fig. 7. 

Shrub, 1-1.5 m. Branchlets erect, terete, rather 
stoutish, very densely and ± imbricately set with 
spiral, r: ascending leaves in their upper and with 
wartlike scars in the lower defoliate part, tips 
rather densely covered with very shortly or not 
stalked scales, lower parts glabrescent, smooth. 
Leaves narrow lanceolate, gradually acuminate, 
tip obtuse, base attenuate into the petiole, stiff- 
coriaceous, shining above only, mostly glabrous 
at full maturity, but sometimes with vestiges of the 
scales above, rather densely lepidote beneath 
(scales sessile, ± rounded, margin entire to 
undulate, narrow, centre rather large and dark, 
markedly impressed), edge thickened revolute 
and crenulate by impressed scales, (8-)l 1-16(-18) 
by 2-3 mm, midrib i impressed above, obtusely 
raised beneath especially in the lower half, nerves 
obsolete; petiole rather slender, 1-2 mm. Umbels 
(4-)6-12-flowered. Perulae ovate to obovate, 
apiculate, dorsally lepidote and sericeous in the 
upper part, fimbriate, up to 7 by 4 mm. Bracteoles 



Jan. 1966] 



Ericaceae (Sleumer) 



491 




Fig. 7. Rhododendron adinophyllum Merr., Mt 
Goh Lembuh, 3000 m, N. Sumatra (van Steenis 
9062) (van Steenis, 1937) 

filiform, hairy, up to 6 mm. Pedicels slender, all 
over densely longish and whitish patent-pilose, 
laxly lepidote upwards, 1-1.3(-1.5) cm. Calyx 
oblique, small, undulate, whitish hairy and 
lepidote outside and at the margin, glabrous 
inside, subacutcly obtusely 5-lobed (1-1.5 mm). 
Cttrolla tubular, bright crimson to scarlet, all over 
subpatently long-hairy and laxly lepidote outside, 
glabrous inside, tube (10-)12 13 mm, c. 3 mm a 
at the base, r. 5 mm below the lobes, the latter 
straight to subpatcnt, obovale, apex rounded, 4-5 
by 4 mm. Stamens 10, slightly or not cxserted. 



unequal; filaments linear, glabrous; anthers ob- 
long-ellipsoid, c. 1.5 mm. Disk glabrous. Ovary 
ovoid-conical, densely clothed with overlapping 
scales, 2.5-3 by 2 mm, rather abruptly attenuate 
to the glabrous style (4-5 mm); stigma elongate- 
obconical, subtruncate. Capsule elongate-ovoid 
to subovate-oblong, densely lepidote, 5-ribbed, 
c. 11 by 4 mm, crowned by a glabrous style c. 
1 cm long with a turbinate-capitate stigma, on a 
glabrescent pedicel up to 1.8 cm. 

Distr. Malesia: N. Sumatra (Atjeh, Gajolands: 
Mts Losir, Lembuh, Putjuk Angasan and Kemiri). 

Ecol. In open places, summit heath scrub, 
2600-3460 m. Fl. Jan. -April. 

18. Rhododendron cyrtophyllum Wernh. Trans. 
Linn. Soc. II, Bot. 9 (1916) 97; Sleum. Reinwardtia 
5 (1960) 61. 

Shrub. Branchlets rather stoutish, terete, tips 
densely covered with sessile or very shortly stalked 
roundish scales, lower parts glabrescent, ± smooth, 
internodes 2-4 cm. Leaves in 3-5-merous pseudo- 
whorls at the upper nodes, sometimes scattered 
but close together, elliptic or ovate-elliptic, apex 
gradually attenuate, obtuse, apiculate, base round- 
ed or subtruncate, thin-coriaceous, rather stiff, 
subdensely to laxly lepidote on both faces initially 
(scales round, entire, marginal zone narrow, 
centre thick, very little or not impressed), glabres- 
cent above with age, persistently lepidote beneath, 
edge much revolute, 2.5-3 by 1.2-1.9 cm, midrib 
slightly impressed above, very prominent beneath, 
nerves 5-6 irregular pairs, spreading, a little raised 
above, more distinctly so beneath, reticulation 
obscure; petiole stoutish, lepidote, 2-3 by 0.8 mm. 
Umbels 5-8-flowered. Ferulae ovate, obtuse, 
glabrous except some scales near the top dorsally, 
up to 10 by 6 mm. Pedicels rather slender, lepidote, 
0.7-1.2 cm. Calyx oblique, obscurely 5-lobed, c. 
2.5 mm 0. Corolla pinkish red, tubular below, 
dilated upwards, c. 3.5 cm long in all, very densely 
lepidote outside, glabrous inside, tube curved, 
c. 4 mm at base, 7-10 mm below the lobes, 
the latter - spreading, rounded, c. 8 mm 0'. 
Stamens 10; filaments linear, sparsely pubescent 
below, glabrous upwards, c. 2.5 cm; anthers broad- 
oblong, 2.5 mm. Disk glabrous. Ovary densely 
lepidote and laxly patently short-hairy, r. 5 by 2 
mm; style rather abrupt with the ovary, glabrous 
c. 2 cm. 

Distr. Malesia: New Guinea (Mt Carstensz, 
along the Utakwa R.), between 2520 and 3050 m. 
Once collected. Fl. Jan. 

19. Rhododendron ericoides Low ex Hook. /. in 
Hook. Ic. PI. (1852) t. 887; Wai p. Ann. 5 (1858) 
446; Stai'f, Trans. Linn. Soc. II. Bot. 4 ( 1894) 198; 
CiiBHS, J. Linn. Soc. Bot. 42 (1914) 104; Mfrr. 
Rn. Born. (1921) 461; Sli UM. Reinwardtia 5 
(1960) 61; Blumea 12 (1963) 91; Jahrb. Deutsch. 
Rhodod. Ges. Bremen (1965) f. 15. — Fig. 8. 

var. ericoides. 

Erect virgatc shrub, (l-)2-3 m, reduced to a 
dwarfed habit and c. 15 cm height at its highest 



492 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 8. Rhododendron ericoides Low ex Hook. /. 
var. ericoides. Mt Kinabalu, c. 3800 m, N. Borneo 
(Sleumer, 1963) 

localities, occasionally epiphytic. Branchlets slen- 
der, densely set with wartlike leaf-cushions in the 
lower defoliate parts (no fine pubescence present, 
which generally is found in 25. R. quadrasianum). 
Leaves spirally arranged, very dense and — im- 
bricate in the upper part of the branchlets, ericoid, 
suberect, linear-lanceolate, the very apex with an 
obtuse glandular point, base attenuate into the 
petiole, coriaceous, glabrous above, laxly lepidote 
beneath (scales disk-shaped, entire, centre darker, 
thick, slightly impressed), margin crenulate by im- 
pressed scales, somewhat thickened and slightly 
or not revolute, 4— 7(-10) by i 1 mm, midrib very 
slightly depressed above, swollen underneath, 
nerves obsolete; petiole thick, 0.5 mm. Flowers 
solitary or in twos, rarely 3 in an umbel, nodding. 
Ferulae ovate- to obovate-oblong, lepidote dorsal- 
ly in the upper part, densely subvillous-fimbriate, 
especially near the apex, up to 4 by 1.5 mm, 
early caducous. Bracteoles linear-spathulate, long- 
ciliate upwards, c. 3 by 0.5 mm, early caducous. 
Pedicels rather slender, 5-8 mm, densely lepidote, 
epilose, — hidden among the leaves for the lower 
half. Calyx slightly oblique, lepidote outside, 
glabrous inside, 5-partite, lobes erect, initially 
appressed to the corolla, spreading in later stages, 
narrowly subovate-triangular, obtuse, c. 1 mm 
long, 1-1.3 mm wide at the base. Corolla bright 
scarlet- or purplish red, tubular, not or slightly 
oblique and curved, (1-)1.5 cm long in all, sub- 
densely pale greenish lepidote outside, the marginal 
part of the lobes excepted, glabrous inside, tube 
0.8-1 cm long, c. 3 mm at the base, 4-5 mm at the 
throat, lobes broadly ovate to rounded, suberect 
or a little spreading, margin erose-denticulate, 
2-3 mm 0. Stamens 10, ± equalling the corolla or 



slightly exserted, if the corolla lobes are more 

expanded; filaments filiform, glabrous; anthers c. 
0.7 mm. Disk glabrous. Ovary conical, green, 
densely lepidote, c. 1.8 mm, gradually attenuate 
to the red style, the latter glabrous, finally exserted, 
up to 14 mm; stigma club-shaped. Capsule oblong- 
ovoid, often a little oblique and curved, 6-7 by 
2.5-3 mm. 

Distr. Malesia: North Borneo (Mt Kinabalu). 

Ecol. Shrub in primary mossy forest from 
2440 m upwards, abundant, and a very common 
constituent of the Ericaceous (scrub)forest on 
exposed sunny places, also in crevices on the open 
granite core of Mt Kinabalu, running almost to 
the summit, reduced there to a spreading wind- 
clipped dwarf shrublet at c. 4000 m. Fl.fr. March- 
Nov. 

var. silvicolum Sleum. Reinwardtia 5 (1960) 62; 
Blumea 12 (1963) 91. 

Differs from the typical plant (with which it has 
the large calyx lobes in common) by less densely 
arranged and wider leaves, 8-10 by 1.6-1.8(-2) 
mm. Bushy c. 1 m high undershrub. Corolla (pale) 
red. 

Distr. A/a/ei/a: North Borneo (Mt Kinabalu). 

Ecol. Forest and more open vegetation, 1525- 
3000 m. Possibly of hybrid origin. 

20. Rhododendron nanophyton Sleum. Reinward- 
tia 5 (1960) 62. 

var. nanophyton. 

Dwarf shrub. Branchlets erect, slender, tips — 
papillose-puberulous, densely foliate, lower parts 
set with the scars and the thickish cushions of 
fallen leaves. Leaves scattered, lanceolate, apex 
short-attenuate, subacuminate, somewhat obtuse 
at the very top, base cuneate, coriaceous, laxly 
lepidote on the upper surface for a rather long 
time, finally glabrescent, persistently lepidote 
beneath (scales lax to subdense, round, marginal 
zone narrow, entire or a little wavy, centre dark 
red-brown, thick, distinctly impressed), not or 
slightly recurved on edge, crenulate by impressed 
scales, 4—6 by 1.5-2(-2.5) mm, midrib slightly 
impressed above, not much raised beneath, nerves 
obsolete; petiole rather slender, semiterete, c. 
1 mm. Flowers solitary or in twos. Perulae ovate 
to narrow-ovate, acuminate, minutely pubescent 
and laxly lepidote in the upper part outside, ciliate, 
up to 5 by 2 mm. Bracteoles filiform, subspathulate 
at the end and densely ciliate, 4-5 mm. Pedicels 
very slender, densely lepidote, 5-6 mm. Calyx 
minute, disk-shaped, wavy and obscurely 5-lobed 
at the spreading limb, lepidote outside, glabrous 
inside. Corolla dark brick-red, tubular, somewhat 
oblique, — straight, 1.3-1 .4(-l .5) cm long in all, 
2.5 mm at the base, c. 3.5 mm at the mouth, 
densely set with thickish scales outside, margin 
of the lobes excepted, glabrous inside, lobes 
slightly unequal, ovate to nearly rounded, irregu- 
larly crenulate (but not erose-denticulate) at the 
apex, c. 3 by 2 mm. Stamens 10, subequal, as long 
as or slightly longer than the corolla; filaments 



Jan. 1966] 



Ericaceae (Sleumer) 



493 



linear, dilated towards the base, glabrous; anthers 
broadly oblong, 1.3-1.5 mm. Disk glabrous. Ovary 
conical-ovoid, 5-furrowed, densely lepidote, c. 2 
by 1.2 mm, gradually tapering to the style, the 
latter thick-filiform, glabrous, 7-8 mm; stigma 
shortly club-shaped, rather small. 

Distr. Malesia: SW. Central Celebes (Lati- 
modjong Range: Rantemario), in open country at 
3100-3300 m. once collected. Fl. June. 

var. petrophilum Sleu.m. Reinwardtia 5 (1960) 63. 

Differs from the type variety by less distinctly 
papillose-puberulous branchlets, obovate-oblong 
leaves 2.5-4(-6) by 1.5-2(-2.5) mm, obtuse and 
mostly a little retuse at the ape.x and papillose- 
puberulous in the lower half, shortly but distinctly 
ciliate calyx lobes, minutely pubescent pedicels 
and subglobose-obovoid anthers 1 mm. 

Distr. Malesia: SW. Central Celebes (Lati- 
modjong Range), frequent on rocks just below the 
western edge of the plateau of bivouac Heinrich, 
at 3200-3250 m, once collected. Fl. June. 

21. Rhododendron schizostigma Sleum. Reinward- 
tia 5 (1960) 63. 

Weak shrub, 0.3-1.5 m. Branchlets erect, 
slender, tips laxly lepidote, lower parts set with 
the scars and the thick leaf-cushions of the fallen 
leaves. Leaves scattered, dense together and sub- 
imbricate, ovate or elliptic-ovate or elliptic, apex 
shortly obtusely acuminate (terminal gland small), 
base subtruncate-rounded, or more rarely broadly 
cuneate, coriaceous, very laxly lepidote and early 
glabrescent above, laxly and persistently lepidote 
beneath (scales round or nearly so, margin entire 
or subentire, centre mostly rather small, not or 
faintly impressed), edge subcartilagineous and 
flat, crenulate by depressed scales, (5-)6-8(-l2) 
by (3-)3.5-5(-8) mm, midrib minutely or not 
impressed above, slightly raised beneath, nerves 
obscure; petiole c. 1 mm. Flowers solitary or in 
twos. Ferulae ovate-acuminate, membranous, 
glabrous, glossy, up to 1.3 by 0.8 cm. Bracteoles 
linear-subspathulate, glabrous, up to 1 cm. Pedicels 
stoutish, 10-14 mm at anthesis, densely covered 
with subsessile scales. Calyx minute, oblique, disk- 
shaped, wavy, lobes obsolete. Corolla tubular, 
bright red, 2.6-2.9(-3.2) cm long in all, 3-4 mm 
at base, c. 6 mm below the lobes, very laxly 
lepidote below, glabrous upwards outside, glabrous 
inside, tube l-l.li-l.S) cm, lobes broad-elliptic, 
6-7(-8) by4-5(-6) mm, margin undulate. Stamens 
10, unequal, c. 2.5 cm; filaments linear, glabrous, 
somewhat dilated towards the base, filiform up- 
wards; anthers brown, broadly subobovatc-elliptic, 
1.5-1.7 by I mm. Disk glabrous, lobes thick and a 
little prominent. Ovary obliquely ovoid-conical, 
very densely lepidote, r. 4 by 2 mm, rather abrupt 
with the stoutish glabrous style (r. 1.5 cm at 
anthesis); stigma gradually thickened or turbinate, 
its numerous lobes irregularly club-shaped. 

Distr. Malesia: New Guinea (Lake Habbema 
and Hubrccht Mts). 

Ecol. In open vegetation, near Lake Habbema 
pleniirul in heavy ground moss in thickets, or in 



shrubberies of peaty ridges, 3100-3450 m. Fl. 
Feb., Aug. 

22. Rhododendron meliphagidum J. J. S. in Fedde, 
Rep. 30 (1932) 162; Sleum. Reinwardtia 5 (1960) 
64. 

Shrub, up to 5 m. Branchlets terete, tips initially 
laxly set with sessile scales, finally glabrescent, 
smooth; internodes 1.5-6.7 cm. Leaves scattered, 
8-1 1 together at the 2 or 3 upper nodes, obovate, 
apex rounded, shortly retuse, base cuneate into 
the petiole, coriaceous, laxly to subdensely lepidote 
on both faces initially (scales sessile, round, sub- 
entire, centre thick, impressed), — glabrescent 
above with age (marginal zone of the scales then 
obliterated, the impressed centre remaining visible 
for a long time), less quickly or not glabrescent 
beneath, entire, edge distinctlybutslightlyrevolute, 
1.9-4.3 by 0.7-1.9 cm, midrib impressed above, 
strongly raised beneath especially in the lower 
2/3, nerves 4-5 pairs, anastomosing, faintly im- 
pressed above, not rarely obsolete on both sides; 
petiole somewhat flattened, grooved above, lepi- 
dote, c. 2 mm. Flowers solitary. Outer perulae 
subulate, densely lepidote, inner ones ovate, 
acuminate, in part apiculate, puberulous and 
lepidote dorsally, up to 7 by 3 mm. Pedicels 
rather slender, laxly hairy (especially above the 
base) and lepidote, 8-10 mm. Calyx oblique, 
small, obtusely 5-lobed (1 mm), lepidote dorsally, 
margin rather long whitish ciliate. Corolla brown- 
ish orange or yellow, tubular, slightly oblique, c. 
3 cm long in all, laxly lepidote outside, glabrous 
inside, tube c. 2.5 cm long, c. 4 mm at base, 
c. 6 mm below the lobes, the latter ovate, obtuse, 
erect, 5-6 by 4—5 mm, ciliolate. Stamens 10, 
2.5-2.9 cm; filaments linear, flat, nearly 1 mm 
wide at the base, filiform upwards, glabrous; 
anthers suborbicular, c. 1 mm. Disk glabrous. 
Ovary oblong-conical, densely lepidote, c. 3 by 
1.8 mm, gradually developing into a rather thick, 
nearly columnar (1 mm o), glabrous style, c. 2.3 
cm long at anthesis; stigma club-shaped. 

Distr. Malesia: Moluccas: Buru, summit of 
Mt Togha. at 1800-2050 m, a very common shrub 
on open spots; Central Ceram, in primeval forest 
of Ml Murkele, at 2000-2500 m, abundant, on 
limestone. Fl. Feb.. July. 

23. Rhododendron vinkii Sleum. Blumea 12 (1963) 
91. 

Epiphytic shrub. Branches numerous, divaricate; 
branchlets terete, 1.5-3 mm o. densely covered at 
the ends with suborbicular, sessile or subsessile 
scales; internodes 2-4(-5) cm. Leaves in 3-5- 
mcrous pseudowhorls at the upper 2-3 nodes, 
oblong or obovate-oblong, apex broadly attenuate, 
obtuse, rarely slightly retuse, base cuncatc- 
attenuate into the petiole, subcoriuceous, densely 
lepidote on both faces initially, glabrescent with 
age above, persistently lepidote beneath (scales 
roundish or almost so. thickish, impressed, distant 
mutually for 2 or 3 limes their diameter), (3.5-) 
4-6(-7.5) by (l.3-)l.5-2(-2.5) cm, edge slightly 
rcvolule especially towards the base of the lamina. 



494 



Flora Malesiana 



[ser. I, vol. 6* 



midrib a little impressed above, as thick as the 
petiole and much prominent, gradually narrowed 
and less prominent distally below, nerves 6-8 
irregular pairs, somewhat impressed above, ob- 
scure beneath, reticulation faintly impressed above, 
or almost inconspicuous at all; petiole subterete, 
grooved above, densely lepidote, (5-)6-8 by c. 
1 mm. Umbels 2- or 3-flowered. Outer perulae 
ovate, inner ones obovate to spathulate, glabrous 
dorsally, caducously lepidote-fimbriate, thin, up 
to 1 cm. Bracteoles filiform. Pedicels slender, 
rather densely patent-puberulous and lepidote, 
1.2-1.8 cm at anthesis, slightly elongate in later 
stages. Calyx obliquely patellar, indistinctly ob- 
tusely 5-lobed, c. 2.5 mm 0. Corolla subobliquely 
tubular, light red, 2.5 cm long in all, subdensely 
set with rather thickish subrotund scales outside, 
laxly hairy inside, tube erect, c. 2 cm long, 3—4 mm 
at base, c. 6 mm at the throat, lobes spathulate- 
obovate, c. 5 by 6-7 mm. Stamens 10, slightly 
exserted from the throat; filaments linear and 
shortly hairy below, glabrescent in the upper 1/4; 
anthers broad-oblong, hardly 2 mm. Disk densely 
erect-hairy. Ovary cylindric, rather abrupt with 
the style, c. 4 by 2 mm, yellowish tomentose, 
the hairs covering the scales; style rather thickish, 
1.2-1.6 cm at anthesis, densely patent-hairy below, 
less so upwards, glabrous but for c. 2 mm below 
the capitate stigma. 

Distr. Malesia: W. New Guinea (hinterland of 
Fak-Fak), 840 m, in primary Agathis forest on 
clayey soil. Once found. Fl. Feb. 

24. Rhododendron ciliilobum Sleum. Reinwardtia 
5 (1960) 64; Blumea 12 (1963) 92. 

Erect shrub. Branchlets rather short, terete, 
subdensely clad with rubiginous sessile or sub- 
sessile scales, not hairy at all. Leaves several 
together at the tips of the branchlets in a kind of 
pseudowhorl, obovate, apex very shortly retuse, 
rarely rounded, base cuneate, coriaceous, laxly 
lepidote above, though ± glabrescent with age and 
becoming a little rough, subdensely lepidote 
beneath (scales nearly round, sessile, marginal 
zone narrow and thin, subentire, centre rather 
large, thick and manifestly impressed), margin 
very shortly revolute basally, otherwise flat, 
(l.l-)1.3-1.8(-2) by (0.5-)0.6-0.8(-l) cm, midrib 
faintly sunk above, obtusely raised beneath, ± 
evanescent towards the top of the lamina, nerves 
obsolete above, hardly or not visible beneath; 
petiole terete, lepidote, subrugose, 2-3 mm. 
Flowers solitary or in twos. Perulae ovate to 
obovate-spathulate, apiculate, sericeous and lepi- 
dote in the upper dorsal part, densely white- 
fimbriate, up to 6 by 3 mm. Bracteoles filiform, 
hairy at the apex, up to 5 mm. Pedicels rather 
slender, densely to more laxlysubstipitate-lepidote, 
otherwise whether or not laxly short-hairy, 6-8 
(-10) mm. Calyx minute, wavy and spreading, 
or reflexed at the narrow margin, lepidote and 
very shortly pubescent dorsally, c. 2 mm 0, lobes 
very short and obtuse. Corolla tubular, c. 2 cm 
long in all, c. 3 mm at the base, c. 5 mm below 
the lobes, laxly lepidote outside, glabrous inside, 



lobes ovate to suborbicular, 4 by 3-4 mm, erect 
initially, finally !- spreading, manifestly erose- 
denticulate and ciliate at the apex. Stamens 10, 
h as long as the corolla at anthesis, or finally 
somewhat exserted; filaments filiform, glabrous; 
anthers 1.3 mm. Disk glabrous. Ovary subovoid- 
cylindric, c. 2.5 by 1.5 mm, very densely lepidote, 
gradually tapering to the style, the latter thick- 
filiform, glabrous, c. 1.7 cm; stigma club-shaped, 
irregularly lobed. Capsule c. 1.3 by 0.55 cm, 
obliquely subcylindric, deeply 5-grooved. 

Distr. Malesia: W. New Guinea (Wissel Lakes 
region), in heath vegetation at c. 1755 m. Two 
collections. Fl. March, April. 

A form without hairs at the calyx and corolla 
lobes is known from the Balim Valley, at 1600 m. 

25. Rhododendron quadrasianum Vidal, Rev. PI. 
Vase. Filip. (1886) 170; Merr. Publ. Gov. Lab. 
Philip, n. 29 (1905)43; Philip. J. Sc. 1 (l906)Suppl. 
Ill; ibid. 2 (1907) Bot. 292; ibid. 3 (1908) Bot. 
382; Elm. Leafl. Philip. Bot. 3 (1911) 1106; 
MiLLAis, Rhodod. (1917) 233; Merr. En. Philip. 
3 (1923) 244; Millais, Rhodod. ser. 2 (1924) 218; 
CoPEL. / Philip. J. Sc. 40 (1929) 139, t. 1; H. J. 
Lam, Blumea 5 (1945) 574; Sleum. Reinwardtia 5 
(1960) 64. — R. retusum (non Benn.) F.-Vill. 
Nov. App. (1883) 353. 

See for further synonyms under the varieties. 

KEY TO THE VARIETIES 

1. Ovary exclusively lepidote, not hairy at all. 

2. Corolla lepidote and, at least laxly, pubescent 

outside. 

3. Corolla 20 (rarely up to 25) mm long, 

strictly tubular ... 4. var. malindangense 

3. Corolla 10-15(-16) mm long, slightly sub- 

campanulate-dilated distally. 
4. Pedicels pubescent and ± densely lepidote 
at anthesis. Perulae dorsally lepidote and 
pubescent. 
5. Leaves rather broadly and shortly obovate, 
15-20(-28) by (5-)6-8(-10) mm 

2. var. davaoense 

5. Leaves it elongate-obovate, (10-)12-20 
(-22) by 3-5 (rarely up to 6) mm 

3. var. rosmarinifolium 
4. Pedicels pubescent and but very sparsely or 
not lepidote at anthesis. Perulae outside 
glabrous or almost so. 

6. Leaves obovate, (ll-)12-18(-20) by 4-7 
(-8) mm .... 5. var. marivelesense 

6. Leaves narrow-obovate, 8-10(-lI) by 2-3 

(-4) mm 6. var. intermedium 

2. Corolla mostly exclusively lepidote, rarely 
lepidote and also with some hairs at the base, 
or glabrous. 
7. Leaves(15-)17-30by(7-)9-13mm. (Pedicels 
pubescent aw^ lepidote) 1. var. quadrasianum 
7. Leaves 6-16(-20) by 2.5-6 mm. 
8. Leaves ± broadly obovate, (3-)4-5(-6) mm 
wide. Pedicels lepidote and laxly pubescent, 
sometimes exclusively lepidote 

7. var. selebicum 



Jan. 1966] 



Ericaceae (Sleumer) 



495 



8. Leaves narrowly and — elongately obovate, 
2.5-4(-5) mm wide. Pedicels very laxly 
lepidote, nearly always epilose 

8. var. cuneifolium 
1. Ovary lepidote and rather densely hairy to 

almost villous. 
9. Leaves narrowly obovate, (5-)9-16 mm long. 
10. Leaves 3-5 mm wide . . 9. var. villosum 
10. Leaves 1-1.5 mm wide. 

10. var. angustissimum 

9. Leaves broadly obovate, (ll-)16-22 by (5-) 

7-10 mm 11. var. borneense 

1. var. quadrasianiun. — R. quadrasiamim Vidal, 
Rev. PI. Vase. Filip. (1886) 170; Copel./. Philip. 
J. Sc. 40 (929) 140, t. 1, f. 1, 2 {not typical). 

Shrub, 0.3-1 (-3) m, occasionally treelet, erect 
and terrestrial, or rarely epiphytic. Branches 
greyish, the cork peeling offlongitudinally. Branch- 
lets terete, slender, brownish greyish, striate, 
younger parts generally — densely patent-puber- 
ulous, tips moreover laxly to subdensely lepidote; 
internodes 0.5-8(-10) cm, the lower ones leafless. 
Leaves close together or in 4-8-merous pseudo- 
whorls at the upper 2-3 {^) nodes, obovate, 
subsessile, apex broadly obtusely attenuate or 
mostly rounded, very slightly or not retuse, the 
terminal gland small and on the undersurface 
within the edge, base cuneate into the petiole, 
— coriaceous, dark green and shining above, 
dilutely so beneath, laxly lepidote initially, gradu- 
ally glabrescent with age above, pits of the scales 
remaining visible for a long time, subdensely lepi- 
dote beneath (scales rather small, round or nearly 
so, marginal zone narrow, entire to crenulate, 
centre thick, rusty-brown. - distinctly impressed), 
not rarely provided with a fine puberulence in the 
lower part of the blade especially when young, 
margin : revolute, (15-) 17-30 by (7-)9-13 mm, 
midrib sunk above, prominent beneath especially 
in its lower half, nerves 2-4 pairs, curved-ascend- 
ing and anastomosing, minutely depressed above, 
slightly raised beneath, not rarely : inconspicu- 
ous, reticulation 0; petiole very short (blade is 
decurrent to nearly the base), up to 1.5 mm. 
Flowers solitary or in twos (sometimes 3, rarely 4 
in an umbel). Flower buds ovoid, c. 10 by 6 mm. 
Outer perulae ovate to suborbicular, inner ones 
broadly obovate-spathulate, obtuse or minutely 
apiculate. lepidote and sericeous upwards dorsally, 
short-ciliate. Bracteolcs filiform, glabrous, up to 
6 mm. Pedicels rather slender, c. 10 (sometimes 
up to 15) mm, rather densely shortly patent- 
pubescent, laxly to subdensely lepidote. Calyx 
small, disk-shaped, 5-lobcd, lepidote and • hairy, 
lobes broad-ovate and obtuse, c. 0.5 mm. Corolla 
red, tubular in outlmc, slightly oblique, sub- 
campunulatc-dilutcd upwards, rather shortly 5- 
lobcd, 12-I4( 15) mm long in all, • shining and 
subdensely lepidote. moreover with some scattered 
hairs at the very base (but not hairy all over the 
lube) outside, glabrous inside, tube c. 10 mm, 
2.5-3 mm o at the base, 4 5 mm at the throat, 
lobes somewhat irregular, broadly clliptic-obovate 
or suborbicular, 3-4 by 2.5-4 mm. finely subdenti- 



culate-erose or entire at the apex. Stamens 10, 
as long as or slightly shorter than the corolla; 
filaments linear, glabrous; anthers subglobose, c. 
1 mm. Disk glabrous. Ovary obliquely cylindrical, 
gradually attenuate into the style, c. 2 by 1.5 mm, 
5-ribbed, densely lepidote; style glabrous, c. 8 
mm; stigma subcapitate, irregularly lobed. Capsule 
obliquely cylindrical, densely rusty-lepidote, 1-1.2 
(-1.5) by 0.25-0.35 cm. Seeds c. 2 mm including the 
narrow tail at both ends. 

Distr. Malesia: Philippines (Luzon, Albay 
Prov.: Mayon Volcano). 

Ecol. Rocky open slopes, 1330-2100 m, 
abundant. Fl.fr. May-June, Sept. 

Vern. Kutmii. 

Note. This is the "typical" form, i.e. the first 
one described of this rather variable species, 
collected again and again on Mayon Volcano, 
but not found elsewhere. Within the general aspect 
of R. quadrasiamim s. lat., this is a homogeneous, 
glabrescent, big-leaved form. 

CoPELAND /. stated that a special form of R. 
quadrasianum occurs on almost every mountain in 
the Philippines. I had to reduce his infraspecific 
taxa to a more reasonable number in order to 
avoid overlapping. 

2. var. davaoense (Copel./.) Sleum. Reinwardtia 
5 (1960) 65. — /?. quadrasianum Vidal f. davaoense 
Copel. /. Philip. J. Sc. 40 (1929) 143, t. 1, f. 9, 
10. — R. quadrasianum Vidal/. negrosense Copel. 

f.l.c. 142, t. l,f. 6. 

Treelet or shrub, sometimes epiphytic. Leaves 
rather broadly and shortly obovate, 15-20(-28) 
by (5-)6-8(-10) mm. Corolla 10-15(-16) mm 
long in all, subcampanulate-dilated distally, lepi- 
dote and at least laxly pubescent outside. Pedicels 
pubescent and - densely lepidote at anthesis. 
Ovary only lepidote. 

Distr. Malesia: Philippines (Mindanao, Leyte, 
Negros, S. Luzon). 

Ecol. Dense moss forest on summits. 1600- 
2440 m. F/./r. Jan.-Dec. 

Vern. Tongog, Bag., lasimancahoy, Negros. 

3. var. rosmarinifolium (Vidal) Copel. /. Philip. 
J. Sc. 40 (1929) 144. t. 1. f. 13-18; Sleum. Rein- 
wardtia 5 (I960) 65; Blumea 12 (1964) 92. — 
R. rosmarinifolium Vidal, Rev. PI. Vase. Filip. 
(1886) 172; Rendle, J. Bot. 34 (1896) 355; 
Merr. Publ. Gov. Lab. Philip, n. 29 (1905)43; 
Philip. J. Sc. 2 (1907) Bot. 292; ibid. 3 (1908) Bot. 
382; MiLi.Ais, Rhodod. (1917) 236; Merr. En. 
Philip. 3 (1923) 245; Miilais, Rhodod. scr. 2 
(1924) 225. — R. cuneifolium (non Siaim ) Rindle, 
J. Bot. 34 (1896) 355; MtRR. Publ. Gov. Lab. 
Philip. /;. 29 (1905) 43. R. quadrasianum WwM. 
var. intermedium MtRR. Philip. J. Sc. 3 (1908) Bot. 
382./)./).; En. Philip. 3 ( 1923) 244. /»./?. R. quadra- 
sianum Vidal /. halconensc Copel. / Philip. J. 
Sc. 40 (1929) 141. t. I, f. 5. — R. quadrasianum 
Vidal /. banaluioense Copel. f. I.e. 145, t. I, 
f. 24. — R. quadrasianum Vidal /. pulogense 
Copel./ I.e. 144. t. 1. f. 19-23. 28; H. J. Lam. 
Blumea 5 (1945) 575. p.p. 



496 



Flora Malesiana 



[ser. I, vol. 6* 



Shrub, (0.5-)l-2 m, rarely treelet up to 6 m. 
Leaves ± elongate-obovate, (10-) 12-20 (-2 2) by 
3-5 (rarely up to 6) mm. Corolla 10-15(-16) mm 
long in all, subcampanulate-dilated distally, red 
to orange, lepidote and at least laxly pubescent 
outside. Pedicels pubescent and -±, densely lepidote 
at anthesis. Ovary only lepidote. 

Distr. Malesia: Philippines (Mindoro, Biliran, 
Luzon). 

Ecol. Mossy (oak-)forest, also in secondary 
growth and open places, locally common, 1300- 
2400 m. Fl.fr. Jan.-Dec. 

V e r n . Bikbiki, sisisip, Ig. 

4. var. malindangense (Merr.) Copel. /. Philip. J. 
Sc. 40 (1929) 142, t. 1, f. 7, 8; Sleum. Reinwardtia 
5 (1960) 66. — R. malindangense Merr. Philip. J. 
Sc. 3 (1908) Bot. 256, 381; Millais, Rhodod. 
(1917) 206; Merr. En. Philip. 3 (1923) 244; 
MiLLAis, Rhodod. ser. 2 (1924) 183. 

Shrub, 2 m, or treelet up to 7 m, sometimes 
epiphytic. Leoveioblong-obovate, (1 5-)20-25(-30) 
by (5-)6-8(-10, rarely up to 13) mm; petiole 
2-3 mm. Corolla strictly cylindric, red, 20(-25 ) mm 
long in all, 4-5 mm 0, laxly pubescent and lepidote 
outside, lobes 5 by 3-4^ mm. Anthers 1 mm. 
Pedicels rather densely pubescent and laxly 
lepidote. Ovary only lepidote. 

Distr. Malesia: Philippines (Mindanao, Cami- 
guin). 

Ecol. Mossy forest, 1000-1600 m. Fl.fr. March- 
Sept. 

Vern. Malagas, Bag., mogd, Sub., tagum-tagiim, 
Mbo. 

5. var. marivelesense (Copel. /.) Sleum. Rein- 
wardtia 5 (1960) 66. — R. quadrasianurn Vidal/. 
marivelesense Copel. /. Philip. J. Sc. 40 (1929) 
141, t. l,f. 3, 4, 25-27. 

Mostly small tree. Leaves (rather broadly) 
obovate, (11-) 12-1 8 (-20) by 4-7 (-8) mm. 
Corolla c. 15 mm long in all, tubular, pubescent 
and lepidote outside. Pedicels pubescent, prac- 
tically not lepidote. Ovary only lepidote. 

Distr. Malesia: Philippines (Luzon, Mindoro). 

Ecol. Mossy forest, 1 200-2500 m. Fl.fr. March- 
Nov. 

6. var. intermedium Merr. Philip. J. Sc. 3 (1908) 
Bot. 382, s. str.; En. Philip. 3 (1923) 244; Copel./. 
Philip. J. Sc. 40 (1929) 143, t. 1, f. 11, 12, 29; 
Sleum. Reinwardtia 5 (1960) 66. 

Small tree or shrub. Leaves narrow-obovate, 
8-10(-ll) by 2-3(-4) mm. Corolla tubular below, 
subcampanulate distally, c. 10 cm long in all, 
pubescent and lepidote outside. Pedicels pubescent, 
practically elepidote. Ovary exclusively lepidote. 

Distr. Malesia: Philippines (Luzon). 

Ecol. Open exposed ridges, 2000-24jOO m. 

Vern. Kutmii, Ig., tongog. Bag. 

7. var. selebicum J. J. S. Bull. Jard. Bot. Btzg III, 
13 (1935) 443; Bot. Jahrb. 68 (1937) 199; Sleum. 
Reinwardtia 5 (1960) 67. — R. quadrasianurn 



ViDAL /. monodii H. J. Lam, Blumea 5 (1945) 
576. 

Shrub or treelet up to 4 m. Leaves ± broadly 
obovate, 6-14 by (3-)4-5(-6) mm. Corolla lepi- 
dote, but epilose outside, (10-)12-15 mm long in 
all, red or orange, sometimes tube yellow, limb 
orange. Pedicels laxly hairy and laxly to subdensely 
lepidote, sometimes exclusively lepidote. Ovary 
exclusively lepidote. 

Distr. Malesia: Central and SE. Celebes. 

Ecol. Light forest or open places, 2335-3000 m. 
Fl. June-Nov. 

8. var. cuneifolium (Stapf) Copel./. Philip. J. Sc. 
40 (1929) 145; Sleum. Reinwardtia 5 (1960)67; 
Blumea 12 (1963) 92; Jahrb. Deutsch. Rhodod. 
Ges. Bremen (1965) f . 6 — R. cuneifolium Stapf, 
Trans. Linn. Soc. H, Bot. 4 (1894) 198, t. 15, f.B, 
3; GiBBS, J. Linn. Soc. Bot. 42 (1914) 104; Millais, 
Rhodod. (1917) 150; Merr. En. Born. (1921) 
A6\,p.p.; Millais, Rhodod. ser. 2 (1924) 121. — 
R. quadrasianurn Vidal f. pulogense (non Copel./) 
H. J. Lam, Blumea 5 (1945) 575, p.p. 

Terrestrial, or sometimes epiphytic shrub or 
treelet. Leaves i elongate, rather narrow-obovate, 
6-16 by 2.5^(-5) mm. Pedicels laxly lepidote, 
nearly always epilose. Corolla glabrous or nearly 
so, red, pink, (pale) brick- or salmon-red, 10-15 
(-20) mm long in all. Ovary only lepidote. 

Distr. Malesia: North Borneo (Mt Kinabalu) 
and SW. Central Celebes (once found near Rante- 
pao, N of Makale). 

Ecol. On Mt Kinabalu common undershrub in 
mossy forest clearings, or in ridge or summit scrub 
forest, (1 525-) 1650-2865 m, in Central Celebes 
in grass and fern wilderness at 1200 m. Fl. Jan.- 
Dec. 

9. var. villosum J. J. S. [ex Merr. En. Born. (1921) 
463, nom. nud.] Bull. Jard. Bot. Btzg III, 13 (1935) 
444; H. J. Lam, Blumea 5 (1945) 576, incl.f lutea 
H. J. Lam et f rubra H. J. Lam, I.e.; Sleum. 
Reinwardtia 5 (1960) 67; Blumea 11 (1961) 114; 
ibid. 12 (1963) 92. — R. cuneifolium {non Stapf) 
RiDL. J. Str. Br. R. As. Soc. n. 63 (1912) 61. — 
R. cuneifolium Stapf var. subspathulatum {non 
RjDL.) Merr. Sarawak Mus. J. 3 (1928) 541. 

Shrub, apparently epiphytic in general. Leaves 
narrow-obovate, (5-)9-16 by 3-5 mm. Corolla 
pubescent (sometimes rather sparingly so) and 
lepidote, red or yellow, c. 15 mm long in all. 
Pedicels lepidote and pubescent. Ovary lepidote 
and rather densely pilose to nearly villous. 

Distr. Malesia: Borneo. 

Ecol. Both in open mossy forest (rather 
common on Mt Dulit and Mt Kinabalu) and in 
low scrubby "kerangas" vegetation on bare sand- 
stone rocks (as on summit of Mt Tanggoi), and in 
elfin woodland on rugged hill crest (Mt Pagon, 
1675 m), (820-)1350-1680 (sometimes up to 1900) 
m. Fl. Jan.-Dec. 

10. var. angustissimum Sleum. Blumea 12 (1963) 
92. — R. ericoides {non Low ex Hook./) Burtt, 



Jan. 1966] 



Ericaceae (Sleumer) 



497 



Proc. R. Soc, B, 161 (1964) 83, in text. Epiphytic 
shrublet; branchlets short-pubescent. Leaves 
narrow-oblong-obovate. densely arranged especi- 
ally towards the top of the young shoots, c. 10 
by 1-1.5 mm. Pedicels slender, c. 5 mm, hairy. 
Calyx lobes very short, ciliate, tube hairy. 
Corolla tubular, slightly dilated distally, red, 
densely covered with roundish scales, sparsely so 
by longish white hairs, c. 10 by 3-4 mm. Stamens 
exserted; anthers 0.5 mm. Ovary subvillous. 

Distr. Malesia: Borneo (Sarawak: Mt Mulu), 
in submontane forest, c. 1300 m. Fl. June. 



11. var. borneense J. J. S. [ex Merr. En. Born. 
(1921) 462, nom. nud.] Bull. Jard. Bot. Btzg III, 
13 (1935) 444; Sleum. Reinwardtia 5 (1960) 67; 
Blumea 12 (1963) 92. 

Shrub, probably epiphytic in general. Leaves ± 
broad-obovate, (1 1-)16-22 by (5-)7-10mm. Corol- 
la red, hairy and lepidote outside, c. 15 mm long 
in all. Pedicels hairy and laxly lepidote. Ovary 
densely pubescent and lepidote. 

Distr. Malesia: Borneo. 

Ecol. In primary forest, common on Mt Kemul, 
1800 m. Fl. Oct. 



2. Subsection Siphonovireya 

Sleum. Reinwardtia 5 (1960) 68. — Rhododendron sect. Hadranthe Schltr, Bot. Jahrb. 55 
(1918) 157,/7./7.— Fig. 9-10. 

Scales disk-shaped, entire or nearly so (subcrenate), their marginal zone narrower and thinner 
than the mostly thick centre. Corolla salver-shaped, tube elongate and ± narrow, straight or 
slightly curved, lobes equalling 1/4 or less of the total length of the corolla, spreading ± hori- 
zontally. 

Distr. Malesia: limited to New Guinea. Fig. 16. 

KEY TO THE SPECIES 

1. Leaves distinctly prominent-reticulate underneath 26. R. habbemae 

1. Leaves obscurely or not reticulate underneath. 
2. Style lepidote and patent-hairy in the lower third. 

3. Ovary exclusively lepidote 27. R. cinchoniflorum 

3. Ovary lepidote and hairy. 

4. Corolla tube 2-2.5 cm; lobes much reflexed. Style widely exserted 28. R. protandrum 

4. Corolla tube c. 3 cm; lobes erecto-patent. Style included 29. R. inundatum 

2. Style lepidote to various degree, not hairy at all. 
5. Corolla tube strictly cylindric, or but slightly dilated in the upper part. 
6. Capsule c. 10 by 0.5-0.6 cm. Petiole (9-)l l-16(-22) mm. Corolla tube 7-8(-8.5) cm 

30. R. agathodaemonis 
6. Capsule (2.5-)3-4(-5) by 0.4 cm. Petiole (3-)5-8 (rarely up to 12) mm. Corolla tube 4-5.5 (rarely 

up to 7) cm 31. R. herzogii 

5. Corolla tube gradually dilated from the base upwards 32. R. incommodum 



26. Rhododendron habbemae Koord. Nova 
Guinea 8 (1912) 877 Vliahbemai'); J. J. S. ibid. 
12 (1914) 142; ibid. 12 (1917) 510; Sleum. Rein- 
wardtia 5 (1960) 68. — R. bodenii Wernh. Trans. 
Linn. Soc. IF, Bot. 9 (1916) 94. 

Shrub. Branchlets smooth, tips exclusively laxly 
lepidote. Leaves in 4-9 cm spaced, 3-5-merous 
pseudowhorls, obovate or obovate-oblong or 
obovatc-clliptic, apex broadly attenuate, mostly 
;r obtuse, often apiculatc by a small protruding 
gland, base broadly cuneate to truncate-rounded or 
subcordatc, coriaceous, rigid, shining and h 
olivaceous above, dull and paler beneath, laxly 
to subdcnscly covered with small, rounded 

scales, which arc ;: immersed on both faces in 
fully mature leaves, 3.5-6 by 1.5-3 cm, midrib 
strongly impressed above, much raised beneath, 
nerves 6-8(-!0) pairs, rather straight, inarching 
near the edge, very slightly impressed above, 
prominent beneath, reticulation rather inconspicu- 
ous or very slightly impressed above, dense and 
manifestly raised beneath; petiole grooved above, 
4-7 by c. 1.5 mm. Umbeh 5-8-flowercd. Pcruiac 



ovate to obovate, outer ones :b acuminate, inner 
ones obtuse, i lepidote outside, glabrous inside, 
up to 1.5 cm. Bracteoles filiform to linear-spathu- 
late, glabrous, 1-1.5 cm. Pedicels rather slender, 
±densely lepidote, (0.6-) 0.9-1 .5 cm. Calyx small, 
shortly and rather irregularly 5-dentate. Corolla 
salver-shaped, white, tube narrow-cylindric, 
straight or slightly curved, its base only somewhat 
dilated (5-6 mm) and 5-pouched, rather densely 
covered with small rounded scales outside as are 
the lobes, glabrous inside, (3.3-)4-5(-5.5) cm, 
lobes obovate, obtuse, 1-1.4 by 0.6-1 cm. Stamens 
10. unequal; filaments linear, laxly to very laxly 
hairy, 3.5-4 cm; anthers oblong, 3 by I mm. Disk 
glabrous or almost so. Ovary subcylindric, 5- 
ribbcd. very densely lepidote, 6-7 by 1.5 mm; 
style slender, 2.7-3.5 cm, mostly somewhat ex- 
serted from the throat, clad with short spreading 
hairs and scales at the base, exclusively hairy 
above, the upper part glabrous; stigma turbinate. 
Distr. Malesia: New CJuinea (Main Range 
from Mt Carstcnsz to the Hell wig Mts and Mt 
Goliath). 



498 



Flora Malesiana 



[ser. I, vol. 6* 



Ecol. Both in forest on wet mossy ground and 
on humus- and moss-covered exposed rocks, 
2000-3450 m. Fl. Jan. -Dec. 

27. Rhododendron cinchoniflorum Sleum. Rein- 
wardtia 5 (1960) 68. — Fig. 9. 

Shrub. Branchlets rather slender, terete, tips 
lepidote; intcrnodes 2-8 cm. Leaves in 4-6-merous 
pseudowhorls at the end of the upper internodes, 
ovate-elliptic or mostly elliptic, apex shortly i 
obtusely acuminate, not rarely apiculate by a 
small gland, base subtruncate-rounded to slightly 
cordate, coriaceous, subdensely and ± cadu- 




Fig. 9. Rhododendron cinchoniflorum Sleum. a. 

Habit, X 2/3, b. stamen, x 2, c. ovary and style, 

X 2 {a-c Eyma 5264). 

cously lepidote above, densely and persistently so 
beneath (scales nearly round, ± wavy at the mar- 
gin, very distinctly impressed, the pits becoming 
blackish with age), (1.6-)2-2.5(-3) by 1-1.6 cm, 
midrib impressed above, strongly prominent 
beneath, nerves 3-4 pairs, very slightly impressed 
above in fully mature leaves, somewhat prominent 
in younger ones, generally not well visible, 
especially beneath, reticulation 0; petiole lepidote, 
1.5-3 by 1-1.5 mm. Umbels 4— 6(-7)-flowered. 
Outer perulae ovate-acute, inner ones obovate, 
lepidote dorsally along the middle line, up to 1 by 
0.7 cm. Bracteoles linear, c. 6 by 1 mm. Pedicels 



rather slender, densely rufous-Iepidote, 5-8 mm. 
Calyx disk-like, c. 2.5 mm 0, limb spreading or 
reflexed at anthesis, lepidote outside, glabrous 
inside, margin undulate and shortly 5-lobed. 
Corolla salver-shaped, colour not known (probably 
white), tube nearly straight, somewhat 5-saccate 
at the very base, very densely lepidote outside, 
glabrous inside, 2.5-3 cm by c. 2.5 mm, lobes ~ 
spreading horizontally, obovate-eliiptic, 1-1.2 by 
0.8-0.9 cm, laxly lepidote outside, glabrous inside. 
Stamens 10, unequal, not or slightly exserted 
from the throat; filaments filiform, laxly hairy; 
anthers broadly oblong, 2-2.5 mm. Disk low, 
i densely white-hairy. Ovary cylindric-conical, 
densely exclusively lepidote, c. 4 by 2 mm, 
abruptly narrowed to the style, slightly 5-ribbed; 
style c. 2.3 cm, lepidote and shortly patent-hairy 
in the lower third, gradually glabrescent upwards; 
stigma shortly turbinate, minutely crenulate. 
Distr. Malesia: W. New Guinea (near the 
Wissel Lakes), 2400 m. Once found. Fl. Oct. 

28. Rhododendron protandrum Sleum. Blumea 1 1 
(1961) 114. 

Shrub c. 2.5 m, divaricately branched. Branch- 
lets rather slender, juvenile parts densely lepidote 
and c. 2 mm 0; internodes (2-)3-6(-8) cm. 
Leaves in 4-5-merous pseudowhorls at the upper 

2 or 3 nodes, elliptic, apex — broadly subacumi- 
nate, or sometimes obtusely apiculate, base broadly 
attenuate or subcuneate, coriaceous, subdensely 
lepidote initially, glabrescent above with age, i 
persistently lepidote beneath (scales small, round- 
ish, marginal zone narrow, centre thick, yellowish, 
slightly impressed), very narrowly marginate, 3-5 
by (1.6-)2-3 cm, midrib narrowly immersed above, 
as thick as the petiole and much prominent in the 
lower part beneath, nerves 4(-6) spreading, rather 
straight pairs, faintly impressed above, very 
slightly raised beneath, or not rarely :r obsolete 
on both faces, no reticulation; petiole grooved 
above, 4-6 by c. 1 mm. Umbels 5-8(-10)-flowered. 
Perulae membranous, glabrous dorsally, glandular 
fimbriate apically, outer ones ovate, apiculate or 
obtuse, inner ones obovate-oblong to oblong, up 
to 1 by 0.8 cm. Bracteoles subspathulate-linear, 
glabrous, up to 8 by 1 mm. Pedicels slender, 
5-7(-10) mm, subdensely lepidote. Calyx sub- 
obliquely patellar, c. 2.5 mm 0, shortly and 
irregularly 5-lobed. Corolla tubular, ± horizon- 
tally expanded at the lobes, pure white or greenish 
white, or pale pink at the tube and white at the 
lobes, densely lepidote outside, set with short 
retrorse hairs in the lower third, glabrous for the 
rest inside, tube straight or slightly curved, c. 
2.5 cm, 2-2.5 mm except the slightly dilated 
very base, lobes subspathulate-oblong, 5-6 by c. 

3 mm, which become + reflexed in full anthesis. 
Stamens 10, much exserted from the corolla tube 
early at the anthesis (when the style is still in- 
cluded in the corolla tube), erect, ri recurved 
with the upper part of the filaments towards the 
end o*" the anthesis (when the style becomes much 
exserted); filaments almost filiform, white, laxly 
or more densely hairy in the lower 1/4; anthers 



Jan. 1966] 



Ericaceae (Sleumer) 



499 



oblong, base obtuse, 1.8 by 0.5 mm. Disk slightly 
prominent, laxly erect-hairy at the upper margin 
only. Ovary cylindric. c. 5 by 1.5 mm, subdensely 
subappressedly short-hairy and lepidote. tapering 
to the slender style, the latter lepidote to almost 
the top and laxly hairy in the lowest 10 mm; 
stigma subcapitate, very shortly 5-lobed. Sub- 
mature capsule narrow-fusiform, reddish, laxly 
lepidote and hairy, 2.5-3 cm by 2.5-3 mm, 
rather deeply 5-furrowed lengthwise. 

Distr. Malesia: New Guinea (Sibil R. Valley). 

Ecol. in shrub vegetation on poor, impervious, 
white clay, 1260 m, common. Fl. June; /r. Aug. 

29. Rhododendron inundatum Sleum. Blumea 12 
(1963)92. 

Terrestrial shrub, c. 1 m; branchlets reddish, 
striate, slightly pruinose when dry, variously 
obtusely compressed, laxly and in caducously 
lepidote; the 2 ultimate internodes 6-12 cm by 
3-4 mm. Leaves erecto-patent, in 4-5-merous 
pseudowhorls at the upper 3 nodes, elliptic to 
elliptic-obovate, apex very broadly attenuate or 
mostly rounded, sometimes apiculate, base broadly 
cuneate to the petiole, coriaceous, green above, 
yellowish green below, early glabrescent above, 
still subdensely lepidote with age (scales small, 
roundish, distant from each other for about their 
diameter, marginal zone thin, narrow, centre 
small, much impressed), slightly revolute in dry 
specimens, 3.5-5.5 by (1.5-)2-3(-3.3) cm, midrib 
narrowly immersed above, as thick as the petiole 
at the base, nerves 6-7 pairs, with some additional 
less conspicuous ones between, rather straight, 
obscurely anastomosing before the edge, slightly 
impressed above, a little prominent or indistinct 
below, no reticulation; petiole 6-9 by 1-1.5 mm. 
Umbels :r 4-flowered. Outer perulae ovate- 
acuminate, submucronulate by a protracted apex, 
densely set with thickish scales and very short 
hairlets at base and the middle outside, otherwise 
glabrous, inner ones : spathulate, thinner, edge 
minutely lacerate, up to 1 by 0.6 cm. Bracteoles 
linear, sparsely lepidote, not hairy, c. 1.2 cm. 
Pedicels pinkish, • densely lepidote, epilose, 
1-1.2 cm by c. 0.5 mm. Calyx a little oblique, 
patellar, obscurely lobed, c. 2.5 mm 0. Corolla 
tubular below, somewhat expanded at the lobes, 
white, 4-4.3 cm long in all, subdensely lepidote 
except the marginal parts of the lobes outside, 
laxly hairy at the tube inside, tube rather straight, 

: 3 cm, base 2.5-3 mm o, gradually widened to 
5 mm at the throat, lobes erecto-patent, obovatc- 
spathulatc, r. I by 0.7-0.9 cm. Slanicns 10, 
reaching halfway the corolla lobes; filaments 
narrow-linear and densely patently white-hairy 
in the lower 3/4, filiform and glabrous upwards; 
anthers oblong, base obtuse, 2 mm. Disk hardly 
prominent, glabrous. Ovary subcylindric, densely 
lepidote and laxly patently white-hairy, 4-5 by c. 

1.5 mm, continuous with the style, the latter 
rather slender, as long as the corolla tube, densely 
lepidote and laxly patently hairy in the lower, 
exclusively densely lepidote in the upper half; 
stigma subcapitate. 



Distr. Malesia: New Guinea (Balim R. Valley 
near Wiligimaan), once found. 

Ecol. Temporarily inundated grassplain on 
peaty soil, at c. 1650 m, locally common. Fl. June. 

Vern. Tiimuk, Dani. 

30. Rhododendron agathodaemonis J. J. S. in Fedde, 
Rep. 12 (1913) 209; Nova Guinea 12 (1914) 142, 
t. 37 A; ibid. 12 (1917) 510; Sleum. Reinwardtia 5 
(1960) 69. — R. hellwigii {non Warb.) Koord. 
Nova Guinea 8 (1912) 878, t. 153. 

Shrub or small, compact tree, 1.2-6 m. Branch- 
lets terete, tips laxly to densely rusty-lepidote. 
Leaves in 7-20 cm spaced, 4-5-merous pseudo- 
whorls, elliptic to obovate-elliptic, apex broad- 
acuminate to obtuse, sometimes apiculate by a 
small gland or a little retuse, base broadly attenuate 
into the petiole or mostly rounded-obtuse, coriace- 
ous, rather stiff, entire, somewhat revolute to- 
wards the base, sparsely to subdensely minutely 
lepidote on both surfaces especially beneath 
(scales round, entire or shallowly undulate at the 
marginal zone, centre ± impressed), (3.5-)5-7 
(-7.5) by (2.5-)3.5^.8 cm, midrib slightly im- 
pressed above, distinctly raised though somewhat 
flattened beneath, nerves 6-8(-10) pairs, rather 
straight, anastomosing w'th each other and the 
primary veins near the edge, very slightly im- 
pressed above, a little raised or inconspicuous 
beneath, reticulation obscure; petiole somewhat 
flattened, sparsely lepidote, (9-)l l-l6(-22) by 
1.5-2 mm, grooved above. Umbels showy, (6-) 
8-10(-l2)-flowered. Perulae ovate to obovate, 
lepidote dorsally, c. 1 .5 by I cm. Bracteoles filiform 
below, linear-spathulate upwards, laxly lepidote, 
c. 12 mm long, I mm wide apically. Pedicels 
rather stoutish, 6-15(-18) mm, i densely lepidote. 
Calyx very small, annular, irregularly denticulate. 
Corolla salver-shaped, with a scent of carnations, 
pure white or somewhat rose tinged at the tube, 
the latter slightly curved, cylindric at least below, 
sometimes somewhat dilated in the upper half, 
7-8(-8.5) cm long, c. 3 mm at the base, 4-6.5 mm 
in the middle and the upper half, sparsely to sub- 
densely lepidote outside, laxly hairy and not rarely 
sparingly lepidote in the lower half inside, lobes 
rounded to obovate, spreading in a right angle, 
1-1.5 by 0.8-1.2 cm, laxly lepidote or mostly 
glabrous outside, glabrous inside, Stamens 10, 
slightly or not cxsertcd from the throat; filaments 
very slender, subdensely (rarely more laxly) set 
with spreading hairs in the lower half, glabrous 
above; anthers orange, oblong, thccac apiculate at 
base, 3-4 by 0.8 mm. Disk glabi ous. Oranelongate- 
conical, densely lepidote, 7-10 by 2 mm; style 
slender, slightly exscrte'd, rather densely to laxly 
lepidote almost to the top, epilose; stigma short- 
turbinate, crenulate. Capsule red-brown, cylindric, 
attenuate at both ends, obtusely 5-angled, c. 10 
by 0.5-0.6 cm. 

Distr. Malesia: New Guinea (Mts llubrecht 
and Wichmann on the southern, and near Habbe- 
ma Lake on the northern side of the Orange Mts). 

Ecol. Moss forest, in damp grt)und or swampy 
open places. 3050-3450 m. /•/. leb., Aug.. Nov 



500 



Flora Malesiana 



[ser. I, vol. 6^ 



31. Rhododendron herzogii Warb. Bot. Jahrb. 16 
(1892) 15, 25; K. Sch. & Laut. F1. Schutzgeb. 
(1901) 487; Schltr, Bot. Jahrb. 55 (1918) 160, 
f. 6; Sleum. Reinwardtia 5 (1960)69; Blumea 11 
(1961) 115; ibid. 12 (1963) 93; Jahrb. Rhodod. 
Ges. Bremen (1963) f. 13, 18. — R. carringtoniae 
{noil F. V. M.) Lane-Poole, For. Res. Papua (1925) 
130; White & Francis, Proc. R. Soc. Queensl. 39 
(1928) 68; Stonor, Rhodod. Yearbook 6 (1951) 
49, f. 50. — /?. agathodaemonis (non J. J. S. 1913) 
J. J. S. Nova Guinea 18 (1936) 99. — Fig. 10. 

Erect shrub, (0.5-)l-2 m. Branchlets terete, 
subdensely lepidote at the ultimate internodes 
(3-13 cm), glabrescent and pruinose below. Leaves 
in 4-6-merous pseudowhorls at the upper nodes, 




Fig. 10. Rhododendron herzogii Warb. Edie Creek 

near Wau, E. New Guinea, 1850 m (Sleumer 

4129) (Sleumer, 1961). 

elliptic or obovate-elliptic, sometimes nearly 
obovate or elliptic-oblong, apex broadly ± 
obtusely acuminate, sometimes rather blunt, base 
broadly cuneate to almost rounded, coriaceous, 
dark to medium green above, paler to yellowish 
green beneath, slightly revolute towards the base, 
subdensely lepidote when young, though early 
glabrescent above, rather densely and ± persist- 
ently impressed-lepidote beneath (scales roundish, 
pale yellow, their marginal zone narrow and 
shallowly irregularly crenate), (2.5-)4-6(-9) by 



(1.5-)2-4(-5) cm, midrib slightly grooved above, 
robust and prominent beneath, nerves (6-)8-10 
(rarely up to 12) pairs, slightly curved, ± parallel 
to each other, rather obscurely anastomosing, 
faintly impressed above, raised beneath, reticula- 
tion obscure; petiole semiterete, grooved above, 
lepidote, (3-)5-8 (rarely up to 12) by 1.5-2 mm. 
Umbels 5-IO-flowered. Flower buds obovoid- 
oblong, c. 2 by 1 cm, almost whitish green when 
fresh. Ferulae ovate to subobovate, thin, densely 
lepidote dorsally except the glabrous marginal 
part, scales almost transparent-silvery in fresh 
specimens, up to 1.5 by 1 cm. Bracteoles narrow- 
linear, up to 3 cm. Pedicels rather stoutish, 
densely lepidote, (4-)6-8 mm. Calyx very small, 
oblique, disk-shaped, subentire. Corolla salver- 
shaped, pure white, or slightly pinkish at the tube 
externally, fragrant, tube straight or very slightly 
curved, 4-5.5 (rarely up to 7) cm long, c. 2.5 mm 
at base and middle, dilated upwards to c. 3.5 mm, 
densely lepidote outside, rather densely hairy and 
sparingly or not lepidote in the lower 2/3 of the 
inside, lobes spreading, orbicular to elliptic- 
obovate, 8-9 by 5-6 mm, i lepidote outside, 
glabrous inside. Stamens 10, somewhat exserted at 
least in part beyond the recurved corolla lobes; 
filaments white, very slender, ± densely and 
patently hairy in the lower 2/3 only; anthers orange, 
oblong, 3-3. 5 (-4) by 0.8 mm. Disk very laxly 
hairy or glabrous. Ovary narrowly cylindric, very 
densely lepidote, 6-7 by 1.5 mm, rather abruptly 
tapering to the style, the latter mostly somewhat 
exserted from the throat, densely lepidote below, 
less so upwards, sometimes glabrous distally; 
stigma green, short-obconical. Capsule reddish, 
cylindric, rather straight, lepidote, (2.5-)3-4(-5) 
by 0.4-0.5 cm. 

Distr. Malesia: New Guinea (Main Range 
from the Wissel Lakes to the Owen Stanley Range, 
and on Mt Saruwaged). 

Ecol. Terrestrial, more rarely epiphytic, on 
earthen cliff faces, landslides, steep eroded banks 
or slopes, also in mountain forest or heath-scrub, 
on infertile sandy or clayey soil, not rarely found 
in (fern) regrowth, as coloniser in sluiced areas or 
in gullies of old garden land, (1530-) 1600-2500 m. 
Fl. fr. Jan.-Dec. 

Vern. Lon, rohn, Enga, kibimuma, Chimbu: 
Masul, wumbambegon, Wahgi: Minj, mimbamb, 
Hagen: Togoba, ron^igehpa, Mairi: Watabung. 

32. Rhododendron incommodum Sleum. Reinward- 
tia 5 (1960) 70; Blumea 11 (1961) 115; ibid. 12 
(1963) 93. 

Shrub, c. 1 m. Branchlets rather slender, sub- 
terete, lepidote only at the ultimate internodes 
(0.5-6 cm). Leaves in 3-4-merous pseudowhorls 
at the upper nodes, elliptic-oblong or mostly 
elliptic, apex broadly obtusely acuminate, base 
cuneately attenuate to the petiole, coriaceous, 
laxly lepidote initially, finally glabrescent above, 
subdensely and ± persistently lepidote beneath 
(scales small, roundish, entire or nearly so, 
yellowish, ± impressed), slightly revolute towards 
the base, 5-7.5 by 2.5-4 cm, midrib a little impress- 



Jan. 1966] Ericaceae (Sleumer) 501 

ed above, robust and very prominent in the lower obovate, — spreading, very laxly lepidote outside, 

part, gradually less so upwards and fading dis- glabrous inside, 1-1.5 by 0.5-0.7 cm. Stamens 10, 

tally beneath, nerves 4-6 pairs in an acute angle well exserted from the throat; filaments filiform, 

with the midrib, obscurely anastomosing, slightly laxly pubescent especially in their lower part, 

impressed above, respectively raised beneath, and glabrous above, or glabrous at all; anthers 

reticulation obscure; petiole subterete, rugose, obovate-oblong, 2-3 by 1 mm, base obtuse. Disk 

4-20 by C-. 1.5 mm. Umbels — 8-flovvered. Bracte- glabrous. Ovary elongate-conical, very densely 

oles filiform, very laxly lepidote. Pedicels rather lepidote, c. 5 by 1.7 mm, gradually tapering to 

slender, subdensely covered with pale yellow the style, which is laxly lepidote to the top and 

scales, c. 10 by 0.5 mm. Calyx subobliquely disk- attains 5 cm; stigma shortly turbinate, rather 

shaped, wavy, very shortly 5-lobed, c. 2.5 mm small. 

0. Corolla tubular, 5-5.5 cm long in all, red, tube Distr. Malesia: New Guinea (Ogeramnang in 
straight or slightly curved, 3.5-4 cm, gradually the Sattelberg-Saruwaged area, and Purosa in the 
dilated from c. 3 mm o at the base to c. 1 cm at Okapa area. Eastern Highlands). 

the throat, laxly to subdensely clad with small Ecol. Locally common, high epiphyte in mixed 

yellowish scales outside, very laxly hairy and rain-forest, 1830-1950 m. Fl. Sept., Dec. 
sparingly lepidote inside, lobes obovate or oblong- 

3. Subsection Phaeovireya 

Sleum. Bot. Jahrb. 74 (1949) 539; Reinwardtia 5 (1960) 71. — Rhododendron sect. Schistanthe 
ScHLTR, Bot. Jahrb. 55 (1917) 140, p.p. — Rhododendron sect. Zygomorphanthe Schltr, ibid. 
55 (1918) 145, p.p. — Rhododendron sect. Hadranthe Schltr, I.e. 157, p.p. — Fig. 11—15,31. 
Scales — dendroid, each on top of a distinct and ± persistent epidermal tubercle, thus both 
surfaces or at least the undersurface of the leaves densely tubercled and scabrid or rough to the 
touch in later glabrescent stages; marginal zone usually wide, deeply, often narrowly incised 
(then becoming similar to a stellate hair). Corolla various, mostly shortly tubular and ± 
zygomorphous, rarely elongate and funnel- or salver-shaped. 

Distr. Malesia: Central Celebes and New Guinea. Fig. 16. 

KEY TO THE SPECIES 

1. Ovary exclusively and densely stellate-lepidote, i.e. no simple hairs present except possibly at the 
very apex of the ovary and on the style. 

2. Style up to 2 mm. (Leaves 0.8-1.6 by 0.4-0.7 cm, very much revolute). 
3. Anthers dehiscent by anticous pores. Style columnar, 1.5-1.6 mm. 
4. Leavesspirally arranged, equidistant. Corolla c. 1.3 cm long in all, lobes 4-5 mm o. . .33. R. eymae 
4. Leaves in distant pseudowhorls. Corolla (l.5)-l.7-2.2 cm long in all, lobes 7-9 mm o. 

34. R. revolutum 
3. Anthers dehiscent by posticous pores. Style obconical, I mm. Leaves in pseudowhorls. 

35. R. extrorsum 
2. Style at least 5 mm. 

5. Style thick, c. 2 mm o. Ovary 4-5 mm 0. Leaves large, (7-)10-16 by (3.5-)6-IO cm. 
6. Corolla deep blood red, 6.5-7(-8) cm. Style 3-3.5(-4) cm 36. R. hellwigii 

6. Corolla white or cream, may be with a rose tinge, or pink with a reddish throat, (1 l-)l 1.5-12.5 cm. 
Style 5.5-6.5 cm 37. R. supcrbum 

5. Style slender, up to 1 mm 0. Ovary up to 2.5 mm 0. Leaves small or mostly mcdium-si/cd. 

7. Leaves linear, lanceolate or ovate-lanceolate, 0.3-0.9 (-1.3, very rarely up to 1.6) cm wide. 

8. Corolla glabrous outside 38. R, prainianum 

8. Corolla lepidote and/or hairy outside. 

9. Corolla lepidote and hairy outside 39. R. steliigerum 

9. Corolla lepidote or hairy outside. 
10. Corolla exclusively lepidote outside. 

11. Pedicels up to 0.5 cm. Style 0.6 cm 40. R. ncriifniium 

II. Pedicels (0.7-)l-l.3cm. Style 1.7-3 cm. 
12. Leaves linear, strongly revolute, (3-)3.5-5.5 cm by 2-3(-4) mm . . .41. R. huoKlandii 
12. Leaves lanceolate to ovate-lanceolate, slightly revolute, 3-7 by (0.3-)0.6-l(-l .3, very 

rarely up to 1.6) cm 42. R. rariirii 

10. Corolla exclusively whitish hairy outside 43. R. dtlicatuluin 

7. Leaves of various shape besides linear or lanceolate, I cm wide only in the narrowest leaves in 
the same specimen, and at least in part, or mostly all of them much wider. 
13. Style exclusively laxly : patently short-hairy at least at the base, not lepidote at all. 



502 Flora Malesiana [ser. I, vol. 6* 

14. Filaments glabrous or practically so 47. R. dielsianum var. stylotrichum 

14. Filaments hairy to various degree. 

15. Corolla quite glabrous outside from the beginning 44. R. bryophilum 

15. Corolla laxly to subdensely lepidote outside at least initially, may be glabrescent in later stages. 

16. Corolla 3.8-4.5 cm long in all. Anthers 3 mm 45. R. tuberculiferum 

16. Corolla 2.6-3 cm long in all. Anthers 2 mm 46. R. asperrimum 

13. Style bearing no (or practically no) hairs, otherwise whether or not stcllate-lepidote. 
17. Style stellate-lepidote only at the very base, or quite glabrous. 
18. Corolla sparsely to subdensely lepidote outside. 
19. Filaments glabrous or practically so 47. R. dielsianum var. dielsianum 

19. Filaments laxly papillose-hairy 46. R. asperrimum 

18. Corolla completely glabrous outside. 

20. Corolla about halfway 5-lobed, c. 2.5 cm 48. R. stolleanum 

20. Corolla 5-lobed to 1/3, 3.5-3.8 cm 49. R. psilanthum 

17. Style stellate-lepidote at least in the lower third. 

21. Filaments densely hairy below. Disk ± densely hairy 50. R. phaeochitum 

21. Filaments very sparsely hairy or glabrous. Disk glabrous. 
22. Leaves coriaceous, rather stiff, very densely stellate-lepidote ('tomentose') initially (scales 
persistent a rather long time), ± distinctly revolute at the margin. . 51. R. beyerinckianum 
22. Leaves subcoriaceous, less densely stellate-lepidote initially (scales ± fugacious), but 
slightly or not revolute at the margin. 

23. Leaves ovate or oblong-ovate 52. R. leptanthum 

23. Leaves (broadly) elliptic to oblong-elliptic, rarely obovate-elliptic ... 53. R. warlanum 
1, Ovary all over densely set with simple hairs, or distinctly hairy and lepidote to various degree. 
24. Corolla (8-)9-14(-16) cm long in all. 
25. Style glabrous except for some hairs and/or scales at the very base. 

26. Ovary subcylindric, densely covered with ± stiff hairs, the hairs completely hiding the scales. 
Corolla glabrous outside 54. R. gardenia 

26. Ovary ± bottle-shaped, densely covered equally with hairs and scales mixed on the lower (seed- 
bearing), and exclusively lepidote in the upper (constricted) part and the very base of the style. 
Corolla laxly lepidote outside 55. R. thaumasianthum 

25. Style lepidote and/or hairy at least in the lower half. 

27. Style predominantly hairy, and laxly stellate-lepidote in about the lower half, gradually more 
lepidote and less hairy above for about 1/4 of the total length, exclusively stellate-lepidote for 
some mm above, and glabrous in the uppermost part. Flowers 6-7-merous. 

28. Anthers (8-) 10-1 1 (-16) mm 56. R. konori 

28. Anthers 6(-7) mm 57. R. phaeopeplum 

27. Style densely stellate-lepidote and epilose in the lower 2/3, glabrous above. Flowers 5-merous. 

Anthers 6 mm 58. R. opulentum 

24. Corolla up to 7 cm long in all. 
29. Leaves sessile or subsessile (petioles rarely up to 4 mm). 
30. Style glabrous except for some spreading hairs at the base or in the lower 1/4. Ovary predomi- 
nantly lepidote, the hairs scattered between the scales. 
31. Corolla 3.8-^.5 cm; anthers 3 mm 45. R. tuberculiferum 

31. Corolla 2.6-3 cm; anthers 2 mm 46. R. asperrimum 

30. Style hairy and lepidote at least halfway. Ovary predominantly hairy, the tomentum ± covering 

the scales. 

32. Corolla 2.5-2.8 cm. Leaves 1.6^ by 1.2-1.7 cm. 

33. Leaves ovate, base slightly cordate. Umbels 3-flowered. Corolla outside laxly lepidote, and 
sometimes very sparsely hairy too, inside laxly hairy 59. R. spondylophyllum 

33. Leaves elliptic, base ± rounded. Flowers solitary. Corolla outside sparsely hairy, inside glabrous. 

60. R. melantherum 
32. Corolla generally at least 4 cm long. Leaves (3-)3.5-7.5 by (1.6-)2.4-4.5 cm. 

34. Leaves ovate or elliptic or suborbicular, sometimes obovate-elliptic. (Corolla 4-4.5 cm). 
35. Corolla broadly tubular, ± zygomorphic; lobes ± erecto-patent, 1-1.2(-1.5) by 0.8-1 cm. 

61. R. cyatheicolum 
35. Corolla funnel-shaped, ± regular; lobes widely spreading, 1.5-1.9 cm . .62. R. asperum 

34. Leaves oblong-elliptic. (Corolla (4.5-)5-5.5 cm) 63. R. rhodochroum 

29. Leaves distinctly petioled. 

36. Corolla 5-lobed ± halfway, 2-2.3 cm long 64. R. gilliardii 

36. Corolla 5-7-lobed up to 1/3 of its total length. 

37. Leaves deeply impressed-reticulate above, sharply prominently so beneath, i.e. distinctly 
bullate. 

38. Leaves (8-)10-18(-22) by 4.5-12(-13) cm 65. R. bullifolium 

38. Leaves 5-8 by 3-5 cm 66. R. solitarium 



Jan. 1966] 



Ericaceae (Sleumer) 



503 



37. Leaves whether or not slightly impressed-reticulate above, resp. prominently so, i.e. not properly 
buUate. 
39. Corolla ± densely and ± persistently lepidote outside. 
40. Corolla 2.5-3 cm long. 
41. Leaves ovate- or subovate-oblong, coriaceous 67. R. schoddei 

41. Leaves elliptic- to obovate-oblong, subcoriaceous 68. R. phacochristum 

40. Corolla 4-6 cm long. 

42. Leaves oblong, base truncate or slightly cordate. Umbels 4-8(-10)-flowered. 

69. R. haematophthalmum 
42. Leaves broad-elliptic to ovate-elliptic, base broad-attenuate to almost rounded. Umbels 

2-3-flowered 70. R. phaeops 

39. Corolla but laxly and more caducously lepidote, or glabrous outside. 

43. Anthers c. 2.5 mm. Corolla (3-)3.5(-4) cm 71. R. truncicolum 

43. Anthers 3.5-5 mm. Corolla (4-)5-7 cm. 
44. Style with a short though distinct purely lepidote zone (as found in 56. R. konori) above the 
lower zone, where hairs and scales are found together or the hairs are predominant. 

57. R. phaeopeplum 
44. Style without such an upper exclusively lepidote zone. 
45. Corolla white or pink, narrowly funnel-shaped, regular; tube straight or almost so. 
46. Corolla 5-merous. Leaves broad-elliptic, more rarely oblong-elliptic or almost rounded. 

72. R. rappardii 

46. Corolla 6-7-merous. Leaves oblong or elliptic-oblong 73. R. dianthosmum 

45. Corolla red, tubular, — zygomorphous; tube rather straight below, slightly curved and 
gradually dilated upwards 74. R. rubelium 



33. Rhododendron eymae Sleu.m. Reinwardtia 5 
(1960) 74. 

Dwarf shrub, 10-25 cm. Branchlets divaricate, 
terete, tips densely covered with rufous squamular 
scales, otherwise scabrid and dark coloured; inter- 
nodes 0.5-1.5 cm. Leaves scattered and equi- 
distant along the upper internodes, ovate or nearly 
oblong ovate, apex short-acuminate, acutish, base 
subtruncate rounded, very coriaceous, rigid, revo- 
lute to or nearly to the midrib, very young ones 
stellate lepidote on both faces, mature ones 
glabrescent above, though iz persistently lepidote 
beneath (scales squamular, the margin deeply 
stellate-lacerate, subsessile, the minute centre 
deepened, each scale on top of a persistent epi- 
dermal elevation), scabrid 0.8-1.2 by 0.4-0.7 cm 
when extended, only 2-3 mm wide in the rcvolute 
state, midrib obscure above, a little prominent 
beneath nerves obsolete; petiole lepidote, thick, 
0.5-1.5 mm. Flowers solitary or mostly in twos, 
rarely in threes. Pcrulae ovate-acuminate, lepidote 
at the apex dorsally, glabrous otherwise, up to 
1 by 0.6 cm. Bracteolcs filiform, glabrous, up to 
8 mm. Pedicels densely rufous-squamular-lepidotc, 
(0.7-)l-1.2cm by c. I mmat anthesis. Ca/>'jrsmall, 
densely lepidote, irregularly 5-dentatc, teeth broad- 
triangular, subacute, I mm. or one or the other of 
them elongate to 3 mm. Corolla tubular below, 
dilated distally, rather fleshy, sulfureous, c. 1.3 cm 
long in all, tube subcylindric, 8-9 mm long, 3-4 
mm o at the base. 5-6 mm at apex, very densely 
rufous-stcllatc-icpidotc outside, glabrous inside, 
lobes subcrect or a little spreading, subspathulate 
to orbicular, 4-5 mm a, densely lepidote outside, 
the margin excepted, glabrous inside. Stamens 10, 
unequal, only partly a little exceeding the corolla 
tube; filaments filiform, slightly dilated above the 
base, glabrous; anthers broad-oblong, dchi.sccnt by 
anticous pores, c. i .5 by I mm. Disk low, glabrous. 



Ovary ovoid-conical, very densely stellate-lepidote, 
3-4 by c. 2.5 mm, rather abrupt with the style, 
the latter thick-columnar, glabrous, c. 1.6 mm 
long, stigma subcapitate, very shortly 5-lobed. 
Capsule ovoid, shortly contracted distally, 8-9 
by c. 5 mm, densely rufous-stellate-lepidote; style 
persistent, c. 1.5 mm; valves not twisted; calyx a 
little accrescent; fruit-pedicel 1.5-1.8 cm. 

Distr. Malesia: SW. Central Celebes (Lati- 
modjong Range: Mt Rantemario), 3100-3300 m. 
Once collected. Fl.fr. June. 

34. Rhododendron revolutum Sleum. Reinwardtia 
5 (1960) 74. 

Low shrub, 10-40 cm. Branches mostly much 
divided; branchlets terete, c. 1.5 mm 0, tips 
densely stellate-lepidote and scabrid; internodes 
l-3(-4) cm. Leaves in 4-6-merous pseudowhorls 
at the upper 2-3 nodes, ± narrowly ovate, apex 
short-acuminate, subacute, base truncate-rounded, 
coriaceous, very stiff, mostly revolute to the mid- 
rib, densely rufous-brown-lepidote on both faces 
initially (scales irregularly and ± deeply stellate- 
incised, the minute centre extended downwards 
into a stem-like foot, each scale on top of a 
thickish persistant epidermal tubercle), early 
glabrescent and scabrid above, lepidote for a long 
time beneath, 1-1.4 by 0.4-0.7 cm when extended, 
only : 4 mm wide when revolute, midrib very 
slightly impressed above, obscure beneath as are 
the nerves; petiole thick, lepidote, 0.5-1 mm. 
Flowers solitary or in twos. Outer perulae ovate- 
subulate, i lepidote dorsally, inner ones narrowly 
ovate-acuminate, ciliatc and set with caducous 
marginal scales, glabrous otherwise, up to 1 by 0.5 
cm. Bracteolcs filiform, laxly long-hairy and 
lepidote, c. 1 cm. Pedicels very densely rufous- 
stellatc-lcpidotc, 1.2-2 cm at anlhcsis. nearly 1 
mm a. Calyx subobliqucly disk-shaped, densely 



504 



Flora Malesiana 



[ser. I, vol. 6* 



rufous-stellate-lepidote, 3-4 mm 0, lobes obtuse, c. 
1 mm. Corolla tubular-infundibuliform, bright 
pink, (1.5)-1. 7-2.2 cm, tube 0.9-1.4 cm, base 
rounded and 3-4 mm 0, gradually dilated up- 
wards to 5-6 mm, densely stellate-lepidote outside, 
glabrous inside, lobes subspathulate-orbicular, 
wrinkled, 7-9 mm 0, laxly lepidote outside from 
the base to the middle, otherwise glabrous. 
Stamens 10, unequal, hardly exserted from the 
throat; filaments linear, slightly dilated above the 
base, glabrous; anthers oblong elliptic, c. 1.3 mm, 
dehiscent by anticous pores. Disk low, glabrous. 
Ovary ovoid-conical, densely stellate-lepidote, 
apex obtuse or abrupt with the style, c. 5 by 3 mm. 
Style columnar, glabrous, c. 1.5 by 0.4-0.5 mm; 
stigma thick-obconical, slightly 5-lobed. Capsule 
ovoid-ellipsoid, densely stellate-lepidote, c. 8 by 4 
mm, crowned by a 2.5 mm long style; stigma 
manifestly 5-lobed; fruit-pedicel c. 2 cm. 

Distr. Malesia: W. New Guinea (Lake 
Habbema), once found. 

Ecol. Restricted to crevices and sandy niches 
on sterile limestone slopes, 3225 m. Fl. Aug. 

35. Rhododendron extrorsum J. J. S. Nova Guinea 
18 (1936) 95, t. 19, 2; H. J. Lam, Nat. Tijd. N. L 89 
(1929) 97, f. 54 (phot.); Sleum. Reinwardtia 5 
(1960) 75. 

Lax shrub, 0.3-1.2 m, with irregular, often 
tortuous branches covered with grey bark. Branch- 
lets dichotomously ramose, terete, densely rusty- 
scurfy-lepidote at the tips, glabrescent and scabrous 
below, blackish when dry; internodes 0.8-2.3(-4) 
cm. Leaves 4-5 per pseudowhorl at the upper 2-3 
nodes, with some much reduced ones scattered 
along Ihe internodes, normal ones ovate, apex 
short-acuminate, subacute, base rounded-obtuse, 
coriaceous, stiff, dull light green above, light 
greenish yellow beneath, strongly revolute, often 
so to the midrib, very densely stellate-lepidote on 
both faces initially (scales ri deeply irregularly 
lacerate in the marginal zone, not rarely incised to 
the minute centre, each scale shortly dendroid and 
on top of a distinct epidermal permanent eleva- 
tion), rather early glabrescent above, less or later 
so beneath, finally scabrid, 1-1.6 by 0.4—0.7 cm 
when expanded, only 0.3-0.5 cm wide in the revo- 
lute state, midrib slightly grooved above, some- 
what prominent beneath, nerves obscure; petiole 
stout, lepidote, 2-3 mm. Flowers solitary or in 
twos. Ferulae ovate, acuminate, ciliolate, scaly 
along the dorsal middle line, glabrous otherwise, 
up to 1.5 by 0.5 cm. Bracteoles linear to linear- 
subspathulate, glabrous, up to 1 cm long. Pedicels 
densely rufous-lepidote, c. 5 mm by c. 1 mm. 
Calyx oblique, disk-shaped, very shortly obtusely 
5-lobed, densely lepidote, c. 2.5 mm 0. Corolla very 
light lemon-yellow, nearly white, broadly funnel- 
shaped, 5-lobed halfway or slightly more, c. 1.9 
cm long in all, tube wide, gradually dilated from 
3-4 mm at base to 8-9 mm at the throat, sub- 
densely covered with brown scales outside, 
glabrous inside, lobes somewhat spreading, wavy, 
obliquely suborbicular, c. 1.2 cm 0, glabrous 
except for some scales at the very base outside. 



Stamens 10, slightly unequal, 7-8 mm; filaments 
linear, slightly dilated above the base, pale green, 
glabrous; anthers red-brown, oblong, dehiscing by 
distinctly backward pores, c. 1.5 mm. Z>/i/c obsolete. 
Ovary ovoid-conical, obtuse, c. 4 by 3 mm, 
densely stellate-lepidote; style abrupt with the 
ovary, very shortly obconical, hardly 1 mm, 
glabrous; stigma convex, slightly lobed. Capsule 
ovoid, densely rusty-stellate-lepidote, c. \.2 by 
0.6 cm, pedicel 8 by 1 mm. 

Distr. Malesia: W. NewGuinea(Mt Doorman). 

Ecol. Open slope, also in the saddle between 
the summits, scattered though common, 3500 m. 
Fl. Oct. 

36. Rhododendron hellwigii Warb. Hot. Jahrb. 16 
(1892) 15, 26; K. Sch. & Laut. Fl. Schutzgeb. 
(1901) 487; Schltr, Hot. Jahrb. 55 (1918) 157; 
DiELS, ibid. 62 (1929) 487; Sleum. Reinwardtia 5 
(1960) 76. 

Shrub or small tree, terrestrial or apparently 
usually epiphytic. Branchlets 4-6 mm 0, terete, 
glabrous except the somewhat flattened or angular 
and ~ densely rufous-stellate-lepidote tips; inter- 
nodes 2-8(-15) cm. Leaves in 3-5-merous pseudo- 
whorls at the upper 1-2 nodes, or 2 or 3 scattered 
but near together at the nodes, ovate to ovate- 
elliptic, sometimes suborbicular, very coriaceous 
and firm, apex broadly attenuate and obtuse to 
rounded, base rounded, sometimes slightly cord- 
ate, greyish brownish above when dry, brownish 
beneath, rusty-stellate-lepidote on both faces initi- 
ally (scales small, irregularly incised, sessile or very 
shortly stalked, each on top of a minute epidermal 
tubercle which remains visible for a long time 
beneath), early glabrescent above with age, rather 
smooth or often somewhat rough to the touch, 
more persistently lepidote beneath, edge not 
revolute, 8-13(-16) by 5-8.5(-10.5) cm, midrib 
grooved above in the lower half, robust, rather 
flat, distinctly prominent to nearly the top, nerves 
8-10 pairs, rather irregular, the lowest 2 pairs 
from near the base of the blade, all ^r straight 
below, curved-anastomosing to the edge, slightly 
prominent on both faces, veins rather coarse, 
much less visible, veinlets obscure; petiole robust, 
a little flattened, grooved above, ± densely lepidote, 
(1-)1.2-1.6 (rarely up to 2.5) cm by 3-4 nun. 
Umbels 2-3-flowered. Flower buds ovoid, 3-3.5 
by 2.5-3 cm. Outer perulae ovate to suborbicular, 
inner ones broad-spathulate, obtuse or very 
shortly apiculate, densely covered with short, 
stoutish, nearly bullate hairs on both faces, 
glabrescent, remaining scabrous, not ciliate, up 
to 4 by 3.5 cm. Bracteoles linear to subspathulate, 
hairy or almost hirsute below, glabrous distally, 
c. 2-3.5 cm by 1-4 mm. Pedicels very stout, 
densely dark rufous-lepidote, 0.6-U-1.3) cm by 
2-2.5 mm. Calyx rim-like, very shortly obtusely 
5-6-lobed, c. 6 mm 0. Corolla broadly funnel- 
shaped, dark blood red, rather fleshy, 6.5-7 (some- 
times up to 8) cm in all, tube slightly dilated up- 
wards and moderately curved, (4-)4.5(-5) cm 
long, 1-1.3 cm at base, 1.7-2 cm at apex, 
glabrous outside, densely set with short ± retrorse 



Jan. 1966] 



Ericaceae (Sleumer) 



505 



hairs inside in the lower half, lobes 6-7, somewhat 
expanded, broadly spathulute or obovate, (2.5-) 
3-3.5 by 2.5-3(-3.5) cm. Stamens 12-14, unequal, 
slightly e.xserted from the throat; filaments linear 
(1 mm), clad with retrorse or patent hairs in their 
lower -3. glabrous and filiform distally; anthers 
linear-oblong. 7-8 by 1.5 mm, base of each theca 
shortly obtusely bilobed. Disk glabrous, blackish 
when dry. Ovary subovoid-cylindric, c. 10 by 4-5 
imn. densely covered with small flattened scales, 
— abruptly attenuate to the curved style, the latter 
thick, glabrous, sometimes shining. 3-3.5(-4) cm 
by c. 2 mm; stigma shortly thick-capitate, ob- 
scurely 5-6-lobed. 

Distr. Malesia: E. New Guinea (Finisterre and 
Saruwaged Mts). 

Ecol. Terrestrial on steep slopes among grasses 
at 1675-2100 m, mostly epiphytic in mountain or 
mossy forest, 1065-2440 m (also reported up to 
4000 m. which seems to be less reliable). Fl. Jan.- 
May, Sept., Oct. 

37. Rhododendron superbum Sleum. Reinwardtia 
5 (1960) 76; Blumea 11 (1961) 115; ibid. 12 (1963) 
95; ibid. 12 (1964) 339; Jahrb. Rhodod. Ges. 
Bremen (1963) f. 19, iion Hort. ex Lavallee, 1877, 
nom. nud. — R. devrieseanum vel aff. Stonor, 
Rhodod. Yearbook 6 (1951) 50, f. 48. 

Stiff shrub, (0.5-)1.5-2(-6) m. Branchlets terete, 
(5-)6-7 mm 0, tips striate and somewhat flattened 
in dry specimens, densely fugaciously rusty- 
stellate-lepidote, older parts glabrescent, i scab- 
rid; internodes 5-10 cm. Leaves in 4-merous 
pseudowhorls or scattered and ± close together 
at the upper nodes, broad- or ovate-, sometimes 
subobovate-elliptic, very coriaceous, stiff", erect, 
apex broad-attenuate and very shortly acuminate 
or obtuse, base subtruncate-rounded or slightly 
cordate, dark green above, slightly paler 

beneath, dull young ones densely covered on both 
faces with a golden tomentum of rather small 
scales (scales irregularly stellate-lacerate at the 
marginal zone, each on top of a distinct persistent 
epidermal tubercle), early glabrescent above, 
tardily so beneath, finally very rough on both 
surfaces, flat, (8-) 10— 12 by (4.5-)6-8 cm, midrib 
sulcate above in the lower part, less so apically, 
stout and : flattish beneath, c. 4 mm wide at the 
base, gradually narrower apically, nerves c. 8 
pairs, 2 lowest nearly from the base of the blade, 
upper ones patently from the midrib, all rather 
straight below, curved before the edge and anasto- 
mosing, a little prominent on both faces, veins 
still less conspicuous, somewhat impressed above 
in fully mature leaves, slightly raised beneath, no 
proper reticulation visible; petioles flattened, 
grooved above, 1.5-2 cm by 2.5-4 mm. Umbels 
3-5-flowcrcd. Pcrulae ovate, subcoriaceous, red- 
dish carmine, densely set with coarse thick hairs 
and rough to the touch outside, up to 5 by 3 cm. 
Bractcolcs linear to lincar-subspathulatc, laxly 
hairy, up to 5 by 0.5 cm. Pedicels robust, densely 
rusty-stcllatc-lcpidotc, (0.8 )l.2-2.4 cm by 2-2.5 
mm. Calyx oblique, cup-shaped, Icpidote dorsally, 
7-9 mm 0, lobes obtusely deltoid, manifestly 



shortly fimbriate, 1.3-2 mm, sometimes much 
larger and petaloid. Corolla funnel- or widely 
trumpet-shaped, fleshy or waxy in texture, white 
to cream or very light yellow, sometimes faintly 
flushed with pink, and shades to deep pink towards 
the tube, or pink with reddish throat, outside of 
tube sometimes ± uniformly pink, scented, (1 1-) 
11.5-12.5 cm long in all, 9-10(-12) cm across the 
limb in dry, 13-14 cm long in fresh specimens, 
tube gradually dilated from the base upwards, 
6.5-7.5 cm, c. 1 cm at base, 2-2.5 cm at apex, 
laxly lepidote or mostly glabrous outside, covered 
with white, ± retrorse and rather dense hairs in the 
lower half inside, glabrous otherwise, lobes 6-7, 
i spreading, broadly obovate or spathulate, 
4.5-5 by 3-3.5 cm, glabrous. Stamens 12 or 14, 
slightly longer than the corolla tube, unequal; 
filaments light green, linear, c. 1 mm wide at the 
base, very densely ± patently hairy below, less so 
or even glabrous and narrower in the upper '/2 
to 5/4; anthers linear-oblong, white, 8-9 by c. 
1.5 mm, thecae minutely lobulate at the base. 
Disk c. 1 mm high, glabrous. Ovary thickly 
conical-cylindric, densely clad with copper- 
coloured, stellate scales, c. 1 1 by 4 mm, not rarely 
abruptly narrowed to the red or green style, the 
latter completely glabrous, 5.5-6.5 cm by c. 2 mm; 
stigma capitate, 7-8 mm across in fresh specimens, 
obscurely 6-7-lobed. Capsule cylindric, shortly 
narrowed at both ends, widest above the middle, 
c. 5 by c. 1.3 cm. Seeds 7-8 mm including tails. 

Distr. Malesia: New Guinea (Southern, 
Western, and Eastern Highlands: Mt Giluwe, Mt 
Sugarloaf, Yaki Valley, Minj-Nona Divide, Mt 
Hagen, Bismarck Mts, Marafunga above Goroka; 
Mt Saruwaged area). 

Ecol. Generally epiphytic on the upper branches 
of large trees, apparently with preference just 
below the true moss forest level; terrestrial on 
Mt Michael and Mt Giluwe, in low shrub and on 
edge of cloud forest, ( 1 525-)2300-3050 m. in Uinba 
(Minj-Nona Divide) even in fallow native gardens 
at 1970 m. Fl.fr. April-Sept. 

Vern. Marugump, Enga, wamp-wamp, Mendi, 
kukump, Minj. 

Note. Stonor described this as growing 
commonly, but individually, its huge, lily-like 
flowers making glowing spots of colour among the 
greens of the innumerable other epiphytes. The 
flower has an exquisite scent, reminiscent of 
Carnations or the Regale Lily; he thinks, that 
this and other high-epiphytic Rhododendrons with 
a delicious scent like R. ai^'ailiodacmonis "are 
pollinated by the Lory parrots, some of which live 
in the moss-forests, and the more so as wc know 
parrots to be among the few birds with a sense of 
smell". 

38. Rhododendron prainianum Koord. Neva 
Guinea « (1909) l«7; Sii dm. Kcinwardfia 5 (I960) 
78. 

Epiphytic or terrestrial shrub, up to 2 m. 
Branchlets terete, slender, c. 1.5 mm «, few-bran- 
ched, tips very densely covered with rubiginosc, 
stellate scales, glabrescent below; internodes 2-6 



506 



Flora Malesiana 



[ser. I, vol. 6* 



cm. Leaves in 3-4-merous pseudowhorls, linear- 
lanceolate or linear, subsessiie, straight or sub- 
falcate, aE>ex gradually attenuate, subacute, base 
shortly cuneate, subcoriaceous, initially lepidote 
on both faces (scales stellate-incised at the marginal 
zone to various degree, dendroid, each scale on 
top of a minute epidermal persistent tubercle), 
early glabrescent and scabrid above, tardily so 
beneath, edge very slightly or not revolute, 5-10 
by (0.3-)0.4-0.7 cm, midrib strongly impressed 
above, prominent beneath, nerves i obscure; 
petiole slender, densely lepidote, 2(-3) mm. 
Flowers solitary or mostly in twos. Ferulae ovate, 
rostrate-acuminate, stellate-Iepidote outside main- 
ly at the apex, up to 15 by 6 mm. Bracteoles linear, 
laxly lepidote, 1-1.5 cm. Pedicels very slender, 
densely lepidote, 9-1 1 mm. Calyx cup-shaped, 
c. 1 mm 0, glabrescent outside, with obscure 
teeth. Corolla tubular, slightly zygomorphic, 
membranous, lilac, glabrous in- and outside, 
2.5-2.7 cm long in all, tube cylindric, straight or 
slightly curved, 1.5-1.8 cm long, c. 3 mm at 
base, not more than 4 mm at apex, lobes spreading, 
obovate-spathulate to suborbicular, 6-8 by 5-7 
mm. Stamens 10, 2.5-2.8 cm; filaments linear 
below, filiform distally, glabrous; anthers broad- 
oblong, 1.5 by 1 mm. Disk finely short-pubescent 
at the upper margin. Ovary elongate-conical, very 
densely rufous-stellate-lepidote, c. 3^ by 1 mm, 
gradually tapering to the slender style (1.5-2 cm), 
which is lepidote in the lower "/j to 1/3. bearing 
sometimes several short hairs in the middle 
(practically epilose), glabrous apically; stigma 
short-obconical. 

Distr. Malesia: New Guinea (Hellwig Mts; 
Orion Mts: Sibil R. Valley). 

Ecol. On poor clayey soil, at 900-1300 m. 
Fl. May-Aug. 

39. Rhododendron stelligerum Sleum. Blumea 11 
(1961) 115. 

Shrub, 1-1.5 m, with sparse erect, much branch- 
ed twigs. Branchlets slender, tips very densely 
covered with rusty stellate scales, older parts 
glabrescent, ashy; internodes (1.5-)3-7 cm by 
1.5-2 mm. Leaves in 4-5-merous pseudowhorls 
at the upper (2-3) nodes, sublinear-lanceolate, 
attenuate at both ends, apex subacute, coriaceous, 
initially very densely clothed with rusty stellate 
scales, glabrescent and remaining rough to the 
touch above with age, more persistently lepidote 
beneath (scales partly covering each other, each on 
top of a minute, pers'stent epidermal tubercle, 
marginal zone rather wide, deeply and irregularly 
stellate-incised or -lacerate, ends of the laciniae 
filiform, centre rather large, dark chestnut brown, 
tapering downwards into a kind of short stem 
or foot), much revolute at the edge, (2-)2.5^.5 cm 
long, (0.3-)0.4-0.8 cm wide in the revolute state, 
midrib hardly indicated above, obtuse and much 
prominent beneath, nerves inconspicuous; petiole 
very densely lepidote, 4-6 by c. 1 mm. Flowers 
2 or 3 per umbel, rarely solitary. Flower buds 
ovoid-oblong, c. 1 cm; outer perulae broadly 
subulate, more inner ones ovate, subabruptly 



narrowed into an acumen of 2-3 mm, slightly 
keeled and lepidote in the upper part dorsally, 
innermost ones ovate-oblong, obtuse, -± glabrous 
dorsally, all densely lepidote-fimbriate, up to 1 by 
0.5 cm. Bracteoles filiform, laxly caducously 
lepidote, up to 1 cm. Pedicels entirely covered 
with almost golden stellate scales, 1.2-1.6 cm by 
0.7-1 mm. Calyx disk-shaped, suboblique, shortly 
5-lobed (or one or two lobe(s) sometimes elongated 
into a narrow lacinia up to 8 mm), c. 2.5 mm 0, 
densely lepidote dorsally. Corolla tubular below, 
gradually subcampanulate-dilated upwards, not 
rarely somewhat curved, markedly zygomorphous, 
deep pink when fresh, magenta in dry specimens, 
subdensely set with golden stellate scales and ± 
crisp, white hairs outside at the tube and the base 
of the lobes, glabrous for the rest, 3.5-4 cm long 
in all, tube (2-)2.5-3.2 cm long, 5-6 mm at base, 
7-9 mm at throat, lobes erecto-patent, broadly 
spathulate or obovate, 0.7-1 (-1.2) by 0.6-0.8 cm. 
Stamens 10, unequal, i long exserted from the 
tube, the longest attaining half the corolla lobes; 
filaments linear, 1 mm wide below, 0.5 mm wide 
in the upper part, quite glabrous; anthers obovate- 
oblong, 2-2.5 by 1.5 mm. Z)/.vA: slightly prominent, 
glabrous. Ovary subcylindric, c. 5 by 2.5 mm, 
very densely clothed with deeply stellate-incised 
golden scales, but no hairs, gradually narrowed 
into the style, which is covered with dense or more 
laxly arranged scales and hairs to the lower y^, 
glabrous for the rest; stigma thick-turbinate, 
shortly 5-lobed. Capsule subcylindric, slightly 
curved, densely stellate-Iepidote, not hairy at all, 
2-2.5 by 0.4 cm, splitting into 5 twisted and sub- 
reflexed valves. Seeds very narrow, almost filiform, 
4 mm, long-tailed at both ends. 

Distr. Malesia: New Guinea (Star Mts: on 
Mt Antares), once collected. 

Ecol. In semi-alpine shrub-vegetation, 3000 m. 
Fl. July. 

40. Rhododendron neriifolium Schltr, Bot. Jahrb. 
55 (1918) 149; Sleum. Reinwardtia 5 (1960) 78. 

'"Epiphytic shrub. Branchlets elongate, flaccid, 
terete, laxly foliate, densely stellate-Iepidote. 
Leaves opposite, linear-lanceolate or nearly linear, 
apex acute, base nearly rounded, lepidote on both 
faces initially, glabrescent above, entire, 3.5-7 by 
0.5-0.9 cm; petiole 2-3 mm. Flowers solitary. 
Perulae and bracteoles not known. Pedicels 
densely stellate-Iepidote, up to 5 mm. Calyx disk- 
shaped, densely stellate-Iepidote. very shortly 5- 
lobed, 2.5-3 mm 0. Corolla tubular, bright pink- 
red to purplish, zygomorphous and slightly curved, 
rather densely stellate-Iepidote outside, glabrous 
inside, 2.7-3.3 cm, lobed to the upper V3. lobes 
suborbicular, erecto-patent. Stamens 10, very 
slender, up to 2.5 cm; filaments filiform, laxly 
hairy; anthers oval, c. 2 mm. Ovary oblongoid, 
densely stellate-Iepidote, c. 6 mm. Style subulate, 
laxly hairy, 6 mm; stigma capitate, 6-gibbous." 

Distr. Malesia: New Guinea (Schrader Mts), 
in mountain forest at c. 2070 m. Fl. May-June. 

Note. Neither type material preserved nor 
recollected; description after the original diagnosis. 



Jan. 1966] 



Ericaceae (Sleumer) 



507 



41. Rhododendron hooglandii Sleum. Reinwardtia 
5 (1960) 75; Blumea 12 (1964) 339. —Fig. 31 b— <:. 

Straggling, much branched shrub, 0.7-1.8 m, 
also epiphytic. Branchlets densely set with stalked 
scales, warty and rough to the touch at the ultimate 
1-2 internodes, early covered with greyish or 
whitish cork below; internodes 2-6(-8) cm. Leaves 
in 4~6-merous pseudowhorls at the upper l-2(-3) 
nodes, linear, apex subobtuse, base narrowed to 
the petiole, coriaceous, densely covered on both 
faces with yellowish stalked, deeply stellate- 
incised scales which go early, leaving the stalks, 
thus making the surface of the lamina scabrous, 
revolute to near the midrib, (3-)3.5-5.5 cm by 2-3 
(-4) mm, midrib faintly impressed above, as thick 
as the petiole and somewhat prominent beneath, 
nerves not visible; petiole stoutish, densely stalked- 
lepidote, (4— )5-8 mm. Umbels (l-)2-flowered. 
Outer perulae subulate, inner ones ovate-sub- 
acuminate, almost blunt, lepidote along the dorsal 
middle line, glabrous otherwise, lepidote-fimbriate, 
3-7 by 1^ mm. Bracteoles filiform, minute. 
Pedicels rather robust, densely and exclusively 
clothed with yellowish stalked scales, c. 1 cm. 
Calyx obliquely patellar, very obtusely and 
shortly 5-lobed, densely set with nearly sessile 
scales, c. 3 mm o. Corolla tubular, somewhat 
curved and zygomorphous, pinkish red, 2.8-3 cm 
long in all, densely covered outside with sub- 
stipitate or sessile, subentire to variously dented 
scales, glabrous inside, tube 2-2.3 cm, c. 5 mm 
at base, hardly dilated upwards, lobes broadly 
obovate, 5-8 mm o. Stamens 10, as long as or 
slightly longer than the corolla; filaments linear, 
glabrous: anthers subquadratc-oblong, 1.5 by 1 
mm. Disk glabrous. Ovary nearly conical, gradu- 
ally tapering to the style, very densely lepidote, 
not hairy at all, c. 5 by 2 mm. Style red, rather 
slender, with some scales at the very base, glabrous 
for the rest, c. 2 mm; stigma subcapitate, greenish 
yellow, 5-lobed. Capsule c. 1 .8 cm. 

Distr. Malesia: New Guinea (Eastern High- 
lands: Goroka Subdistr., Kerigomna; Western 
Highlands: Upper Minj R. Valley, Minj-Nona 
Divide). 

Ecol. Forest grassland edge or alpine thickets, 
not common, 3000-3410 m. Fl. July. 

42. Rhododendron rarum Schltr, Sot. Jahrb. 55 
(1918) 150; Si.ELM. Reinwardtia 5 (1960) 78; 
Jahrb. Rhodod. Gcs. Bremen (1963) f. 17; Blumea 
12(1963)97; ihicL 12 (1964) 339. — Fig. 11. 

Epiphytic slender shrub, up to I m high, much 
branched. Branchlets terete, 1-2 mm 0, laxly 
Leaved, tips densely rusty-squamular-lcpidote, 
older parts glabrcsccnt, somewhat scabrid and 
early covered with greyish cork; internodes (1.5-) 
3-10 cm. Leaves opposite or mostly in 3-5-merous 
pseudowhorls at the upper 3-4 nodes, often 
different in size in the same whorl, with some 
much reduced leaflets between them, in general 
larger in the lower whorls, lanceolate to ovatc- 
lanccolatc, apex gradually long-acuminalc, sub- 
acute, base very broadly attenuate or mostly h 
rounded, subcoriaccous, rather thin, deep green 



above, pale green below with brown sparse scales, 
densely lepidote initially on both faces (scales ± 
deeply stellate-incised, ± dendroid, each scale on 
top of a minute epidermal persistent tubercle), 
rather early glabrescent above, persistent for some 
time beneath, finally also glabrescent there, a little 
rough to the touch on both sides in later stages, 
slightly revolute at the margin especially towards 
the base, 3-7 by (0.3-)0.6-l(-1.3, rarely in part 
in the lowest ones up to 1.6) cm, always widest in 
the lower third, midrib faintly impressed above, 
prominent beneath, nerves obscure; petiole slender, 
densely lepidote, (l-)2-3 mm. Flowers solitary or 
in twos, sometimes in 3-4-flowered umbels. Flower 
buds ovoid, acute. Outer perulae spreading, sub- 
uliform, inner ones ovate, rather abruptly subulate- 
acuminate, ciliate, all densely but ± fugaciously 
squamular-stellate-lepidote outside, glabrous in- 
side, up to 1 by 0.6 cm. Bracteoles linear to fili- 
form, rt glabrous, up to 1 cm. Pedicels slender, 
densely stellate-lepidote, (0.7-)l-1.3 cm. Calyx 
small, disk-like, c. 2 mm 0, not or very shortly 
obtusely lobed, lepidote dorsally. Corolla (deep) 
pink, crimson to scarlet or blood red, tubular in 
the lower 2/3, expanded at the limb, rather mani- 
festly zygomorphous, (2.3-)2.5-3.5(-3.8) cm in all, 
tube slightly curved, (1.8-)2-2.5(-2.7) cm long, 
3-4 mm at base, gradually widened to 6-8 mm 
at apex, ± densely stellate-lepidote and epilose 
outside, glabrous or very laxly hairy inside, lobes 
broad-obovate to suborbicular, (6-)7-9 by 6-8 
(-9) mm. Stamens 10, unequal, well exserted from 
the throat, in part as long as the corolla; filaments 
narrow-linear, glabrous below, very laxly hairy 
in the middle, glabrous distally; anthers oblong, 
c. 2 by 1 mm. Disk glabrous or very minutely 
pubescent at the upper margin. Ovary subcylindric, 
densely stellate-lepidote, c. 6 by 2-2.5 mm, 
gradually tapering to the style, the latter stoutish, 
generally lepidote in the lower Vi and exclusively 
laxly patent-hairy above to or almost to the top, 
rarely ± glabrescent, ± equalling the corolla in 
length not before the latter is developed; stigma 
clavate-capitatc, obscurely 5-lobed. Capsule nar- 
row-oblong, 3-4 by 0.4-0.5 cm, valves twisted. 
Seeds c. 4 mm including the tails. 

Distr. Malesia: New Guinea (Western and 
Eastern Highlands: Mt Hagen, Nondugl; Bis- 
marck Mts). 

Ecol. Low epiphytic on tree trunks or rotten 
stumps in mountain forest, also in light moss- 
forest, 1585-3350 m, plentiful locally. Fl. April- 
Nov. 

Vern. Sezupezup, Enga, Kepilam, wamhuberi, 
Enga, Poio, remono kire, Kaugcl. 

43. Rhododendron dclicatulum Slhum. Reinwardtia 
5 (I960) 78. 

var. dclicatulum. 

Epiphytic shrub. 30-50 cm. Branchlets very 
slender, 1-2 mm o. terete, laxly foliate, tips 
dcn.sely stellate-lepidote, glabrcsccnt below; inter- 
nodes 3-6 cm. Leaves in 3-4-mcroiis pseudowhorls 
at the upper nodes, lincar-lanccolatc, apex gradu- 



508 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 1 1. Rhododendron rarum Schltr. Marafunga near Goroka, 2400 m, E. New Guinea (Sleumer4137) 

(Sleumer, 1961). 



ally acuminate and somewhat curved, subacute, 
base broadly attenuate into the petiole to nearly 
rounded, subcoriaceous, the very young ones 
rather densely covered on both faces with deeply 
stellate-incised, sessile or shortly dendroid scales, 
these on top of persisting epidermal tubercles, 
early glabrescent and rough to the touch above, 
tardily so beneath, slightly revolute at the margin, 
(3-)4— 6.5 by 0.4-0.7 cm, mostly widest below the 
middle, midrib sunk above, prominent beneath, 
nerves obsolete; petiole lepidote, c. 1 by 0.5-0.7 



mm. Umbels ± 3-flowered. Outer perulae subulate, 
inner ones ovate, with a subulate acumen for the 
ultimate 3-10 mm, rather densely stellate-iepidote 
dorsaliy, i glabrous internally, fimbriate, up to 
2 by 0.6 cm. Bracteoles linear-subspathulate, very 
laxly hairy, up to 1.5 cm by 1 mm. Pedicels slender, 
densely rufous-stellate-lepidote, c. 1.2 cm. Calyx 
small, oblique, annular, membranaceous, ob- 
scurely lobed, stellate-iepidote dorsaliy, shortly 
fimbriate, c. 2 mm 0. Corolla tubular below, 
widened at the limb, somewhat zygomorphous, 



Jan. 1966] 



Ericaceae (Sleumer) 



509 



pink, c. 2.7 cm long in all. tube 1.5-1.8 cm long, 
c. 3 mm o at base. c. 6 mm at apex, equally sub- 
densely shortly whitish hairy (but not lepidote) 
outside, — laxly hairy inside, lobes 5, suberect- 
patent, broadly obovate-spathulate or suborbicu- 
iar. 5-7 by 5-6 mm. Stamens 10, almost equalling 
the corolla in length; filaments filiform, very 
laxly hairy below, glabrous distally; anthers 
broad-oblong, 1.3 by c. 0.8 mm. Disk very shortly 
yellowish hairy at the upper margin, glabrous 
below. Ovary conical-cylindric, densely dark 
rusty-stellate-lepidote, 5 by 1.8 mm, gradually 
tapering to the style, which is rather slender, as 
long as the corolla and covered with spreading 
yellowish hairs to the top; stigma subcapitate. 

Distr. Malesia: W. New Guinea (Bernhard 
Camp, Idenburg R.), in mossy forest at 1800 m. 
Once collected. Fl. Jan. 

var. lanceolatoides Sleum. Blumea 11 (1961) 116. 

Leaves wider, ovate-lanceolate, (2.5)-3.5-5 by 
0.7-1.2 cm. 

Distr. Malesia: W. New Guinea (Star Mts), 
on peaty soil at 1500 m. Once collected. Fl. July. 

44. Rhododendron bryophilum Sleum. Reinwardtia 

5 (1960) 79; Blumea 12 (1963) 97. 

Shrub, epiphytic in high trees. Branchlets 
slender, 1-2 mm o, terete, laxly leaved, tips brown, 
densely covered with caducous star-shaped 
scales, older parts glabrescent and verruculose; 
internodes 2-6(-8) cm. Leaves opposite or in 3-4- 
merous pseudowhorls at the upper 3 or 4 nodes, 
oblong, apex short-attenuate, subobtuse, base 
broadly attenuate or mostly subtruncate-rounded, 
thinly coriaceous, subdensely lepidote on both 
faces initially (scales minute, deeply stellate-incised 
and subdendroid, early caducous, each on top of a 
persistent epidermal tubercle), rather early glabres- 
cent and subdensely tuberculate or scabrid, 
slightly or not revolute at the margin, 4-5.5 by 
(l-)l.3-2.2 cm, widest in the middle, midrib 
lepidote for a long time, very slightly sunk above, 
prominent beneath, nerves 4-5(-6) pairs, curved- 
anastomosing, faintly visible on both faces; petiole 
flattened, 2(-3) by c. 1 mm, lepidote. Umbels i 
3-flowercd. Ferulae ovate to obovate-spathulate, 
apiculate or obtuse, membranous, glabrous, 
minutely fimbriate, up to 8 by 6 mm. Bracteoles 
filiform, subglabrous, up to I cm. Pedicels very 
slender, densely minutely stellate-lcpidotc, cpilosc, 
10-12 by 0.4 mm. Calyx suboblique, small, 
obscurely 5-lobcd, stellate-lcpidotc outside, c. 
2 mm o. Corolla tubular, distinctly zygomorphous, 
pale salmon, thin, 3.5-4 cm long, quite glabrous 
outside, laxly hairy at the tube inside, tube cylin- 
dric, curved. (1 .8-)2.2-2.8 cm, 4-5 mm at base, 
widened to 7 mm at the throat, lobes spathulate, 
10-12 by c. 7 mm. Stamens 10, nearly equalling 
the corolla in length; filaments narrow-linear, 
glabrous immediately above the base, laxly hairy 
upward.s. but again glabrous distally; anthers 
oblong. 1.8-2 by I mm. Disk very shortly hairy at 
the upper margin, glabrous elsewhere. Ovary 
clongate-ovoid, densely and exclusively stcllatc- 



lepidote, r. 4 by 1 .5 mm, gradually attenuate to the 
style, the latter relatively thick, laxly covered 
with short, subpatent hairs to nearly the top, c. 
1.8 cm; stigma shortly club-shaped. 

Distr. Malesia: N. New Guinea (Cycloop Mts), 
in forest, 1065-1830 m. Fl. March-June. 

45. Rhododendron tuberculiferum J. J. S. Med. 
Rijksherb. 25 (1915) 4; Nova Guinea 12 (1917) 
504; ibid. 12 (1918) t. 199 a, 203 b-f; Sleum. 
Reinwardtia 5 (1960) 80. 

Shrub, 1-1.5 m. Branchlets terete, 2-3 mm 0, 
tips clad with dendroid stellate scales, older parts 
glabrescent and densely verruculose; internodes 
1.8-1 1 cm. Leaves 5-7 crowded towards the upper 
2-3 nodes, or in pseudowhorls, ovate to ovate- 
elliptic or oblong-elliptic, rarely lanceolate, coria- 
ceous, stiff, ± erecto-patent, apex obtusely 
acuminate or obtuse, sometimes apiculate, base 
rounded-obtuse to slightly cordate, subsessile, 
densely rufous-lepidote on both faces when very 
young (scales ± stellate, subsessile or subden- 
droid, each on top of a minute epidermal per- 
manent tubercle), early glabrescent especially 
above, manifestly verruculose and rough to the 
touch on both faces, slightly to strongly recurved 
at the margin, (1.7-)2^.8 by (0.9-)l.2-2.1 cm, 
midrib sunk above, prominent beneath, nerves 
obscure; petiole densely lepidote or early verru- 
culose, 1-4 by 1.5-2 mm. [/m6e/5 2-5(-6)-flowered. 
Outer perulae triangular, subulate-acuminate at 
the apex, inner ones ovate-acuminate to spathulate 
and shortly apiculate or obtuse, glabrous on both 
sides, subdenticulate, up to 1 .8 by 0.6cm. Bracteoles 
linear, glabrous, up to 2 cm by c. 1 mm. Pedicels 
rather slender, densely lepidote, 1.5-2.5 (some- 
times up to 3.2) cm. Calyx disk-shaped, oblique, 
lepidote dorsally. very shortly obtusely 5-lobed, 
c. 4 mm 0. Corolla r: zygomorphous, tubular 
below, funnel-shaped towards the limb, red, 3.8- 
4.5 cm, ; densely lepidote outside, margin of the 
lobes excepted, laxly hairy to the lower % of the 
tube inside, glabrous for the rest, tube cylindric, 
2.5-3.2 cm, slightly curved, 4-6 mm at base, 
widened to '■- 8 mm at apex, lobes 5 (rarely 6) 
suborbicular, retusc, 9-12 mm 0. Stamens 10 
(rarely 12), subcqual, 2.8-3.5 cm; filaments linear, 
rather laxly hairy below, less so upwards, glabrous 
distally; anthers oblong, base obtuse, c. 3 by 1 mm. 
Disk minutely pubescent, or practically glabrous. 
Ovary elongate-conical, very densely lepidote, 
with some hairs between the scales in the upper 
part respectively at the base of the rather abrupt 
style, c. 7 by 2.5 mm. Style patently hairy at base, 
otherwise glabrous, c. 1.7 cm; stigma capitate. 

Distr. Malesia: W. New Guinea (Wichmann 
and Hubrecht Mts), in open places at 3000-3100 
m. Fl. Feb. 

46. Rhododendron aspcrrimum Sleum. Blumea 12 
(1963) 97. 

Hpiphytic, sometimes scandcnt. divaricately 
branched shrub. Branchlets rather slender; inter- 
nodes 3-5(-7) cm by c. 2 mm. Leaves in 3-4- 
mcrous pseudowhorls at the upper 2-3 nodes. 



510 



Flora Malesiana 



[ser. I, vol. 6* 



subsessile, ovate, apex very broadly attenuate to 
rounded, base a little cordate, coriaceous, covered 
by a tomentum of rufous dendroid stellate scales 
as are the branchlets and petioles initially (scales 
stellate-incised or -lacerate to almost the centre, 
the latter small, protracted downwards into a short 
stipe, each on top of an epidermal marked eleva- 
tion), early glabrescent with age above, more 
tardily so beneath, rough to the touch by the 
persistent epidermal warts, which are paler than 
the blade, (2-)2.5-4(-4.5) by (1.5-)2-2.8(-3.2) 
cm, midrib impressed above, prominent beneath, 
nerves subinconspicuous on both sides, no reticula- 
tion; petiole 2 by 1.5 mm. Umbels (3-J4-6- 
flowered. Ferulae thin, ovate to spathulate, laxly 
caducously lepidote or finally glabrous, up to 
1.5 by 0.7 cm. bracteoles linear to filiform, 
glabrous, c. 1 cm. Pedicels rather slender, very 
densely stellate-lepidote, not hairy, 1.1-1.5 cm. 
Calyx obliquely patellar, shortly or hardly 5- 
lobed, densely lepidote dorsally, 2-3 mm 0. 
Corolla tubular, zygomorphous towards the limb, 
pink to purplish, 2.6-3 cm long in all, initially 
densely, later more laxly lepidote or finally ± 
glabrescent outside, laxly papillose-hairy at the 
tube inside, tube curved, c. 2 cm long, 4—5 mm 0, 
slightly dilated towards the throat, lobes unequal, 
erecto-patent, obovate-spathulate, c. 1 by 0.6 
cm. Stamens 10, subequalling the corolla in length; 
filaments linear, laxly papillose-hairy in the lower 
half, glabrous above; anthers obovate-elliptic, 
base obtuse, 2 by 1.3 mm. Disk glabrous below, 
hairy at the upper margin. Ovary subcylindric, 
c. 4 by 1.5 mm, densely set with rufous stellate 
scales and more laxly with paler hairs, rather 
abrupt with the style, the latter lepidote and hairy 
at the very base, and exclusively laxly patently 
hairy in the lower Vi (rarely glabrous even there), 
glabrous for the rest; stigma subcapitate. 

Distr. Malesia: NW. New Guinea (Wandam- 
men Peninsula: Wondiwoi Mts). 

Ecol. Primary forest, 800-1800 m. Fl. March. 

47. Rhododendron dielsianum Schltr, Bot. Jahrb. 
55 (1918) 150; Sleum. Reinwardtia 5 (1960) 80; 
Blumea 12 (1963) 97. — R. laureola Schltr, Bot. 
Jahrb. 55 (1918) 151. 

var. dielsianum. 

Epiphytic or terrestrial shrub, 0.3-0.9(-1.5) m, 
much branched. Branchlets erecto-patent, terete, 
1.5-3 mm 0, tips densely rusty-stellate-lepidote, 
or almost -tomentose, older parts glabrescent, 
rather smooth, brownish; internodes 1.5-8 cm. 
Leaves opposite or mostly in 3^-merous pseudo- 
whorls at the upper 2-3 nodes, elliptic-oblong or 
oblong, apex short-acuminate, apiculate, sub- 
acute, base broadly attenuate into the petiole, or 
rarely subrotundate, subcoriaceous, densely 
stellate-lepidote on both faces initially (scales 
rather small, deeply stellate, sessile or shortly 
dendroid and on top of rather low persistent 
epidermal elevations), early glabrescent above, 
tardily beneath especially near or on the m'drib, 
finally glabrescent and a little rough to the touch 



on both faces, margin not or slightly revolute, 
3-5.5 by (0.8-)1.2-2 (sometimes up to 2.7) cm, 
midrib somewhat sunk above, prominent beneath, 
nerves c. 6 pairs, very faintly impressed above, 
raised beneath, mostly subinconspicuous on both 
faces; petiole rather slender, (2-)3-5 mm. Flowers 
solitary or in twos, sometimes in 3(-5)-flowered 
umbels. Outer perulae subulate, inner ones ovate- 
subulate to ovate-apiculate, caducously stellate- 
lepidote in the upper half, short-fimbriate, up to 
1.5 by 0.6 cm. Bracteoles filiform, subglabrous, 
up to c. 1 cm. Pedicels slender, densely rusty- 
stellate-lepidote, 1-1.3 cm at anthesis. Calyx 
slightly or not oblique, disk-shaped, densely 
stellate-lepidote dorsally, c. 2.5 mm 0, very 
shortly obtusely 5-lobed. Corolla tubular to 
about ^/s of its total length, slightly curved and 
zygomorphous, sparsely lepidote outside, gla- 
brous or sparsely pubescent inside, (pale) pink, 
(2.5-)3-3.5 cm, tube 2-2.5 cm long, slightly 
saccate and 4—5 mm at the base, dilated to 8-10 
mm at apex, lobes broad-spathulate to sub- 
orbicular, somewhat expanded, c. 6-8 by 5-7 mm. 
Stamens 10, unequal, the longest nearly equalling 
the corolla; filaments linear to filiform, glabrous 
or very slightly hairy; anthers c. 2 by 1 mm. Disk 
rather prominent, glabrous. Ovary subcylindric, 
densely stellate-lepidote, 5-6 by 1.5 mm, gradually 
tapering to the slender style, which is glabrous or 
lepidote at the very base only, and slightly exceeds 
the throat; stigma subcapitate, slightly 5-gibbous. 

Distr. Malesia: E. New Guinea (Upper Sepik 
R. region; Kani, Finisterre and Saruwaged Mts; 
Kuno area: Tagan R.; once found in the SE. part 
near Ueli). 

Ecol. Epiphyte on tall trees in ravines and 
wooded slopes, in dense shade, terrestrial in 
secondary grassland, 1200-1940 m. Fl. May-Sept. 

Vern. Raubiu, Ueli. 

var. stylotrichum Sleum. Reinwardtia 5 (1960) 80; 
Blumea 11 (1961) 116. 

Slender erect shrub, 1.5-3 m. Corolla pink. 
Differs from var. dielsianum by the style which is 
laxly set with short spreading hairs in its lower 
half. 

Distr. Malesia: New Guinea (Eastern High- 
lands: Upper Dunantina Valley and the road 
between Kami and Lufa). 

Ecol. Hillside secondary grasslands, at 1700- 
1920 m, locally common. Fl. Aug. 

Vern. Hakessa, hongkempa, sosomepari, Asaro: 
Kefamo, gagin, Chimbu: Masul. 

48. Rhododendron stolleanum Schltr, Bot. Jahrb. 
55 (1917) 143; Sleum. Reinwardtia 5 (1960) 80. 
"Epiphytic, much branched shrub, c. 1 m. 
Branches and branchlets erecto-patent, terete, 
lepidote-tomentellous initially. Leaves opposite, 
lanceolate to lanceolate-ligulate, apex acute or 
obtuse, base broadly cuneate or rounded, coriace- 
ous, sparsely brown-lepidote above, covered 
beneath with a brown lepidote tomentum initially, 
4-8 by 1.2-3.2 cm; petiole 3-4 mm. Umbels 2-4- 
flowered. Pedicels slender, densely lepidote- 



Jan. 1966] 



Ericaceae (Sleumer) 



511 



tomentellous, 1.3-1.5 cm. Calyx patelliform, very 
shortly and bluntly 5-lobed, c. 3 mm o. Corolla 
tubular, 5-lobed — halfway, c. 2.5 cm long in all, 
glabrous outside, minutely puberulous inside, lilac, 
lobes obliquely obovate-oval, obtuse, glabrous on 
both faces. Stamens 10, c. 2 cm long; filaments 
filiform, sparsely puberulous or almost glabrous; 
anthers oblong, c. 1.5 mm. Ovary narrow-oblong, 
very densely and shortly stellate-lepidote-tomen- 
tellous, c. 8 mm including the subulate glabrous 
style; stigma obliquely capitate, 5-gibbous."' 

Distr. Malesia: E. New Guinea (Upper Sepik 
R. region: camp 'Felsspitze', c. 141^ 30'-4^ 10'), 
in low mountain forest. 

Note. As the holotype specimen is lost and no 
isotype is preserved, the above given description 
is a translation of the original, rather incomplete 
one. The position of R. stolleanum both in and 
within siibsect. Phaeovireya thus remains some- 
what doubtful; yet most of the characters given 
are those of R. dielsianum. 

49. Rhododendron psilanthum Sleum. Reinwardtia 
5 (1960) 81. 

Shrub. Branchlets slender, c. 2 mm o, terete, 
tips densely rusty-stellate-lepidote, early glabres- 
cent and smooth elsewhere; internodes c. 6.5 cm. 
Leaves opposite or in 3-merous pseudowhorls, 
elliptic or oblong-elliptic, apex shortly sub- 
acutely acuminate, apiculate, base attenuate into 
the petiole, subcoriaceous, stellate-lepidote on 
both faces initially (scales minute, deeply incised, 
dendroid, on top of minute epidermal tubercles), 
early glabrescent above, tardily so beneath, but 
finally glabrous and somewhat rough on both 
faces, edge slightly revolute, 3.5-^.5 by l.3-2(-2.2) 
cm, midrib a little impressed above, raised beneath, 
nerves inconspicuous; petiole densely lepidote, 
5-6 by c. 1 mm. Umbels 3-flowered. Pedicels slen- 
der, very densely stellate-lepidote, 1.2-1.5 cm by 
c. 0.5 mm. Calyx minute, oblique, in form of a 
short cup, subentire, r. 2 mm 0. Corolla tubular 
below, dilated at the limb, slightly zygomorphous, 
red, completely glabrous in- and outside, 3.5-3.8 
cm in all, tube c. 2.5 cm, at the base 5-gibbous 
and 4-5 mm o, widened to c. 8 mm at apex, lobes 
expanded, broadly obovatc-spathulate, 1-1.2 by 
0.8-1 cm. Stamens 10, slightly unequal, c. 3 cm; 
filaments narrow-linear, glabrous; anthers obovate- 
oblong. f. 2.2 by 1 mm, very shortly biappen- 
diculate at the base. Disk prominent, glabrous. 
Ovary subovoid-cylindric, c. 4 by 2 mm, very 
densely stellate-lepidote, rather abruptly attenuate 
into the style, the latter 2.5 cm in length, stellate- 
lepidote at the very base, otherwise entirely 
glabrous; stigma subcapitutc. 

Distr. Malesia: .SW. Central C'elebcs (Lati- 
modjong Range, Pintcalbn, spur of Mt Pokapind- 
jang), on the ridge at 2400-2600 m. Once found. 
Fl. June. 

50. KhodcMlcndron phaeochitiim F". v. M. Trans. 
R. .Soc. Vict. n.s. I, 2 (1889) 23; Sleum. Rein- 
wardtia 5 (I960) 83; Blumca 12 (1964) 339. 

hpiphytic shrub, c. I m high, branches stout, 



5-6 rrun 0. Branchlets slender, about 2 mm 0, 
densely dark rufous-stellate-lepidote at the upper 
1 or 2 internodes, glabrescent and densely verru- 
culose or scabrid below; internodes (2-)3-8 cm. 
Leaves opposite or 3 (-4) in a pseudowhorl or 
crowded near the upper nodes, elliptic or mostly 
obovate-elliptic, apex ± rounded-obtuse, apiculate, 
base broadly cuneate into the petiole or rounded, 
coriaceous, rather stiff, initially covered with a 
close dark rufous tomentum of stellular scales 
on both faces (scales very small, deeply incised, 
subsessile, each on top of a minute epidermal 
tubercle), early glabrous and somewhat lustrous, 
finally smooth above, lepidote for a long time 
beneath, ultimately glabrescent there but remain- 
ing manifestly verruculose and rough to the touch, 
entire, edge slightly revolute, 4-6.5 (rarely up to 
14) by 2.5-4 (rarely up to 7) cm, midrib slightly 
impressed above, rather stout and prominent 
beneath, nerves 5-6(-7) pairs, straight below 
only, irregularly curved-ascending and anastomos- 
ing in the upper part, faintly prominent in young, 
a little sunk above in fully mature leaves, promi- 
nent beneath, reticulation rather dense and con- 
spicuous, visible on both sides; petiole densely 
stellate-lepidote, 5-8 by 1-1.5 mm. Umbels 2-4 
(-5)-flowered. Perulae membranous, ovate to 
obovate-spathulate, — obtuse, glabrous, cadu- 
cously fimbriate, up to 1.5 by 0.8 cm. Bracteoles 
linear-spathulate, subglabrous, up to 1.5 cm by 
1 mm. Pedicels densely stellate lepidote (seemingly 
stellate-hairy), 1.4-2.5 cm by 1 mm. Calyx 
reduced to an almost annular membrane, irregu- 
larly dentate, stellate-lepidote dorsally, fimbriate, 
c. 3 mm 0. Corolla tubular below, expanded at the 
limb, ri zygomorphic, pink to deep red, subdenscly 
stellate-lepidote outside both on the tube and the 
base and mediane of the lobes, laxly hairy in the 
lower half of the tube inside, 3.5-4.5(-5) cm in all, 
tube slightly curved, 2-2.5(-3) cm long, 4-5 mm 
at base, widened to c. 8 mm at apex, lobes broad- 
obovate to suborbicular, (8-)10-15 by 8-12 mm. 
Stamens 10, slightly unequal, mostly a little 
exserted from the corolla; filaments linear and 
densely pubescent below, filiform and glabrous 
distally; anthers oblong, 2.5-3 by I mm. Disk ± 
densely white-hairy at the upper margin. Ovary 
cylindric, densely and exclusively rusty-stcllatc- 
lepidote, 5-7 by 2-3 mm, tapering to the stoutish 
style, which is nearly as long as the stamens and 
equally rusty-stcllatc-lcpidotc and may be very 
laxly hairy at least in its lower half, often to 
nearly the top, glabrous otherwise; stigma broadly 
obconical-capitatc. 

Distr. Malesia: E. New Guinea (Western and 
Eastern Highlands: Nondugl, Chimbu Divide, 
Goroka region; originally from Mt Musgrave in 
Central Distr.). 

Ecol. Epiphytic, or terrestrial on ridges (also 
limestone ridges). 2135-2560 m. /•/. April-Dec. 

Vcrn. Tegwa, Chimbu. 

51. Rhododendron bcycrinckianiim Koord. Nova 
Guinea K (r^l2) S76. t. l.M); J. J. •s.ibid. 12(1914) 
137; ibid. 12 (1917) 504, incl. var. longipetiolatiim 



512 



Flora Malesiana 



[ser. I, vol. 6" 



J. J. S. I.e.; ibid. 18 (1936) 94; Sleum. Reinwardtia 
5 (1960)81; Blumea 11 (1961) 116;/6/rf. 12(1963) 
98. — R. dasylepis Schltr, Bot. Jahrb. 55 (1918) 
153,f.4;STONOR, Rhodod. Yearbook 6 (1951 ) 50, 
f. Si. — R. schultzei ScHVYK, Bot. Jahrb. 55 (1918) 
154. — R. saavedranum Diels, ibid. 62 (1929) 
486. — R. chrysopeplon Sleum. ibid. 74 (1949) 540. 
Slender terrestrial or epiphytic, occasionally 
scandent shrub or treelet, 0.5-2.5 (rarely up to 4) 
m, loosely branched, erect. Branchlets stoutish to 
rather slender, often only 1.5 mm at apex, 
densely to very densely covered with rusty stellate 
scales in form of a detersile tomentum, which goes 
early in the older parts, these brownish and scabrid 
by numerous fine tubercles; internodes 1.5-8 
(sometimes up to 15) cm. Leaves opposite or 
mostly in 3-5(-7)-merous pseudowhorls, often 
rather variable in shape and size in the same whorl, 
lanceolate-oblong to oblong or obovate-oblong to 
elliptic, sometimes ovate or broad-elliptic, sub- 
obovate or suborbicular, apex broadly gradually 
attenuate, apiculate, subacute or mostly obtuse, 
base broad-attenuate into the petiole or rounded, 
coriaceous or at least rather stiff, erecto-patent, 
very densely rufous-lepidote ('tomentose') on 
both faces initially (scales irregularly stellate- 
lacerate up to the centre, dendroid, easily remov- 
able, each on top of a minute epidermal persistent 
tubercle), early glabrescent and somewhat shining 
and scabrid above, tardily so beneath, though 
finally glabrescent and scabrid here too, ± dark 
olivaceous green above, paler and more brownish 
beneath, margin mostly strongly revolute and 
more densely and/or more persistently stellate- 
lepidote than the intervenium beneath, generally 
(3-)4-6 by (l-)1.5-3.5 cm, occasionally up to 10 
by 5 cm in the lower leaves, midrib slightly im- 
pressed above, robust and much prominent 
beneath, nerves (4-)5-6(-7) pairs, rather straight 
below, curved-anastomosing before the edge, very 
slightly impressed above respectively raised 
beneath, often inconspicuous or completely 
covered by the tomentum beneath, reticulation 
dense, visibly prominent on both faces in fully 
mature leaves, occasionally ± impressed above; 
petiole densely stellate-lepidote, (3-)5-l 1 (some- 
times up to 20) by c. 2 mm. Umbels 2-4(-5)- 
flowered. Outer perulae subulate, completely 
covered with rusty stellate scales, inner ones ovate- 
acuminate, apiculate, lepidote at the dorsal upper 
part only, fimbriate, up to 1 by 0.5-0.7 cm. 
Bracteoles linear and glabrous or caducously 
lepidote below, subspathulate and laxly hairy 
apically, up to 1 cm. Pedicels densely and exclu- 
sively rusty-stellate-lepidote, at anthesis 1-1.5 cm 
by 0.7-1 mm, up to 2.2 cm in fruit. Calyx patellar, 
densely stellate-lepidote, shallowly obtusely 5- 
lobed, 3(-^) mm 0. Corolla tubular basally, 
widening distally, somewhat zygomorphous, most- 
ly pink to deep purplish pink or dark red, some- 
times pale pink or cream, rarely greenish or white, 
densely covered with ± caducous golden-brown 
stellate scales on the tube and the lower part of the 
lobes outside, completely glabrous inside, (2.5-) 
3-3.5 cm in all when dry (up to 4.5 cm when fresh!), 



tube straight or slightly curved, (1.5-)2-2.5 cm 
long, 4-6(-7) mm at base, somewhat dilated to- 
wards the mouth, lobeserecto-patent, suborbicular, 
7-10(-l2) by 6-9(-ll) mm. Stamens 10, unequal, 
mostly not quite equalling the corolla in length, 
sometimes finally exserted; filaments linear, ± 
reddish, glabrous or rarely laxly hairy; anthers 
obovate-oblong, 2-2.5 by 1.2 mm, brown to dark 
red. Disk low, glabrous, i.e. epilose as far as not 
covered with stellate scales initially. Ovary 
elongate-conical or -subobovoid, densely stellate- 
lepidote, 6-8 by 2.5-3 mm, rather abruptly 
tapering to the stoutish style, the latter as long as 
the stamens and exclusively stellate-lepidote to the 
top at anthesis, but accrescent and glabrescent 
from the top backwards in later stages; stigma 
broad-obconical, slightly 5-lobed. Capsule thick- 
fusiform, often a little wider in the upper half, 
densely rusty-lepidote, 2-4 cm by 5-6 mm, crowned 
by the style remains. Seeds very narrow, tailed at 
both ends, 3-4 mm. 

Distr. Malesia: New Guinea (Main Range 
from the Wissel Lakes to Mt Victoria, also in the 
Nassau and Bismarck Mts, and on and around 
Mt Saruwaged). 

Ecol. Epiphytic both in mossy forest and on 
fern trees in grassland, terrestrial in more open 
mossy Nothofagus-Podocarpus-Pandamts forest and 
along banks of creeks in open alpine grassland, or 
on edges of Equisetum-Carex bog, locally common 
as undergrowth in openings or native rest clearings 
within the stunted moss forest or on (may be old) 
landslides, occasionally a small shrub growing on 
exposed cliff face, 1500-3615 m, reported from 
Mt Saruwaged between 3600 and 4000 m. Said 
to grow also on limestone. Fl. fr. Jan. -Dec. 

Vern. Kibimume, Chimbu: Masul, djilimp, 
wumbemp, Wahgi: Minj, wambu, Hagen: Togoba, 
kwonem, Chimbu: Goromagua, oganga, Hagen: 
WankI, kume, tikoro, Mendi, rohn, Enga. 

Note. R. beyerinckianiim is conceived here in a 
broad sense, comprising forms with larger leaves 
and apparently longer capsules (3^ cm) in the 
mossy forest zone, and such with smaller and 
narrower leaves, respectively shorter capsules 
(2-2.5 cm) from open country above 2800 m, and 
forms which differ much in the colour of the 
corolla, ranging from pure white over I'ght 
yellow and pink to (the normal) dark red, of 
which the paler colours generally are found in 
specimens with larger leaves, apparently growing 
at lower altitude and/or in shadowy places. The 
species recently has come into cultivation, i.e. 
the smaller leaved and dark red flowering form 
from the Western and Eastern Highlands. 

Nearest to R. phaeochitum of which the fruit 
is not yet known, and possibly to be united with 
that species in the future. 

52. Rhododendron leptanthum F. v. M. Trans. R. 
Soc. Vict. n.s. 1, 2 (1889) 24, in obs.; Sleum. 
Reinwardtia 5 (1960) 83; Blumea 11 (1961) 116; 
ibid. 12 (1963) 98; Jahrb. Rhodod. Ges. Bremen 
(1963) f. 9, 10, non Hayata, 1915. — Fig. 12. 
Small, rather straggling shrub, 0.5-2(-3) m. 



Jan. 1966] 



Ericaceae (Sleumer) 



513 



terrestrial or epiphytic, with thick roots and 
pendulous branches. Branchlets rather slender. 
c. 1.5 mm at the densely red-brown stellate- 
lepidote tips, glabrescent and a little scabrid in 
older parts; internodes 2-12 cm.Le«»r5oppositeor 
mostly in 3-4-merous pseudowhorls. ovate or 
oblong-ovate, apex gradually — shortly acuminate, 
subacute, base — rounded, subcoriaceous, firm, 
equally subdenseiy lepidote on both faces (scales 
small, deeply stellate-lacerate and short-dendroid, 
each on top of minute paler, often nearly white 



flowered. Ferulae membranous, ovate-acuminate, 
outer smaller ones — densely lepidote. inner 
larger ones glabrous, fimbriate, up to 1.2 by 
0.6 cm. Bracteoles linear-spathulate, — glabrous, 
up to 1.5 cm by 2 mm. Pedicels rather slender^ 
densely stellate-lepidote,0.7-1.3 cmbyO.5-0.8mm. 
Calyx in form of a low cup, c. 2.5 mm o, very 
shortly obtusely 5-dentate. Corolla tubular, 
slightly zygomorphous, deep pink to salmon or 
carmine, (2-)2.5-3(-3.4) cm long in all, laxly to 
subdenseiy stellate-lepidote outside except a wide 




Fig. 12. Rliododciidron Icpiunihum F. v. M. Lpiphylic shrublet, showing thick roots, Edic Creek near 
Wau, E. New Guinea, 1850 m (Sleumer 4127) (Sleumer, 1961). 



tubercles beneath, the latter less distinct on the 
upper surface), early glabrescent above, ultimately 
so beneath, margin but slightly or not rcvolutc, 
(2-)3-6.5(-7) by (1.3-)l.5-3(-3.2)cm, midrib im- 
pressed above, prominent beneath, nerves in 5 7 
pairs, rather straight, anastomosing near the edge, 
slightly impressed above, somewhat raised beneath, 
veins laxly reticulate, mostly rather distinctly 
raised beneath, finer reticulation of the veinlcts 
visible above, inconspicuous beneath; petiole 2-4 
(-5) mm. densely lepidote. Umhch l-V-S)- 



marginal part of the lobes, glabrous inside, tube 
slightly curved, 1.5-2 cm, (3-)4-5 mm o at base, 
widened to 7-8 mm at apex, lobes broad-spathu- 
late to suborbicular, (5-)7-IO(-l2) byf>-9(-l2) 
mm. Sianu'iis 10, subequalling the corolla in 
length, slightly unequal; filaments red when fresh, 
narrow-linear, glabrous or with some scattered 
hairs; anthers c. 1 .6 by I mm. Di.\k glabrous. Ovary 
elongate-conical, densely rufous-stcllatc-lepidotc, 
3-5 by 1.5-2 mm. gradually tapering to the style, 
the latter stellate-lepidote to or almost to the top, 



514 



Flora Malesiana 



[ser. I, vol. 6* 



±as long as the stamens; stigma broadly obconical 
capitate. Submature capsule fusiform, curved, c. 
4 cm by 3 mm, densely stellate-lepidote, on fruit- 
pedicel 1.5-2 cm. 

Distr. Malesia: E. New Guinea (Morobe to 
Milne Bay Distr.). 

Ecol. Terrestrial on rocks or dry exposed cliff 
face, or in opening of Castanopsis forest, epiphyte 
on big branches of high trees or fallen trunks in 
mossy forest on Nothofagus, sometimes gregarious, 
1370-2250 m. Fl. Jan. -Oct. 

Vern. Zavi, Gerebi: Movoi. 

53. Rhododendron warianum Schltr, Bot. Jahrb. 
55 (1918) 151; Lane-Poole, For. Res. Papua 
(1925) 130; White & Francis, Proc. R. Soc. 
Oueensl. 39 (1928) 68; Sleum. Reinwardtia 5 
(1960) 83. 

Small epiphytic or terrestrial shrub, creeping 
on the ground, occasionally erect and up to 5 m, 
with lax, spreading branches. Branchlets slender, 
terete, the upper 1-2 internodes densely rusty- 
stel'ate-lepidote, early glabrescent and rather 
smooth elsewhere; internodes 1-10 cm. Leaves 
opposite or mostly in 3-4-merous pseudowhorls, 
elliptic or oblong-elliptic, sometimes broad-elliptic 
or rarely subobovate-elliptic, apex broadly attenu- 
ate, acute to nearly rounded, base ± rounded, 
subcoriaceous, though mostly rather firm, lepidote 
initially on both faces (scales deeply and irregu- 
larly stellate-incised, subsessile or short-dendroid, 
each on top of a minute epidermal tubercle), early 
glabrescent above, rather tardily so beneath, 
rather smooth then as the remaining tubercles 
are not much conspicuous, faintly or not recurved 
at margin, (2.5-)3-5(-7) by (1. 3-) 1.5-3 (-4.4) cm, 
midrib impressed above, prominent beneath, 
nerves 5-7 pairs, very slightly impressed above 
and raised beneath, often ± inconspicuous, as 
are the veins and veinlets; petiole densely stellate- 
lepidote, mostly rather flattened, 2-5 by c. 1 mm. 
Flowers solitary, or in 2-3(-6)-flowered umbels. 
Outer perulae narrowly to broadly ovate-acumin- 
ate, ± densely stellate-lepidote outside, inner ones 
obovate-spathulate and ± glabrous, fimbriate, 
up to 1.5 by 0.6 cm. Bracteoles linear-subspathu- 
late, glabrous, up to 1.2 cm. Pedicels rather 
slender, densely rusty-stellate-lepidote, (1-)1.2-1.5 
cm by c. 0.5 mm. Calyx small, very shortly cup- 
shaped and obtusely 5-lobed, c. 3 mm 0. Corolla 
tubular basally, dilated at the limb, ± distinctly 
zygomorphous, waxy white with pinkish flush or 
pale pink to rose, 3-3. 5 (-4) cm in all, laxly stellate- 
lepidote outside, glabrous inside, tube curved, 
2-2.5 cm long, c. 4 mm at base, dilated to c. 6 
mm at throat, lobes broad-spathulate to sub- 
orbicular, 9-12 by 7-9(-ll) mm. Stamens 10, 
unequal, as long as or slightly longer than the 
tube; filaments red, narrow-linear, glabrous or very 
laxly hairy; anthers broad-oblong, c. 1.5 (-2) by 
0.8 mm. Disk prominent, glabrous. Ovary sub- 
cylindric, densely stellate-lepidote, 5-6 by 1.5 mm, 
gradually attenuate to the style, the latter slender, 
c. 1.5 cm, stellate-lepidote at anthesis at least in the 
lower half, often to 4/5 of the total length, gla- 



brescent distally; stigma short-obconical. 

Distr. Malesia: E. New Guinea (Morobe to 
Milne Bay Distr.). 

Ecol. Epiphyte in Nothofagus forest, also 
terrestrial on limestone precipices or in moss 
forest rest c'earings and heath-like vegetation, 
1065-2600 m, locally common. Fl. Jan. -Dec. 

Vern. Rohn, Enga: Kepilam. 

54. Rhododendron gardenia Schltr, Bot. Jahrb. 

55 (1918) 158; Sleum. Reinwardtia 5 (1960) 83. 
Epiphytic or terrestrial shrub, 1-2.5 m. Branch- 
lets terete, stout, 6-8 mm 0, glabrous and pruino.se 
in the older parts, fugaciously rusty-stellate- 
lepidote at the tips; internodes 15-20 cm. Leaves 
opposite or ± in 3(4-)-merous pseudowhorls at 
the upper 2 nodes, patent, elliptic or oblong- 
elliptic, rarely subobovate-elliptic, apex shortly 
obtusely acuminate, not rarely rounded or a little 
retuse, base broad-attenuate into the petiole to 
rounded, very coriaceous, stiff", subdensely lepidote 
initially on both faces (scales minute, i deeply 
stellate-lacerate and i manifestly dendroid, each 
on top of a paler minute epidermal elevation), 
early glabrescent and rough to the touch by the 
persistent tubercles on both sides, especially 
beneath, slightly or not revolute at the narrowly 
cartilagineous margin, 8-15 by 5.5-9 cm, midrib 
robust in the lower half of the blade, and i im- 
pressed above, very prominent beneath, flattish and 
wrinkled in dry specimens, nerves 8-10 pairs, ± 
straight below, curved and indistinctly anastomos- 
ing before the edge, very slightly prominent on 
both faces or somewhat sunk above, often becom- 
ing inconspicuous towards the edge, no proper 
reticulation visible; petiole lepidote initially, 2-3 
(-4) cm by 2.5-4 mm. Umbels 3-4-flowered. 
Bracteoles linear, laxly hispid-hairy, up to 2.5 cm 
by 2 mm. Pedicels stout, cylindric, densely covered 
with rufous sessile scales, moreover laxly hispid- 
hairy in the upper part and more densely so 
immediately below and at the calyx, 1-1. 5 (-2) cm 
by 2-2.5 mm. Calyx oblique, cup-shaped, lepidote 
and hirsute outside, ± glabrous inside, 8-10 mm 
0, irregularly obtusely 5-8-lobed, lobes ciliate, 
2-3 mm. Corolla funnel-shaped, snow-white or 
suff"used with pink especially at the tube, rather 
fleshy, fragant, 10-11 cm long and across the 
limb, tube straight, c. 5 cm long, 8-10 mm at 
base, gradually dilated to 1.5-2.5 cm at apex, 
glabrous outside, ± densely clothed with retrorse 
whitish hairs in its lower half to 3/4 inside, lobes 
5-8, spreading, broadly obovate-cuneate, 5-6 by 
3-4(-5) cm, glabrous. Stamens 10-16, slightly 
unequal, c. 7 cm long at full anthesis; filaments 
linear and densely patent-hispid-villous below, 
glabrous and filiform distally; anthers linear- 
oblong, 10-12 by 1.5 mm, base obtuse. Disk 
prominent, very densely hairy. Ovary oblongoid- 
cylindrical, densely covered with subappressed, 
yellowish, ± stiff" hairs, which cover completely 
the numerous scales, c. 1.3 cm by 5 mm, rather 
abruptly attenuate to the robust style, which is 
glabrous except some hairs at the very base, c. 
6 cm by 1.5 mm; stigma large, subcapitate. 



Jan. 1966] 



Ericaceae (Sleumer) 



515 



obtusely 5-8-gibbous, c. 4 mm 0. 

Distr. Malesia: New Guinea (Sepik R. region 
and Bele R. Valley 25 km NE. of Lake Habbema). 

Ecol. High epiphyte in forest, but also a 
terrestrial shrub in sandy openings in mossy 
forest on crest of mountain ridge, 1400-2300 m. 
Fl. June-July, Oct. -Nov. 

55. Rhododendron thaumasianthum Sleum. Blumea 
12 (1963) 98. — R. magnificiim Sleum. Rein- 
wardtia 5 (1960) 84, non F. K. Ward, 1935. 

Shrub, apparently epiphytic. Branchlets flattened 
at the ultimate internodes (10 cm), densely covered 
with fugacious rusty-stellate scales, glabrescent. 
Leaves la.xly scattered in the upper half of the 
upper internode, the highest 2 leaves opposite 
or 3 in a pseudowhorl, elliptic orsubovate-elliptic, 
thick-coriaceous and rigid, apex nearly rounded, 
base subtruncate-rounded, very young ones 
lepidote on both sides (scales small, stellate- 
lacerate, dendroid, each on top of a distinct 
epidermal tubercle), early glabrescent above, tard- 
ily so beneath, at full maturity very densely set 
with pale tubercles, and correspondingly very 
rugose or scabrid on both surfaces, edge — 
manifestly revolute, 8-13 by (4-)5-7 cm, midrib 
as thick as the petiole below, gradually narrower 
towards the apex of the lamina, slightly impressed 
above, prominent beneath, nerves c. 8 pairs, rather 
straight and markedly raised below, ± evanescent 
upwards on both sides, no proper reticulation; 
petiole a little flattened, grooved above, 2-2.5 cm 
by 3-4 mm, densely lepidote respectively finally 
verruculose. Umbels 3-5-flowered. Outer perulae 
ovate, obtuse, scabrid by numerous — bullate, 
stiff, short hairs in- and outside, c. 2 by 1-1.5 cm, 
middle ones obovate, 2.5-4 by 2-3 cm and equally 
rough, innermost ones spathulate, clad with 
longer, stiff" and — appressed hairs at the base 
dorsally, minutely tubercled upwards, 4-7 by 
2.5-3 cm. Bracteoles linear-spathulate, c. 5 cm 
long, c. 2 mm wide at the apex, subdensely his- 
pidulous. Pedicels densely rusty-stellate-'epidote, 
not hairy at all, (6->10-l5 by 2-2.5 mm. Calyx 
annular, membranous, very short, wavy, obscurely 
lobcd, lepidote outside, glabrous inside, lobes very 
laxly and shortly whitish ciliate. Corolla funnel- 
shaped, fleshy, colour not known (white or 
pinkish?), laxly stellatc-lepidote at the tube out- 
side, rather densely clad with retrorsc hairs in the 
lower half of the tube inside, c. 9 cm long in all, 
tube rather straight, c. 5 cm long, c. 0.8 cm at 
base, dilated to c. 1 .5 cm at apex, lobes 7, obovate- 
spathulatc, c. 4 by 2-2.5 cm. Stamens 14, 7-8 cm 
long; filaments linear and densely - patently 
subvillous below, more laxly hairy upwards, 
glabrous distally; anthers linear-oblong, 10-11 by 
1.5 mm, base obtuse. Disk densely white-hairy. 
Ovary cylindric and densely hairy and lepidote 
below, constricted upwards in form of a bottle- 
neck and exclusively lepidote there, c. 2 mm long, 
c. 5 mm below, 2.5 mm in the upper part. 
Style seemingly abrupt with the ovary due to the 
thick lepidote "tomcntum" at its very base, but 
really gradually continuing the ovary, completely 



glabrous above the base, blackish when dry, c. 
6.5 cm;, stigma thick-capitate, 7-lobulate. 

Distr. Malesia: New Guinea (Wissel Lakes), 
once found. 

Note. Similar to R. konori, but style completely 
glabrous. 

56. Rhododendron konori Becc. Malesia 1 (1878) 
200; Sleum. Reinwardtia 5 (1960) 85; Blumea 11 
(1961) 116; ibid. 12 (1963) 98; Jahrb. Rhodod. 
Ges. Bremen (1963) f. 14, 38. — R. roverenae 
F. V. M. Vict. Nat. 1 (1884) 101, reimpr. Gard. 
Chron. 22 (1884) 712, f. 127; Descr. Not. 4 (1885) 
12; Stein, Jahresber. Schles. Ges. Vaterl. Cult, 
f.d. Jahr 1885 (1886) 412; Sleum. Reinwardtia 5 
(1960) 119; Blumea 11 (1961) 117. — R. devrie- 
seanum KoORD. Nova Guinea 8 (1909) 185; ibid. 
8 (1912) 876, t. 151; J. J. S. ibid. 12 (1914) 141 
{'devriesianum'); Gibbs, Arfak (1917) 170; Kaneh. 
& Hatus. Bot. Mag. Tokyo 56 (1942) 478. — 
R. devrieseanum ssp. astrapiae Foerster in Fedde, 
Rep. 13 (1914) 224. — R. astrapiae Foerster ex 
ScHLTR, Bot. Jahrb. 55 (1918) 160. — Fig. 13, 14. 
Shrub, (0.5-)l-2(-4) m, or treelet, 2-4(-6) m, 
stems mostly few, 2.5-8 cm 0, terrestrial or 
epiphytic. Branches few, suberect, up to 4 cm 
across. Branchlets robust, terete, 3-9 mm 0, the 
upper 1 or 2 internodes caducously stellate- 
lepidote, lower parts glabrescent, somewhat rough 
to the touch, finally smooth, often pruinose in 
dry specimens; internodes 6-20 cm, sometimes 
with several, much reduced scattered early 
caducous leaflets. Leaves scattered in the upper 
third of the upper 1 or 2(-3) internodes, rather 
close together, sometimes crowded into 3-5- 
merous pseudowhorls, elliptic or oblong-elliptic, 
more rarely oblong or ovate-oblong, apex shortly 
subacuminate-attenuate, rather blunt, often round- 
ed, base broadly attenuate to rounded, more 
rarely subcordate, coriaceous and stiff", dark green 
above, yellowish green beneath, densely covered 
with a rufous detersile tomentum on both faces 
when young (scales deeply stellate-lacerate, ± 
dendroid, each on top of a paler permanent 
epidermal tubercle), ..: glabrescent at maturity 
on both sides, the midrib excepted, which remains 
lepidote for a long time, !: rough or even scabrid 
to the touch on both sides, margin slightly or not 
revolute, (7-)10-18(-21 ) by (4-)5-8(-9, some- 
times up to 12) cm, midrib as stout as the petiole 
at base, rather flat or even slightly impressed in 
its middle above, strongly prominent in the lower 
half beneath, nerves (7-)8-10(-12) pairs, rather 
straight and marked below, curved and less visible 
towards the edge, :. anastomosing, slightly im- 
pressed above in fully mature leaves, somewhat 
raised beneath, often rather obscure on both faces, 
reticulation J- inconspicuous; petiole i semi- 
terete, densely lepidote, rough in later stages, (1-) 
1.5-3(-3.5) cm by (2-)3^(-5) mm. Umbels (3-) 
5-8(-l 2) flowered. I'lower buds ovoid, 5-8(-ll) 
by 3-4(-5 )cm. Outer perulae ovate, inner ones 
oblong-ovate to obovate-spathulate. :•. scabrid 
outside by minute stiff bullate hairs or their 
remaining warty bases, up to 7.5 by 3 cm, often 



516 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 13. Rhododendron konori Becc. a. Habit, X 1/3, b. flower, X 1, c. ovary and style, x 2/3, rf. stamen, 
X 2/3 (a-rf Cruttwell 1010 and a drawing made by him from a fresh specimen). 



Jan. 1966] 



Ericaceae (Sleumer) 



517 



of a dull carmine colour outside, sometimes more 
pinkish. Bracteoles linear-spathulate, often lacerate 
distally, subdensely hairy, white, up to 5 cm by 
l-2(— 4) mm. Pedicels densely rufous-stellate- 
lepidote, and whether or not hairy below the 
calyx. (4-J6-15 by 2.3 (^) mm. Calyx rather 
manifestly oblique, patelliform, shortly (1-2 mm) 
irregularly and obtusely (6-)7-lobed or- dented, 
one or the other lobe sometimes up to 1.5 cm, 
stellate-lepidote dorsally and often set with some 
stiff hairs, whitish ciliate or fimbriate. 6-9(-10) 
mm across. Corolla tubular below, funnelform- 
expanded towards the lobes, fleshy, pure white, 
often pinkish in later stages especially distally, or 
sometimes pale to rather deep pink from the be- 
ginning, or pinkish only at the sinuses to the lobes, 
with a sweet fragrance of carnations especially at 
night. (8-)9-14(-16) cm long in all when dry, 
12-16(-17, rarely up to 19) cm long when fresh, 
variable in length probably due to growing going 
on during anthesis, provided with some sparse 
stellate scales at the tube, or completely glabrous 
outside, ~ densely covered with whitish retrorse 
hairs in the lower half of the tube inside, tube 
cylindric, or sometimes slightly 7-angled, (6-)7-8 
(-11) cm long, 0.6-1 (-1.5) cm o at base, widened 
to (l-)1.5-2 cm at apex, lobes 6 or 7, spreading, 
obovate-spathulate to suborbicular, often sinuous 
distally, (3.5-)4-5(-7) by (2.2-)2.5^(-5) cm in 
dry, 5-8 by 4-6 cm in fresh specimens. Stamens 
12 or 14, unequal, mostly exserted for 1-2 cm 
from the tube; filaments densely hairy and linear 
below, glabrous and almost filiform in the upper 
third; anthers orange or yellowish, linear-oblong, 
(8-)10-n(-13, rarely up to 16) by c. 1.5 mm. 
Disk densely white-hairy. Ovary subcylindric, 
somewhat attenuate towards the base, densely clad 
with yellowish, stiff, — forward directed hairs 
which cover numerous scales, 10-15 by 4— 5(-7) 
mm, rather abruptly attenuate to the thick style, 
the latter white to reddish, 4-6(-8.5) cm first, 
accrescent rapidly during and still after anthesis, 
attaining almost the length of the corolla in later 
stages, - densely hairy and laxly stellate-lepidote 
in the lower half, gradually more lepidote and 
less hairy in the following '4, finally exclusively, 
:n caducously lepidote for some mm, glabrous in 
the distal '/s ^o V« part. Stigma yellow, capitate, 
shortly 6- or 7-Iobed, 5-7 mm 0. Capsule ± 
fusiform. 6-9(-12) by 1.2-I.5(-I.8) cm, widest a 
little below the middle, gradually attenuate to the 
apex, rough by the remaining hairs and numerous 
minute tubercles, topped by the persistent style, 
which attains (4.5-)5.5-9 (sometimes even 1 5 ) cm, 
the stigma lobes up to 2 by 1 mm then. Fruit- 
pedicel 2-3 cm by 3-4 mm. Capsule finally splitting 
from the top in form of ligneous, irregularly 
twisted walls, of which the outer part has gone 
before. Seeds narrow-fusiform, flattened, 6-8(-10) 
mm including the tails at both ends, c. 0.03 mm 
wide. 

Distr. Malesia: New Guinea (Tamrau, Ncttoti 
and Arfak Mts in the Vogclkop Peninsula, from 
Mt Cienofa near Kaimana, WisscI Lakes area, 
Mcllwig and Star Mts, Upper .Scpik R. region. 



Western and Eastern Highlands and on and 
around Mt Saruwaged, in the SE. known as far 
as the Maneau Range) and Fergusson I. 

Ecol. Epiphytic in montane forest, terrestrial as 
substage in Nothofagus and Castanopsis-Quercus 
forest and forest edge of clearings, in open fern 
regrowth (there found often with smaller flowers 
and fruits), on burned open summits, also on 
peat bogs, swamps or marsh, generally on poor 
clayey soil or humus, 750 -2400 (-2500) m. Fl. 
Jan. -Dec. 

Vern. Soksaikokohugwe, Arfak, tetjai, Hattam, 
wagadei, warade, Kapauku, wennieran, Arguni, 
wayam, Maneau Ra. 

Note. R. konori has been recently introduced 
into horticulture from various places in New 
Guinea, especially from the Arfak Mts, where it 
grows in abundance around the Anggi Lakes. 
It apparently hybridizes there locally with 62. 
R. asperum, which is often found on the same 
places. Such hybrids have smaller, 6- or 7-merous 
corollas and more ovate and more shortly petioled 
leaves than are found in typical R. konori. It 
is possible that most or all forms of R. konori in 
the Arfak Mts with such smaller corollas and 
smaller fruits are due to introgression of R. 
asperum, though the leaves may be typical R. 
konori. In one place with abundant R. konori and 
248. R. laetum, I found a fruiting specimen which 
was apparently intermediate in leaves (apex more 
pointed, midrib red: laetum characters), in the 
epidermal tubercles of the leaves (less prominent 
than in R. konori) and in the fruit (shorter, and 
more densely hairy than in R. konori). 

57. Rhododendron phaeopeplum Sleum. Rein- 
wardtia 5 (1960) 92; Blumea 12 (1963) 99; Jahrb. 
Rhodod. Ges. Bremen (1963) f. 40. 

Terrestrial shrub. Branchlets terete, 3-5 mm 0, 
densely rusty-stellate-lepidote at the upper inter- 
nodes, glabrescent otherwise and densely papillate- 
verruculosc, rather smooth to the touch, often 
pruinose in dry specimens; internodes 3-10 cm. 
Leaves in 3-5-merous pseudowhorls at the upper 
1-2 nodes, elliptic or ovate-elliptic, apex broad- 
attenuate, obtuse to nearly rounded, apiculate at 
the very top, base generally obtuse or rounded, 
not rarely slightly cordate, thick-coriaceous, very 
stiff, erecto-patent, very young ones densely rusty- 
or brown-lepidote (scales irregularly stcllate- 
lacerate, nearly squamular, the segments ending in 
something like a hair, each subsessile on top of an 
epidermal tubercle, which persists for a long time 
beneath), mature ones olivaceous, dull, ± 
glabrescent, often rather smooth above, rusty- 
brown and covered with a thick dctcrsile tomentum 
of star-shaped scales (no proper hairs!) beneath, 
finally glabrescent also there and very densely 
tubercled and scabrid, distinctly revolute at the 
margin in dry specimens, 4-7(-10) by (2-)2.5-4.5 
(-5) cm, midrib f flat, dilated and a little raised 
in the lower part, very slightly impressed in the 
middle, evanescent towards the top of the lamina 
above, very stout below as a continuation of the 
petiole, gradually decreasing upwards beneath, 



518 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 14. Rhododendron konori Becc. Mt Sensenemes, 2550 m, Anggi Gigi Lake, Arfak Mts, NW. New 

Guinea (Sleumer, 1962). 



Jan. 1966] 



Ericaceae (Sleumer) 



519 



nerves 8-10 pairs, lower ones irregular, upper 
ones rather straight. — raised above, nearly 
inconspicuous beneath in fully mature leaves, 
reticulation absent; petiole somewhat flattened 
above, lepidote, (5-)8-12(-20) by 1.5-3 mm. 
Umbels 4— 6-flowered. Ferulae rigidly membrana- 
ceous, outer ones ovate, — obtuse, laxly lepidote 
dorsally and puberulous below, c. 2 by 1.5 cm, 
inner ones oblong-spathulate, — densely shortly 
subsericeous-hirsutulous on both faces, ciliate, up 
to 4 by 1.5 cm. Bracteoles linear to linear-spathu- 
late, very laxly pubescent, up to 4 cm by 1-3 mm. 
Pedicels very densely rusty-stellate-lepidote, 
whether or not with some hairs immediately 
below the calyx, 1.2-2.3 cm by 1.5-2 mm at 
anthesis. Calyx oblique, disk-shaped, 4-5 mm 0, 
wavy and shortly obtusely 5-6-lobed, ciliate. 
Corolla narrowly funnel-shaped, scented, white 
or pinkish, 6-7 (rarely up to 8) cm long in dry, 
(7-)8-9 cm in fresh specimens, tube very laxly 
stellate-lepidote or entirely glabrous outside, 
densely set with longish, retrorse hairs in the lower 
34 inside, ~ straight, 4—4.5 cm long, 5-8 mm 
at base, c. 1.5 cm at throat, lobes 6 or 7, spreading, 
broadly obovate-spathulate, (2-)3.5(-4.5) by 1.8- 
2.3(-3) cm. Stamens 12 or 14, unequal, -}z equal- 
ling the tube; filaments linear, densely patently 
yellowish hairy below, more laxly so and slender 
upwards, glabrous and filiform at the top; anthers 
oblong, 6(-7) by 1 mm, base obtuse. Disk low, 
laxly to densely pubescent. Ovary nearly fusiform, 
gradually attenuate to the style, c. 10 by 3.5 mm, 
very densely clothed with yellowish, forward 
directed, stiff hairs, which ± cover the scales. 
Style :r as long as the stamens, thick and lon- 
gitudinally 12- or 14-ribbed below as is the ovary, 
densely hairy and lepidote there (the scales well 
visible), upwards almost exclusively clad with 
dark rusty or brown stellate scales and a few or 
practically no hairs for c. 2 mm, more slender and 
quite glabrous for about 10 mm below the thick- 
capitate, 6-7-gibbous stigma. Capsule fusiform, 
densely hairy and lepidote, (4-)5-6 by I cm, 
crowned by the 4—5 cm long style. 

Distr. Malesia: New Guinea (Wissel Lakes 
region and Arfak Mts, above the Warjori R.). 

Ecol. Heathland and open, fern-covered crest 
within the Nothofai^us-Castanopsis forest, 1300 m. 

Note. R. phaeopepliim is something like a small 
edition of R. konori, and possibly in the Wissel 
Lakes region a hybrid of R. konori with a related 
species of the same subsection with smaller 
flowers, may be 72. R. rappardii. Above the 
Warjori R., part of the specimens I found matched 
the type from the Wissel Lakes almost completely, 
but others had petioles up to 2 cm long, larger 
and usually less cordate leaves, and less densely 
hairy pcrulac. All specimens had 6- or mostly 
7-merous, pure white (lowers. Quite a number of 
natural, : intermediate hybrids with the 5-mcrous, 
orangc-rcd flowered 255. R. zoelleri. which is 
abundant on the same place, showed pale pink, 
5- and 6-mcrous corollas on the same specimen, 
longer, more acuminate, thinner, • glossy leaves, 
on which the epidermal tubercles arc much less 



distinct than in the species of subsect. Phaeovireya 
in general, and longer, more slender pedicels. A 
few specimens of R. zoelleri, typical besides white 
corollas with yellowish hue all over or at least at 
the tube, found in the same place, are probably due 
to a slight introgression of R. phaeopeplum. Up 
to now, such "white" R. zoelleri had not yet been 
found despite the fact that it is one of the wide- 
spread and much collected species in New Guinea. 

58. Rhododendron opulentum Sleum. Reinwardtia 

5 (1960) 85. 

Epiphytic shrub, 1 m. Branchlets subterete, tips 
densely rufous-stellate-lepidote,olderpartsglabres- 
cent and scabrid by minute tubercles; internodes 
10-20 cm. Leaves laxly alternate in the upper half 
of the apical internodes, oblong-elliptic or broad- 
oblong, coriaceous, apex short-acuminate, sub- 
acute or subobtuse, base very broadly attenuate to 
rounded, the very young ones ± densely rufous- 
lepidote above and beneath (scales minute, deeply 
stellate-incised, short-dendroid, each on top of a 
minute pale epidermal tubercle, the proper scale 
going early), mature ones glabrescent except the 
midrib, densely minutely tubercled and very rough 
to the touch, margin but slightly wavy or revolute, 
6-10 by 3-4 cm, midrib rather stout, flattened, a 
little raised and sunk in the lower part above, well 
prominent beneath, nerves c. 10 pairs, rather 
dense together, irregular, ± straight below, 
anastomosing near the edge, slightly prominent 
on both surfaces, veins laxly prominent-reticulate, 
finer reticulation of the veinlets rather obscure; 
petiole ± flattened, c. 1 cm by c. 2.5 mm, densely 
lepidote, rough in later stages. Umbels 7-8- 
flowered. Middle and inner perulae spathulate, 
densely whitish subsericeous-hirsute in- and out- 
side, up to 4.5 by 2 cm. Bracteoles linear, very 
densely subappressedly whitish hairy, up to 3 cm 
by 2 mm. Pedicels densely appressed-hairy and 
laxly fugaciously stellate-lepidote, 8-10 by c. 1.5 
mm. Calyx very oblique, annular, obscurely 5-6- 
lobed or -dentate, densely hairy and lepidote 
dorsally, c. 4 mm 0. Corolla elongate-tubular 
below, expanded at the 5 lobes, fragrant, pink 
at the tube, shading to white at the lobes, 9-10 
cm long in all, 4-5 cm at the limb, glabrous out- 
side, rather densely clothed with longish retrorse 
hairs at the tube up to almost the throat inside, 
tube cylindric, 6-7 cm long, 4-5 mm o at base, 
gradually dilated to c. 8 mm at apex, lobes broadly 
obovatc to suborbicular, 2.5-3 by 1.8-2.5 cm, 
glabrous. Stamens 10, exceeding the tube for c. 
I cm; filaments linear and densely set with retrorse 
hairs below, more laxly hairy upwards, filiform and 
glabrous in the ultimate '/^; anthers elongate- 
oblong, base obtuse, c. 6 by 1.2 mm. Disk white- 
hirsute. Ovary clongate-cylindric, very densely 
stellate-lepidote and i laxly short-hairy (the 
hairs i hidden by the scales), 10-12 by c. 2.5 mm, 
gradually attenuate to the style, the latter 5-6 
cm long, rather thick below, very densely stellate- 
lepidote and epilose to the lower 3-4 cm. more 
slender and completely glabrous for the rest; 
stigma large, capitate, manilcslly 5-gibhous. 



520 



Flora Malesiana 



[ser. I, vol. 6* 



Distr. Malesia: W. New Guinea (Bernhard 
Camp, Idenburg R.), once found as a frequent 
epiphyte on large trees in mossy forest at 1800 m. 
Fl. Jan. 

59. Rhododendron spondylophyllum F. v. M. Trans. 
R. Soc. Vict. n.s. 1, 2 (1889) 23; Sleum. Rein- 
wardtia 5 (1960) 86. 

Shrublet. Branchlets slender, 1-1.5 mm 0, tips 
densely lepidote, elsewhere densely finely verrucose 
and scabrid; internodes 3-6 cm. Leaves in 3-4- 
merous pseudowhorls at the upper 1-2 nodes, 
ovate, apex shortly obtusely acuminate, base 
slightly cordate, ± sessile, coriaceous and stiff at 
maturity, glabrous and densely verruculose above, 
densely lepidote for a long time beneath at least 
near the midrib (scales coarsely and irregularly 
incised at the margin, dendroid, each on top of a 
small permanent lighter tubercle), finally glabres- 
cent and very rough beneath too, margin sub- 
revolute, (1.6-)2-3 by 1.2-1.7 cm, midrib mani- 
festly grooved above, prominent beneath, nerves 
± obscure; petiole 0.5-1 by 1 mm. Umbels 3- 
flowered. Pedicels densely set with dark brown, 
short-dendroid, star-shaped, squamular scales, 6-8 
by 0.7 mm. Calyx reduced to a narrow membrane, 
lepidote and very laxly hairy dorsally, obscurely 
5-lobed, c. 2.5 mm 0. Corolla cylindric below, 
expanded at the limb, somewhat zygomorphous, 
2.5-2.8 cm in all, colour not known (probably 
pink or red), tube 1.8-2 cm long, laxly lepidote 
and sometimes very sparsely hairy outside, laxly 
± retrorsely hairy in the lower % inside, c. 4 mm 
at base, c. 6 mm at throat, lobes broadly 
spathulate to suborbicular, lepidote outside in the 
lower half, glabrous otherwise, ± 8 mm 0. 
Stamens 10, nearly as long as the corolla; filaments 
linear, laxly patent-hairy in the lower half; anthers 
broad-oblong, 2 by 1 mm. Disk almost glabrous. 
Ovary conical, yellowish tomentose (hairs covering 
the scales), c. 4 by 2 mm, gradually attenuate to 
the rather slender style, which is densely yellowish 
hairy (with sparse scales) in its lower 2/3 to 3^, 
glabrous distally, and up to 1.2 cm long; stigma 
rather small, turbinate. 

Distr. Malesia: New Guinea (Mt Knutsford, 
Owen Stanley Range), once found. 

60. Rhododendron melantherum Schltr, Bot. 
Jahrb. 55 (1918) 152; Sleum. Reinwardtia 5 
(1960) 86. 

"Epiphytic shrublet, c. 50 cm, much branched. 
Branchlets erecto-patent, densely foliate, terete, 
densely rufous-lepidote in the young parts; inter- 
nodes short. Leaves opposite, elliptic, subsessile, 
apex obtusely acuminate, base cuneate, ± laxly 
rufous-stellate-lepidote beneath, glabrescent above, 
2.4-4 by 1.2-1.7 cm; petiole rufous-lepidote, 2-4 
mm. Flowers solitary. Pedicels slender, densely 
rufous-lepidote, c. 1.5 cm. Ca/yx rufous-lepidote, 
very shortly 5-lobed, c. 2.5 mm 0. Corolla tubular, 
manifestly zygomorphous, dark blood red, slightly 
curved and dilated to c. 1 cm at apex, sparsely 
puberulous outside, glabrous inside, 2.7 cm long 
in all, lobes obovate-suborbicular, truncate-obtuse. 



c. 7 mm 0. Stamens 10, c. 2.3 cm; filaments fili- 
form, sparsely hairy; anthers oblong, blackish red, 
2.5 mm. Ovary oblongoid, slightly 5-grooved 
lengthwise, densely rufous-tomentellous. Style fili- 
form, shortly finely hairy, almost attaining the 
stamens in length; st'gma capitate, 5-gibbous." 

Distr. Malesia: New Guinea (Schrader Mts), 
once found in mountain forest at c. 2070 m. Fl. 
June. 

Note. No type material preserved; description 
after the original diagnosis. 

61. Rhododendron cyatheicolum Sleum. Rein- 
wardtia 5 (1960) 86. 

Epiphytic shrub, 30-40 cm; branches rigid. 
Branchlets terete, 2-5 mm 0, tips stellate-lepidote, 
older parts glabrescent and densely verruculose; 
internodes 3-8 cm. Leaves in 3-5-merous pseudo- 
whorls at the upper 2 nodes, ovate, subsessile, 
apex broadly attenuate, obtuse, base rounded or 
subcordate, coriaceous, rigid, the very young ones 
very densely rufous-stellate-lepidote (scales rather 
small, subsessile or short-dendroid, very early 
caducous), yellowish beneath, scabrid by numerous 
minute whitish tubercles beneath, less so or rather 
smooth above, edge ± manifestly revolute, 3-4.2 
by 2-2.5 cm, midrib slightly sunk above, promi- 
nent beneath, more stoutish towards the base, 
nerves c. 6 pairs, rather inconspicuous above, 
somewhat raised beneath, reticulation not visible; 
petiole 1-2 by c. 2 mm. Umbels 4-5-flowered. 
Perulae membranous, outer ones ovate, apiculate, 
with a prominent dorsal rib, lepidote at the apex 
outside, glabrous otherwise, fimbriate, inner ones 
ovate-oblong to obovate or truncate-spathulate, 
glabrous, up to 1.5 by 0.8 cm. Bracteoles narrow- 
linear below, subspathulate at the apex, up to 1.5 
by 0.1 cm, laxly hairy. Pedicels densely stellate- 
lepidote, not hairy at all, 1.5-2 cm by 0.8 mm. 
Calyx disk-shaped, very shortly obtusely 5-lobed, 
wavy, lepidote and laxly hairy dorsally, ciliate, 
4—5 mm 0. Corolla broad-tubular, somewhat 
zygomorphous, pink, 4-4.5 cm, minutely stellate- 
lepidote and with some hairs here and there out- 
side, the margin of the lobes excepted, laxly sub- 
patently hairy at the tube inside, tube a little 
curved, c. 3 cm long, 5-6 mm at base, dilated to 
10 mm at apex, lobes obovate, erect-spreading, 
1-1.2(-1.3) by 0.8-1 cm. Stamens 10, ± as long 
as the corolla; filaments linear below, rather 
densely hairy in the lower third, glabrescent and 
filiform distally; anthers subobovate-oblong, 3 by 
1 mm, base very shortly biappendiculate. Disk 
glabrous. Ovary ovoid-conical, densely yellowish 
hairy (hairs ± strigose, directed forward, ± 
covering the scales), 5-6 by 3-4 mm, rather 
abruptly attenuate to the style, the latter thickish, 
densely hairy and laxly lepidote below, upwards 
more laxly hairy and lepidote, the ultimate 
glabrous 6-8 mm excepted, as long as or a little 
longer than the corolla; stigma capitate. 

Distr. Malesia: New Guinea (Wharton Range, 
Murray Pass), once collected; common epiphyte 
on grassland tree-ferns, at 2840 m. Fl. June- 
Sept. 



Jan. 1966] 



Ericaceae (Sleumer) 



521 



62. Rhododendron asperum J. J. S. Nova Guinea 12 
(1914) 137. t. 34: Kaneh. & Hatus. Bot. Mag. 
Tokyo 56 (1942) 478; Sleum. Reinwardtia 5 (1960) 
87; Blumea 12 (1963) 99; Jahrb. Rhodod. Ges. 
Bremen (1963) f. 36. — Fig. 15. 

Terrestrial shrub, 0.5-1 (-2) m. Branchiets 
stoutish, tips flattened or obtusangular, densely 
steliate-lepidote, terete in the lower parts, 2-4 
mm o, densely verruculose and rough to the touch; 
internodes 3-10 cm. Leaves in dense 5-10-merous 
pseudowhorls at the upper 2-3 nodes, ovate or 
elliptic or suborbicular.sometimesobovate-elliptic, 
subsessile, apex obtusely or mostly rounded, base 




Fig. 15. Rhododendron asperum J. J. .S. Mt Sense- 

ncmds, 2500 m, Anggi Gigi Lake, Arfak Mts, NW. 

New Guinea (Sleumer & Vink 4303) (Sleumer, 

1962). 



broadly rounded or mostly shortly cordate, 
coriaceous, rigid, greyish green to dark green 
above, rusty below, densely slellatc-lcpidotc on 
both faces initially (scales small, deeply incised, 
each subsessile on top of a small persistent elevated 
light point), early glabrcsccnt above, later so 
beneath, scabrid by numerous minute tubercles 
on both faces especially beneath, edge manifestly 
recurved in dry specimens, (2-)3-5.5(-7) by ( 1 .5-) 
2-4.5(-5.5) cm, midrib a little sunk above, 
strongly obtusely prominent in its lower half 
beneath, gradually less so towards the apex of the 
blade, nerves in 5-9 pairs, spreading irregularly. 



faintly anastomosing before the edge, very slightly 
impressed in fully mature leaves above, somewhat 
prominent beneath, veins — inconspicuous; 
petioles somewhat flattened, 1-2.5 by 1-2 mm. 
Umbels 5-9-flowered. Ferulae membranous, outer 
ones smaller, ovate to suborbicular, shortly 
subulate-acuminate, ^i lepidote and puterulous 
dorsally at the apex, inner ones larger, obovate to 
spathulate, ~ glabrous, up to 2 by 1 cm. Bracteoles 
linear, laxly hairy or nearly glabrous, up to 2 cm 
by 1 mm. Pedicels red, densely steliate-lepidote, 
(l.5-)2-3 cm by c. I mm. Calyx — oblique, in 
form of a flat cup, densely lepidote outside, very 
shortly obtusely 5-lobed, c. 4 mm 0. Corolla funnel- 
shaped, — regular, white or pink (sometimes 
deeply pink, or tube of a more intense colour than 
the lobes), scented, (3.5-)4-4.5 cm long and wide 
at the limb, laxly lepidote outside at the tube and 
the base of the lobes, ± sparsely hairy inside at the 
tube, glabrous otherwise, tube (2-)2.3-2.8 cm, 
c. 4 mm at base, gradually widened to 1 cm at 
apex, lobes much spreading, suborbicular, 1.5-1.9 
cm o. Stamens 10, somewhat exserted from the 
throat; filaments linear, — laxly hairy in the lower 
1/2 to 2/3, glabrous above; anthers oblong, (2.5-) 
3-4 by I mm. Disk low, hirtellous at the upper 
margin only. Ovary cylindric-conical, densely 
covered with whitish ± spreading hairs and stellate 
scales (these ± hidden between and below the 
hairs), gradually attenuate to the style, which is 
rather slender, clad with spreading hairs and some 
scales in the lower V2 to 2/3, glabrous apically, 
2-2.3 cm long; stigma capitate. Capsule sub- 
cylindric, densely rusty steliate-lepidote and hairy, 
1.5-1.8 by 0.6-0.8 cm. 

Distr. Malesia: New Guinea (Arfak Mts). 

Ecol. In heath formation on ridge, edge of 
degradated Nothofagus forest, or burnt open 
summit, on poor clayey soil, granite and quartzite, 
2150-2750 m. Fl. Jan., April, July. 

Note. Forms local hybrids with 248. R. laeium, 
characterized by glossy, shortly acuminate leaves 
with less distinct epidermal tubercles, perulae 
densely hairy dorsally and c. 2.5 cm long, pedicels 
lepidote and pubescent, anthers c. 4 mm long, 
ovary densely hairy; corolla in all shades from 
pale greenish yellow to pink salmon (then often 
with yellow or greenish hue in the lower part of 
the tube). 

63. Rhododendron rhodochroum Sleum. Reinward- 
tia 5 (I960) 87. 

Shrub. Branchiets 5-7 mm 0, terete, obtus- 
angular at the upper internodes, tips apparently 
very early glabrescent, very densely verruculose 
and rough; internodes 4-13 cm. Leaves scattered 
and dense together in the upper third of the 
upper 3-4 internodes, or in 3-5-merous pseudo- 
whorls, oblong-elliptic, apex broadly attenuate 
and obtuse, sometimes apiculate, base slightly 
cordate, coriaceous, rigid, rufous-stellatc-lcpidotc 
on both faces when very young (scales minute, 
subsessile. each on top of a paler permanent 
epidermal tubercle), early glabrescent above, more 
tardily so beneath, finally glabrescent there too and 



522 



Flora Malesiana 



[ser. I, vol. 6 



very rough to the touch on both sides, ± revolute 
at the margin in dry specimens, (3-)4-7.5 by 
1.6-3.5 cm, midrib somewhat impressed in its 
lower, :~ flat in its distal part above, continuing 
the stout petiole beneath and thus very prominent 
below, less so apicaliy, nerves 8-lOpairs, spreading, 
faintly raised on both sides, i evanescent towards 
the edge, often inconspicuous beneath as are the 
veins; petiole 2-4 by 1.5-2.5 mm. Umbels 3-8- 
flowered. Ferulae membranous, ovate-oblong to 
obovate, 4: obtuse, outer ones r" lepidote dorsally, 
inner ones glabrous, up to 1 .5 by 0.8 cm. Bracteoles 
linear, glabrous, up to 1.5 cm. Pedicels densely 
rufous-stellate-lepidote, c. 2 cm by c. 1 mm. 
Calyx disk-shaped, dz reflexed, densely lepidote 
dorsally, lobes subobtuse, 1-1.5 mm, not ciliate. 
Corolla tubular, somewhat curved and zygo- 
morphic, pink, subdensely stellate-lepidote at 
the tube and the lower part of the lobes outside, 
laxly hairy at the tube inside, (4.5-)5-5.5 cm in 
all, tube 3.3-4 cm long, 5-6 mm at base, 9-10 
mm at apex, lobes erecto-patent, broadly obovate, 
1-1.5 by 1-1.3 cm. Stamens 10, unequal, c. 1 cm 
exserted from the tube; filaments linear below, 
subdensely hairy at the base, laxly so in the middle, 
glabrous and filiform distally; anthers obovate- 
oblong, obtuse at the base, 3-3.5 by 1.3 mm. 
Disk glabrous below, hairy at the upper margin. 
Ovary ovoid-conical, densely hairy and stellate- 
lepidote, 6-8 by c. 3 mm, rather abrupt with the 
style, the latter densely patently hairy and laxly 
lepidote in the lower half, glabrous upwards, 
as long as the stamens or slightly exceeding 
them; stigma thick, oblique, capitate, distinctly 
5-gibbous. 

Distr. Ma/raa: W. New Guinea (Lake Habbe- 
ma), at 3225 m, common in more open mossy 
thickets on peat ridges, associated with R. haema- 
tophthalmiim. Fl. Aug. 

64, Rhododendron giUiardii Sleum. Reinwardtia 
5 (1960) 88. 

Shrub; epiphytic or terrestrial? Branchlets terete, 
slender, 2 mm 0, nodose, tips densely rufous- 
stellate-lepidote, very early glabrescent elsewhere, 
early covered with greyish bark; internodes 1.5- 
2.5 cm. Leaves opposite or in 3-merous pseudo- 
whorls at the upper node, broad-elliptic, apex 
almost rounded, apiculate, base very broadly 
attenuate or rounded, subcoriaceous, immature 
ones densely rufous-lepidote on both faces (scales 
it deeply and irregularly incised at the marginal 
zone, each sessile on top of an epidermal tubercle), 
mature ones glabrescent, rather smooth and 
shining above, lepidote for a long time, finally 
glabrescent and somewhat rough by the permanent 
tubercles beneath, the very edge slightly revolute, 
3-5 by 2-3.3 cm, midrib a little sunk above, 
prominent beneath especially in the lower part, 
nerves c. 8 pairs, i patent and parallel to each 
other, curved-anastomosing before the edge, 
minutely impressed above, slightly raised beneath, 
veins not much conspicuous, or only beneath so 
in the glabrescent parts of the lamina; petiole 
lepidote, 5-6 by c. 1 mm. Umbels 4— 6-flowered. 



Ferulae and bracteoles not known. Fedicels very 
densely stellate-lepidote, 2-2.8 cm by c. 0.5 mm 
at anthesis. Calyx minute, disk-shaped, obtusely 
shortly 5-lobed, c. 2.5 mm 0. Corolla (colour not 
known, possibly white or yellowish) tubular in the 
lower, 5-lobed and expanded in the upper half, 
2-2.3 cm in all, densely rufous-stellate-lepidote 
at the tube and the base of the lobes outside, 
laxly set with retrorse hairs at the tube inside, 
glabrous otherwise, tube c. I cm long, 5-pouched 
and c. 5 mm at base, somewhat contracted above, 
but again 5-6 mm at apex, lobes obovate- 
spathulate, sometimes a little retuse, 1-1.3 by 
0.7-0.8 cm. Stamens 10, nearly as long as corolla; 
filaments linear and densely h patently whitish 
hairy below, more slender and glabrous apicaliy; 
anthers obovate-oblong, c. 2 nun. Disk very 
shortly pubescent at the upper margin, glabrous 
otherwise. Ovary elongate-conical, r. 4 by 2 mm, 
densely subappressed-hairy and lepidote (the 
scales well visible among the hairs), gradually 
tapering to the style, the latter rather slender, ± 
densely patent-hairy and less densely lepidote to 
the lower 2/3, glabrous for the rest; stigma broad- 
capitate. 

Distr. Malesia: E. New Guinea (Western High- 
lands: Mt Hagen), once found near timber line, 
2285-3655m. f/. July. 

Note. Remembers R. macgregoriae in the size 
and the form of leaves and flowers, but is com- 
pletely distinct by the peculiar stellular scales on 
distinct tubercles, which are typical for subsect. 
Phaeovireya. 

65. Rhododendron bullifolium Sleum. Blumea 12 
(1963)93. 

Erect, few-stemmed shrub, 3-4 m, or treelet, 
4-6 m, trunk 2-3 m by 5 cm. Branchlets terete, 
3-10 mm 0, covered with a rather persistent, 
though detersile tomentum of rufous stellate 
scales; internodes 6-20(-26) cm. Leaves in (4-) 
6-merous pseudowhorls at the upper 2 or 3 nodes, 
elliptic or mostly ovate-elliptic, more rarely 
broad-elliptic or ovate, slightly inequilateral, apex 
very shortly subacuminate, obtusate or rounded, 
base on the whole rounded, coriaceous, dark green 
and glossy above, paler and dull beneath, young 
ones completely covered with a thick tomentum 
of stellate rufous scales, mature ones glabrescent 
early above, less quickly so beneath especially 
along midrib and nerves (scales + dendroid, 
marginal zone + deeply incised or lacerate, the 
small centre deepened or prolonged downwards 
into a kind of foot), (8-) 10-1 8 (-22) by 4.5-12 
(-13) cm, bullate both in the fresh and dry state, 
margin flat, midrib as thick as the petiole below, 
and hardly prominent in the lower, ± impressed 
in the upper part above, nerves irregular, 10-14 
pairs, curved anastomosing before the edge, much 
impressed above, prominent beneath as are the 
veins, veinlets more distinct and reticulate to- 
wards the leaf margin; petiole densely stellate- 
lepidote initially, 2.5-5 cm by 2.5-3.5 mm. Im- 
mature buds f. 2 by 2 cm, each containing 10-12 
flowers; perulae ovate, outer ones subacuminate. 



Jan. 1966] 



Ericaceae (Sleumer) 



523 



inner ones obtuse, substrigose-sericeous, on both 
faces and lepidote dorsally. Immature corolla 
densely lepidote at the tube outside, glabrous at 
the lobes. Fruit-pedicel densely lepidote, c. 2.5 cm 
by 1.5 mm. Young //-t/Zr 2.5 cm long, fusiform, 
densely set with scales and pale stoutish hairs, as 
is the style, of which only the lower part (for c. 
3 cm) is known. Twisted valves of the mature 
fruit (3-)4-5.5 cm. 

Distr. Malesia: New Guinea (Arfak Mts, Mt 
Sarumot, Anggi Gigi Lake), once found. 

Ecol. Undershrub or -tree in transitional zone 
of Casranopsis to Nothofagus forest, at 2050-2100 
m. Fr. Jan. 

66. Rhododendron solitarlum Sleum. Blumea 12 
(1963) 94. 

Erect, terrestrial shrub, sprawling, one- or few- 
stemmed, 1-1.5 m, branches few and short; 
branchlets and young shoots covered with a rusty 
tomentum of stellate scales, which is :- detersile 
and disappears rather early; internodes 4—7 cm. 
Leaves in 4-6-merous pseudowhorls at the upper 
3(-4) nodes, oblong-elliptic or — broadly elliptic, 
sometimes subobovate-elliptic, apex — rounded, 
not rarely slightly retuse. base broad-attenuate to 
rounded, dark green and glossy above, yellowish 
green below, mature ones glabrescent above, re- 
maining lepidote for a certain time below especially 
at midrib and nerves (scales ir dendroid, marginal 
zone stellate-incised to various degree, the small 
centre deepened and protracted downwards into 
a short foot, the latter on top of an epidermal 
tubercle), 5-8 by 3-5 cm, bullate, midrib as wide 
as the petiole and hardly prominent in the lower, 
gradually narrower and -~. impressed in the upper 
part above, on the undersurface thick as the 
petiole at the base, gradually narrower and less 
prominent distally, nerves in 7-9 pairs, irregular 
and not rarely deeply forked, : transverse and 
straight from the midrib, becoming curved up- 
wards, anastomosing quite a distance before the 
edge dissolving into a coarse network of veins, all 
nerves and veins deeply impressed above, 4: 
sharply prominent beneath, veinlets much less 
conspicuous; petiole densely lepidote initially, 
(0.8-)l-1.5cm by 1.5-2.5 mm. Umbels A-(>-^ov/cv- 
ed. Bracteolcs linear to subspathulate, caducously 
lepidotc-fimbriate, 1-1.5 cm by 1-2 mm. Pedicels 
densely rufous-lepidote, c. 1.5 cm by 1 mm at 
anthesis. Calyx obliquely patellar, c. 3 mm 0, 
lobes sometimes elongate. Corolla tubular below, 
widely funnelform-expandcd at the lobes, white, 
densely stellate-lepidotc outside, the marginal 
part of the lobes excepted, densely set with 
rctrorsc hairs at the tube inside, 5-5.5 cm long in 
all. tube slightly curved, c. 4 cm long, c. 4 mm 
at base, r. 7 mm at apex, lobes obovate-spathulate, 
c. 1.5 by 1-1.5 cm. Stamens 10, c. 5 mm cxserted 
from the throat; filaments linear, laxly hairy in the 
lower y^, glabrous above; anthers oblong, base 
obtuse, 3-3.5 mm. Disk glabrous below, tomcntosc 
upwards. Ovary subcylindric, r. 10 by 2.5 mm, 
densely set with yellowish hairs and scales; style 
continuous with the ovary, densely to more laxly 



hairy and lepidote in the lower 3^ part only, 
slightly exceeding the corolla tube. 

Distr. Malesia: E. New Guinea (Morobe 
Distr.: Edie Creek and Merri Creek; common on 
Mt Kaindi). 

Ecol. Growing solitary in eroded gold workings, 
on rather open, Vaccinium- and grass-covered 
slopes, at c. 1850 m. Fl. June. 

67. Rhododendron schoddei Sleum. Blumea 12 
(1963) 95. 

Terrestrial shrub, c. 1 m. Branchlets rather 
slender, divaricate, densely set with sessile rusty 
scales at the upper internode only; internodes 
2-4.5 cm. Leaves in 3-4-merous pseudowhorls at 
the upper 1-2 nodes, ovate- or subovate-oblong, 
apex gradually or shortly attenuate, tip bluntish, 
base ± rounded, young ones subdensely lepidote 
on both faces, mature ones coriaceous, glabrescent 
and shining aijove, paler and more longer remain- 
ing lepidote below (scales rusty, subdendroid, 
marginal zone stellate-dented or -lacerate to vari- 
ous degree, centre small, often a little deepened 
and prolonged downwards into a short foot, on 
top of a low epidermal tubercle), subrevolute, 
(3_)3.5_5(_6) by 1.5-2.3 cm, midrib narrowly 
impressed above, stout in the lower part, gradually 
decreasing and finally ± evanescent distally below, 
nerves 6-8 pairs ± right from the midrib, curved- 
anastomosing towards the edge, slightly impressed 
above, hardly prominent or obscure beneath, no 
reticulation; petiole reddish, subdensely lepidote, 
1.5-2 cm by c. 1 mm. Umbels (2-)3-flowered. 
Ferulae ovate-acuminate, appressedly lepidote 
dorsally. Calyx obliquely patellar, lepidote dor- 
sally, shortly 5-lobed, c. 3 mm 0. Coro//« deep pink, 
broadly tubular below, expanded at the lobes, 
subdensely to densely lepidote at the tube outside, 
laxly hairy there inside, c. 2.5 cm long in all, 
tube straight, 1 .8-2 cm, 4-5 mm at base, 6-7 mm 
at apex, lobes obovate-spathulate, shortly retuse, 
6-8 mm 0. Stamens 10, subequalling the co- 
rolla; filaments linear and laxly hairy below, more 
filiform and glabrous distally; anthers oblong, 
each theca with a short basal apiculus, c. 2.5 mm. 
Disk prominent and glabrous except the upper 
hairy margin. Ovary subconical-cylindric, c. 5 by 
2.5 mm, slightly 5-angular, densely lepidote and 
short-hairy, more densely and longer hairy at the 
angles, continuous with the style, the latter rather 
slender, hairy in the '/,-'/2 lower part, glabrous 
for the rest; stigma rather ihickish. 

Distr. Malesia: New Guinea (Western High- 
lands: Lagaip Valley). 

Ecol. On limestone ridge in heath-like vegeta- 
tion at c. 2600 m. //. Aug. 

68. Rhododendron phaeochristum Sieum. Blumea 
12 (1963) 95; Jahrb. Rhodod. Ges. Bremen (1963) 
27, in text. 

Terrestrial shrub, (l-)2-3(-4) m; branches 
erect, deeply divided; branchlets slender, tips 
covered with a rusty tomentum of stellate scales; 
internodes (2-)3-6(-10. rarely up to 18) cm. 
Leaves in (3-)4-5(-6)-mcrous pseudowhorls at the 



524 



Flora Malesiana 



[ser. I, vol. 6* 



upper 2-3 nodes, elliptic- to obovate-oblong, apex 
obtusely attenuate or subacuminate, base ± cune- 
ate into the petiole, subcoriaceous, dark green and 
shining above, yellowish green below, young ones 
clad with a rather detersile rusty tomentum of 
star-shaped scales initially, early glabrescent with 
age above, less so below especially at midrib and 
nerves (scales dendroid, marginal zone deeply 
stellate-incised or -lacerate, cupular-depressed to- 
wards the minute centre, the latter prolonged 
downwards into a very slender stem, which rests 
on a distinct epidermal tubercle), slightly revolule 
at the very edge, (3-)4-7(-9) by 2-3 (^) cm, mid- 
rib narrowly impressed above, thick-prominent in 
the lower, decreasingly so in the upper part below, 
nerves 6-8(-9) pairs, rather irregular, curved- 
anastomosing before the edge, very slightly im- 
pressed or not rarely obscure above, hardly 
prominent beneath; petiole densely lepidote initi- 
ally, 8-1 2 (-15) by 1.5-2 mm. Umbels 2-3(-*)- 
flowered. Floral buds subglobose, 1-1.5 cm 0, 
perulae thin, outer ones ovate, densely stellate- 
lepidote at the dorsal midline, caducously lepidote- 
fimbriate, c. 1 cm, inner ones subspathulate, i 
glabrous dorsally, up to 1.2 cm. Bracteoles sub- 
spathulate-linear, glabrous, c. 1 cm by 1-1.5 mm. 
Pedicels rather slender, densely stellate-lepidote, 
c. 1 cm at anthesis. Calyx obliquely patellar, 
minutely 5-Iobed, lepidote dorsally, c. 2.5 mm 
across. Corolla curved-tubular, expanded at the 
lobes, zygomorphous, pale to more deeply carmine, 
2.5-3 cm long in all in dry, 3.5—4 cm in fresh 
specimens, thin-fleshy, glossy and densely set with 
golden scales outside, laxly hairy at the tube inside, 
tube 1.5-2 cm long, c. 3 mm at base, 5 mm at 
throat, lobes subspathulate-obovate, c. 1 cm 0. 
Stamens 10, attaining i half the corolla lobes; 
filaments linear, densely to more laxly hairy in the 
lower 3^ part, glabrous above; anthers obovate- 
oblong, dark purple to almost blackish when fresh, 
base obtuse, 2 by 1 mm. Disk prominent, hairy 
only at its upper margin. Ovary subcylindric, 5 
by 2.5 mm, densely set with rusty stellate scales and 
paler, ± appressed hairs, tapering to the style, 
the latter c. 1.2 cm, base slightly hairy, glabrous 
for the rest; stigma subclavate. Fruit fusiform, 
densely lepidote and hairy, 2(-2.5) cm by 4-5 
mm. 

Distr. Malesia: W. New Guinea (Vogelkop 
Peninsula: Tamrau, Nettoti and Arfak Mts). 

Ecol. In understorey of Nothofagus forest, 
1800-2250 m, often sterile. Fl. Jan. 

Vern. Tschotschai, Manikiong, arun'o, Hattam. 

Note. A natural hybrid of R. phaeochristum 
with apparently 268. R. ciilminicolum var. angiense 
was found once in the Arfak Mts near Lake Anggi 
Gita, 1900 m; in the general appearance, in the 
star-shaped scales and in flowers it shows the 
characters of the first, and in the wider, more 
obovate and blunt leaves (which show a less dense 
tomentum of stellate scales and no trace of 
epidermal tubercles) the characters of the latter. 
On Mt Gwamongga near Lake Anggi Gigi at 
2550 m a sterile plant has been collected, very 
similar in leaf-characters to R. phaeochristum, but 



tubercles much less distinct than normally found 
in this species, probably also due to an ingression 
of a species of ser. Javanica. 

69. Rhododendron haematophthalmum Sleum. 

Reinwardtia5 (1960) 89. 

Erect rigid shrub, 1.5-2.5 m high. Branchlets 
terete, upper internodes 2-8 cm by 2-3 mm, lower 
ones 3-7 mm 0, stellate-lepidote at the innova- 
tions, otherwise very early glabrescent, and very 
rugose by innumerable minute tubercles. Leaves 
mostly scattered, close together at the upper node, 
more rarely along the internode, or in a 3-5- 
merous pseudowhorl, ~ oblong, apex short- 
acuminate, apiculate or obtuse, base truncate- 
rounded, not rarely slightly cordate, very coriace- 
ous, rigid, erecto-patent, densely clad with minute, 
stellate, short-dendroid scales (each on top of a 
paler epidermal tubercle) when very young, early 
glabrescent and very densely scabrid by the 
persistent tubercles on both faces, margin ± 
revolute, 5-9 by 2-3.5 cm, midrib as thick as the 
petiole below, :^ flattened and but slightly promi- 
nent in its lower, — sunk in its upper part above, 
nerves 7-10 pairs, irregular, spreading, rather 
straight, indistinctly anastomosing, — inconspicu- 
ous above, slightly raised or ir evanescent beneath; 
petiole subterete, rugose, 7-17 by 2-3 mm. Umbels 
4— 8(-10)-flowered. Perulae ovate to obovate or 
spathulate, obtuse, membranous, glabrous, up to 
2 by 1 cm. Bracteoles linear, apex subspathulate, 
glabrous, up to c. 2 cm by 1 mm. Pedicels very 
densely rusty-stellate-lepidote, epilose, (1.5-)2-3.5 
cm by 1 mm at anthesis. Calyx oblique, disk- 
shaped, irregularly 5-dentate, densely lepidote 
dorsally, epilose, c. 4 mm 0, lobes short, obtuse, 
not ciliate, or may be 1-2 lobes elongate-deltoid, 
acute, up to 3 mm. Corolla manifestly zygomor- 
phous, tubular below, dilated at the limb, red, 
densely rufous-stellate-lepidote outside, the margin 
of the lobes excepted, very shortly hairy at the 
tube inside, (4-)4.5-5.5 cm long in all, tube ± 
curved, 2.5-3.5 cm, 5-6 mm at base, widened to 
c. 10 mm at apex, lobes irregularly broad-obovate 
or suborbiculate, 1.5-2 by 0.9-1.5 cm. Stamens 
10, unequal, nearly as long as the corolla; filaments 
linear, laxly ± patently hairy below, glabrous 
distally; anthers forming a purple centre in the 
throat of the corolla, broad-oblong to obovate- 
oblong, 3.5-4 by 1.3-1.5 mm. Disk glabrous. 
Ovary cylindric, subabruptly attenuate to the style, 
densely shortly patent-hairy and lepidote (the 
scales well visible among the hairs), 5-6 by c. 3 mm. 
Style rather stout, ± equalling the stamens, 
subdensely to laxly patently hispid-pubescent in 
the lower half, with some scales between the hairs, 
otherwise glabrous; stigma thick-capitate, mani- 
festly 5-gibbous. 

Distr. Malesia: W. New Guinea (Lake Habbe- 
ma). 

Ecol. Common on edges of mossy thickets, 
frequent in shrubberies of peaty ridges or peaty 
treeless area in forest or in mossy thickets, 3000- 
3225 m. Fl. Aug., Oct. 



Jan. 1966] 



Ericaceae (Sleumer) 



525 



70. Rhododendron phaeops Sleum. Reinwardtia 5 
(1960) 90. 

Epiphytic shrub, 1 m. Branchlets terete, 2-3 mm 
o, tips stellate-lepidote, otherwise densely minutely 
verrucose and rough to the touch, early covered 
with cork; internodes 3-6 cm. Leaves in 3-4- 
merous pseudowhorls at the upper 2 or 3 nodes, 
broad-eUiptic or ovate-elliptic, apex shortly attenu- 
ate, obtuse, base very broadly attenuate to nearly 
rounded, coriaceous, rigid, stellate-lepidote initi- 
ally on both faces (scales small, dendroid, on top 
of persistent epidermal tubercles, caducous), early 
glabrescent and scabrid by numerous tubercles on 
both faces especially beneath, edge but shortly 
revolute, 5-8 by 3-5 cm, midrib as thick as the 
petiole, somewhat prominent and grooved along 
the midline in its lower -/}, impressed for the 
rest above, nerves 6-8 pairs, irregular, rather 
straight below, obscurely curved-anastomosing 
before the edge, very slightly immersed above, 
prominent beneath, veins lax, faintly sunk above, 
raised or — obsolete beneath; petiole grooved 
above, densely lepidote, 1-1.5 cm by c. 2 mm. 
Umbels 2-3-flowered. Pedicels densely rusty- 
stellate-lepidote, 1.5-2 cm by c. 1 mm. Ca/j'x disk- 
shaped, densely lepidote, epilose, 5-6 mm 0, with 
5 distinct irregular elongate-triangular subacute 
teeth, these white-ciliate, spreading, 2-4 mm. 
Corolla zygomorphous, tubular below, gradually 
widened upwards, red, 5.5-6 cm long in all, densely 
rusty-stellate-lepidote outside the margin of the 
lobes excepted, subdensely clothed with slender, 
whitish, rather longish retrorse hairs at the entire 
tube inside, tube curved, 6 mm at base, c. 15 mm 
at throat, lobes broad-spathulate to suborbicular, 
1.5-2 cm 0, glabrous inside. Stamens 10, unequal; 
filaments linear, wider and ± densely patent- 
pilose below, narrower and glabrescent to glabrous 
above, 4.5-5.5 cm; anthers brown when fresh, 
broad-oblong, 5-5.5 by nearly 2 mm. Disk 
glabrous. Ovary obconical-cylindric, densely clad 
with subappressed, yellowish, stiff hairs which 
cover the scales, c. 7 by 3 mm, — abrupt with the 
style, the latter rather stout, densely hairy and 
laxly lepidote below, more laxly patent-hairy 
and more densely lepidote upwards, glabrous at 
the upper 5 mm below the capitate stigma. 

Distr. Malesia: W. New Guinea (Bernhard 
Camp, Idcnburg R., c. 139-3 30'), once found, 
in mossy forest at 2150 m. Fl. Feb. 

71. Rhododendron truncicolum Sleum. Reinwardtia 

5(1960)91. 

Small shrub, epiphytic on tree trunks. Branchlets 
slender, 1.5-2.5 mm 0, caducously lepidote at the 
tips, otherwise minutely verrucose, covered with 
grey cork in the older parts; internodes 2-4 cm. 
Leaves in 3-4-mcrous pseudowhorls at the upper 
nodes, elliptic, apex hroad-attenuate, subapiculatc, 
oasc broad-attcnuatc to rounded, subcoriac:ous, 
mature ones glabrescent or with remains of 
former scales, otherwise with minute elevated 
points above, lepidote for a long time beneath 
alonK the midrib fscales stellate, subscssile on top 
of p.ilc persistent epidermal tubercles), otherwise 



glabrescent and somewhat rough to the touch, 
quite flat, (2.5-)3^ by 1.4-2.4 cm, midrib flat 
or slightly sunk above, prominent beneath especi- 
ally towards the base of the blade, much more 
slender than the petiole, nerves c. 6 pairs, irregular, 
rather straight below, anastomosing with each 
other and the veins before the edge, the latter 
forming a lax network which is equally raised on 
both faces, veinlets rather inconspicuous; petiole 
a little flattened, grooved above, densely lepidote, 
4-6 by 1-1.3 mm. Umbels 3-flowered. Ferulae 
membranous, ovate to obovate, apiculate, papillose 
or glabrous dorsally, up to 1 .5 by0.6cm. Bracteoles 
filiform, very laxly hairy or glabrous, up to 1.5 cm. 
Pedicels densely rusty-stellate-lepidote, epilose, 
1-1.3 cm by c. 0.6 mm. Calyx disk-shaped, 
hardly lobed, c. 2.5 mm 0, densely lepidote and 
sparsely pubescent dorsally, sparsely ciliate. 
Corolla zygomorphous, tubular, rosy-pink, (3-) 
3.5—4 cm long in all, laxly stellate-lepidote out- 
side at the tube and the base of the lobes, laxly 
hairy inside at the tube, glabrous elsewhere, tube 
slightly curved, 2-2.5 cm long, 4-5 mm at base, 
8-10 mm below the lobes, these broadly obovate- 
spathulate, 1-1.5 by 0.8-1 cm, irregularly wavy 
and crenulate. Stamens 10, unequal, attaining in 
half the length of the corolla lobes; filaments 
linear, laxly patent-hairy below, glabrous in the 
upper third; anthers broad-oblong, 2.5 by 1 mm, 
thecae very shortly apiculate at the base. Disk 
glabrous below, pubescent at the upper margin. 
Ovary cylindric, densely lepidote and laxly 
pubescent, c. 4 by 1.8 mm, rather abrupt with the 
style, the latter slender, slightly exceeding the 
tube, laxly patently hispidulous and densely lepi- 
dote in the lower third, glabrous for the rest; 
stigma thickly turbinate-capitate. 

Distr. Malesia: New Guinea (Mt Dayman/ 
Mt Maneao, Milne Bay Distr.), once found in 
forest at c. 2135 m. Fl. June. 

72. Rhododendron rappardii Sleum. Reinwardtia 
5 (1960) 93; Blumea 12 (1963) 99. 

Terrestrial shrub, 1.2-2.5 m. Branchlets terete, 
upper internodes 4—10 cm by 2 mm, subdensely 
caducously rusty-stellate-lepidote initially, glabres- 
cent, covered with numerous minute pustules 
and rather scabrid to the touch, pruinosc at the 
older parts (3-5 mm 0), which are smooth re- 
spectively early covered by an ashy cork. Leaves in 
3-5-merous pseudowhorls at the upper 3 or 4 
nodes, broad-elliptic, more rarely oblong- 
elliptic or nearly rounded, apex broad-attenuate, 
obtuse or rounded, base very broadly attenuate or 
mostly rounded, rather thinly coriaceous, though 
firm, fugaciously rusty-or brown-lcpidotc on both 
faces initially (scales rather small, irregularly 
stellate-lacerate, i dendroid, each on a minute 
epidermal tubercle), very early glabrescent and ± 
smooth above, finally glabratc also beneath, but 
permanently rugulose by numerous minute 
tubercles, entire, the edge slightly or not revolute, 
(3.5-)4-7(-7.5) by f2.5-)3-5 cm, midrib dilated 
and grooved in the basal, becoming very narrow 
and : impressed in the upper part above, thick 



526 



Flora Malesiana 



[ser. I, vol. 6* 



and obtusely prominent beneath, nerves 5-6(-8) 
pairs, at an acute angle, rather straight below, 
less conspicuous towards the edge, minutely 
impressed in fully mature leaves above, not or but 
little raised beneath, veins ± indistinct; petiole 
subterete, lepidote, 6-12(-15) by c. 2 mm. Umbels 
5-8-flowered. Outer perulae ovate, obtuse, lepidote 
in the middle outside, otherwise papillate-puberu- 
lous, 1-1.5 by 0.7-1 cm, inner ones oblong to 
spathulate, densely and longer subsericeous- 
pubescent on both sides, up to 3 by 1.5 cm. 
Bracteoles linear, i laxly hairy, up to 2.5 cm by 
c. 1 mm. Pedicels stoutish, densely rusty-stellate- 
lepidote, subdensely or laxly hairy at least below 
the calyx, (5-)8-12 by nearly 1 mm at anthesis. 
Calyx oblique, disk-shaped, rather small, lepidote 
and pubescent dorsally, ciliate, very shortly 5- 
lobed, 2-3 mm 0. Corolla narrow funnel-shaped, 
regular, white or pink, very sparsely stellate- 
lepidote or mostly glabrous outside, di densely 
clad with ± longish retrorse hairs at the tube 
inside, 4-5 (rarely up to 5.5) cm longin all, tube rh 
straight, gradually widened towards the mouth, 
3-4 mm at base, 6-8 mm at the throat, lobes 5, 
irregular, obovate spathulate, 1.2-1. 5 (-1.8) by 
(0.6-)0.8-1.2 cm. Stamens 10, i attaining halfway 
the corolla lobes; filaments linear and almost 
villous below, rather abruptly glabrescent up- 
wards, glabrous and filiform distally; anthers 
elongate-oblong, 3.5-4 by 1 mm, base minutely 
biappendiculate. Disk low, i pubescent. Ovary 
cylindric, gradually attenuate to the style, 6-8 by 
2.5-3 mm, densely clad with appressed, yellow, 
stiff hairs which cover the scales. Style nearly as 
long as the stamens, rather thick below, -}z densely 
patent-hairy and ± laxly lepidote to the lower Yi 
(less densely hairy distally), more slender and 
glabrous for the rest; stigma capitate. 

Distr. Malesia: W. New Guinea (Wissel Lake 
region). 

Ecol. In primary or mostly in secondary forest, 
on ± steep slopes, 1 750-1 850(-2300) m, on 
peaty or clayey soil or stony clay, derived from 
limestone or sandstone, locally common. Fl. 
Jan., March, May, Oct. 

Vern. Wajade, warade, waradei, Kapauku. 

73. Rhododendron dianthosmum Sleum. Blumea 12 
(1963) 100; Jahrb. Rhodod. Ges. Bremen (1963) 
10, in text. 

Epiphytic, much branched shrub, 0.5-2 m; 
branchlets rather slender, densely lepidote, early 
glabrescent and pruinose; internodes (4-)6-16 cm. 
Leaves in 3(-4)-merous pseudowhorls at the 
upper 2 or 3 nodes, oblong or elliptic-oblong, 
apex shortly subacuminate-attenuate, bluntish, 
base broadly attenuate to rounded, coriaceous, 
young ones densely rufous-stellate-lepidote, mature 
ones glabrescent, on both faces set with numerous 
epidermal tubercles and rough to the touch (scales 
dendroid, marginal zone incised to or almost to 
the small centre, the latter deepened or prolonged 
into a short foot), (7-)9-14 by 3-5.5 cm, edge a 
little recurved, midrib not much prominent in the 
lower, gradually flattish in the upper part above. 



as thick as the petiole in the basal, gradually 
decreasing in the upper part below, nerves 8-10 
(-12) pairs, irregular, j^ inarched before the edge, 
slightly prominent on both faces, veins laxly 
reticulate, not much conspicuous; petiole densely 
lepidote initially, 1-1.5 cm by 2-2.5 mm. Umbels 
4-6-flowered. Perulae a sordid red, outer ones 
ovate, more inner ones elliptic-ovate and apiculate, 
innermost ones broadly spathulate, lepidote and/or 
appressed-hairy to various degree, 1-3.5 by 1-1.5 
cm. Bracteoles linear, glabrous, c. 2 cm by 1-2 
mm. Pedicels exclusively and very densely rusty- 
stellate-lepidote, 1-1.5 cm by c. 1.5 mm. Calyx 
obliquely patellar, c. 3 mm across. Corolla tubular 
below, ih horizontally spreading at the lobes, 
6-7-merous, pure white, with a scent of carnation, 
glabrous outside, short-hairy at the tube inside, 
5-6 (when fresh c. 1) cm long in all, tube 2.5-3.5 
(when fresh 3-4) cm, 6-7-pouched at the base and 
c. 6 (when fresh c. 10) mm 0, lobes spathulate, 
3 or 4 of them generally smaller than the others, 
2-2.5 (when fresh 3-3.5) by 1-2 cm. Stamens 
exserted from the throat for c. 1 cm; filaments 
linear, densely or more laxly hairy in the lower %, 
glabrous distally; anthers elongate-oblong, base 
obtuse 3.5-4 by almost 1 mm. Disk glabrous 
below, tomentose at the upper margin. Ovary 
subcylindric, 8-10 by 2-3 mm, covered by a 
tomentum of whitish to yellowish, forward- 
directed hairs, which cover the scales; style 
continuous, rather slender, c. 3 cm, densely or 
more distally laxly hairy in the lower 3^, glabrous 
for the rest, no scales visible; stigma capitate, 
2.5-3 mm across. 

Distr. Malesia: N. New Guinea (Cycloop Mts). 

Ecol. In primary forest, 800-1400 m, descend- 
ing along ravines to c. 500 m. Fl. May-July. 

74. Rhododendron rubellum Sleum. Reinwardtia 5 
(1960) 94. — R. phaeochitum {non F. v. M.) 
Wright, Kew Bull. (1899) 104. 

Erect branching tree, up to 5 m. Branchlets 
terete, 2-3 mm 0, densely stellate lepidote at the 
upper 1 or 2 internodes, glabrescent and early 
covered with ashy cork below; internodes 3-8 cm. 
Leaves in 4-5-merous pseudowhorls at the upper 
1 or 2 nodes, elliptic-oblong or oblong, apex short- 
acuminate, apiculate, base broadly cuneate or 
rarely rounded, coriaceous, immature ones densely 
rusty- or brown-lepidote (scales rather large, 
very narrowly stellate-incised in form of stellate 
hairs, dendroid, each on top of an epidermal 
tubercle), early glabrescent and rough by the 
persistent tubercles above, clothed for a long time 
beneath with a nearly woolly tomentum formed 
by the scales which when gone leave the under- 
surface too tubercled and scabrid, dark green and 
dull above, paler beneath, slightly or not revolute 
at the margin, 4-8. 5 (-9. 5) by 2. 5-3. 5 (-4. 5) cm, 
midrib flat or minutely impressed above, as stout 
as the petiole and very prominent in the lower, 
gradually less prominent distally beneath, nerves 
6-8 pairs, starting at an acute angle, rather 
straight below, curved and anastomosing before 
the edge, forming a lax network with the veins. 



Jan. 1966] 



Ericaceae (Sleumer) 



527 



nerves slightly impressed above, raised beneath, 
veins hardly sunk above, raised beneath, no proper 
reticulation; petiole grooved above, densely 
lepidote, 1-1.7 cm by 1.5-2 mm. Umbels 6-8- 
flowered. Outer perulae ovate, subobtuse, inner 
ones spathulate, both densely subsericeous- 
hirtellous on both sides, up to 1.8 by 1 cm. 
Bracteoles linear to very narrowly spathulate, 
— densely hirsutulous, up to 1.3 cm by 3 mm. 
Pedicels densely rusty-stellate-lepidote, otherwise 
not hairy or only so immediately below the calyx, 
0.8-1.5 by 1 mm. Calyx — deeply 5-dentate, 
rather densely pubescent and lepidote dorsally, 
lobes irregularly triangular, subacute, long-fimbri- 
ate, 1.5-2 mm. Coro//fl tubular, — zygomorphous, 
bright red, (4.5-)5-6 cm at full anthesis, very 
laxly lepidote or glabrous at the tube outside, 
laxly set with retrorse hairs at the tube inside, 
tube rather straight below, slightly curved and 
gradually dilated upwards, c. 4 cm long, c. 4 mm o 
at base, c. 8 mm at throat, lobes erecto-patent, 
irregular, obovate-spathulate, sometimes nearly 
orbicular, 1.2-1.6(-2) by 1-1.5(-1.7) cm. Stamens 



10, slightly shorter than the corolla, unequal; 
filaments linear and — densely patent-hairy below, 
gradually less so above, completely glabrous at the 
ultimate 6-10 mm; anthers elongate-oblong, base 
obtuse, c. 4.5 by 1 mm. Disk glabrous at least 
in the lower part. Ovary elongate-cylindric, gradu- 
ally tapering to the style, very densely hairy in the 
lowest 1 mm (which may be the upp>er margin 
of the disk), densely rufous-Iepidote and hairy on 
the proper ovary (scales more numerous than the 
hairs, and thus well visible), 8-10 by 2.5-3 mm. 
Style as long as or a little longer than the tube, 
laxly pubescent and very sparsely lepidote in the 
lower 1/3 to 2/3, glabrous for the rest; stigma 
thickly turbinate-subcapitate. Capsule 3-3.5 by 
c. 0.6 cm, densely rufous-hirsute and lepidote. 
Seeds filiform, 3-4 mm, with a very narrow tail 
on both ends. 

Distr. Malesia: E. New Guinea (Wharton and 
Owen Stanley Ranges). 

Ecol. In forests, said to be common at the 
Murray Pass (Wharton Range) at 2840 m. Fl. 
June-Sept. 



4. Subsection Malayovireya 

Sleum. Blumea Suppl. 4 (1958) 48; Reinwardtia 5 (1960) 95. — Rhododendron ser. Malayanum 
COPEL./. Am. Midi. Nat. 30 (1943) 605, in text, nom. nud. — Fig. 17—18. 

Scales sessile, flat, slightly impressed, generally of two different sizes (i.e. very many smaller, 
and fewer much larger ones, irregularly mixed), all very dense, touching or partly overlapping 
each other, the centre very dark (sooty to blackish brown) and large in relation to the paler 
marginal zone, the latter irregularly several-lobed or -incised up to the centre (the broad lobes 
again, but more shortly lobed) and becoming silvery in later stages. 

Distr. S. Siam, Malesia (excluding Lesser Sunda Is. and New Guinea). Fig. 16. 

Note. Scales of the same type, but less distinctly inequal and spaced for their greater, partly overlapping 
each other for their smaller number, point to hybrids with species of other subsections, viz Pseudovireya, 
Solenovireya, and Euvireya. The influence of the Malayovireya parent is easily recognized by the dark 
chestnut colour of the scales, chiefly of their centre, whilst the influence of the other parent generally 
results in reducing the size of and increasing the distance between the scales. Species with such :;; inter- 
mediate type of scales are kept in suhsect. Malayovireya. Species as 181. R. pyrrhophorum and 179. 
R. ripleyi var. cryptogonium show the Malayovireya-inf\uence to a minor degree and are placed in subsect. 
Euvireya. .See also the note under subsect. Albovireya. 



^ 







FiK. K). Distribution ot Rhododendron subsect. Malayovireya (- 
and subsect. Siphonovlreya (- 



-), subsect. Phaeovireya ( ) 

■-). 



528 Flora Malesiana [ser. I, vol. 6* 

KEY TO THE SPECIES 

1. Scales very dense, all or most of them touching, respectively partly covering each other, very different 

in size, i.e. many smaller ones mixed with few other ones of almost double their size. 
2. Leaves linear to lanceolate-linear, up to 10 cm by 4 mm. 
3. Inflorescences 4-6-flowered. Corolla i campanulate, L2-1.4 cm; lobes 8-10 by 7-9 mm. 

75. R. himantodes 

3. Inflorescences l-3(very rarely -4)-flowered. Corolla ± tubular, 2-2.4 cm; lobes 5-6(-7) by 
3-4(-5)mm 76. R. vinicolor 

2. Leaves of various shape, at least 6 mm wide. 

4. Leaves sessile or nearly so (petiole up to 4 by 3 mm), base rounded to subcordate-auriculate. (Fila- 
ments pubescent at the base). 

5. Corolla glabrous or rarely very sparsely lepidote in the lower part of the tube outside, 2.4-2.7 
(-3.2) cm. Pedicels shorter than the corolla 77. R. durionifolium 

5. Corolla ± densely lepidote outside at least at the tube, c. 2 cm. Pedicels as long as or mostly 
longer than the corolla 78. R. fallacinum 

4. Leaves manifestly petioled, base mostly cuneate to rounded, very rarely subcordate. 

6. Inflorescences exclusively terminal. 
7. Filaments glabrous. 

8. Style thick, rather short, as long as or shorter than the proper ovary, hardly half as long as the 
corolla 79. R. apoanum 

8. Style slender, ± twice as long as the proper ovary, — as long as the corolla. 

80. R. malayanum var. malayanum 
7. Filaments ± densely short-pubescent in the lower 14 or Vs- 

9. Corolla besides some scales laxly short-pubescent outside . . 80. R. malayanum var. pubens 
9. Corolla besides some scales not hairy outside. 

10. Corolla 4—4.5 cm 81. R. nortoniae 

10. Corolla up to 3.2 cm. 
1 1 . Leaves — distinctly buUate. Pedicels lepidote, not hairy. 
12. Leaves ovate or elliptic-ovate, sometimes ovate-lanceolate, base mostly rounded, or rarely 
attenuate. Corolla short-pubescent inside in the lower half of the tube . . 82. R. acuminatum 
12. Leaves elliptic-lanceolate, base acute or obtuse. Corolla similar but glabrous at the inser- 
tions of the filaments 83. R. fortunans 

1 1 . Leaves not bullate. Pedicels finely patent-hairy under and between the scales. 

80. R. malayanum var. pilosifilum 

6. Inflorescences both terminal and axillary 80. R. malayanum 

1. Scales all or for their greater number spaced from each other, much less different in size. Presumably 
hybrids with species of other subsections. 
13. Leaves linear-lanceolate, (2.8-)3-4(-5) cm by (3-)4-5 (rarely up to 8) mm . . 84. R. lineare 
13. Leafshape otherwise, (4.5-)6-12(-16) by (2-)2.5-4(-5.5) cm. 
14. Filaments glabrous. 
15. Leaves oblong-obovate, apex rounded or slightly retuse 85. R. obscurum 

15. Leaves lanceolate to elliptic-lanceolate, rarely elliptic, gradually attenuate towards the sub- 
acute apex 86. R. variolosum var. variolosum 

14. Filaments rather densely patent-pubescent at least in the lower third. 

16. Ovary and base of the style exclusively lepidote; style glabrous for the rest. Corolla very laxly 
lepidote or glabrous outside 86. R. variolosum var. andersonii 

16. Ovary densely lepidote and whether or not laxly patent-pubescent; style patent-pubescent to 
various length. 
17. Ovary lepidote and laxly pubescent; style manifestly patent-pubescent to its lowest 2 mm, 
glabrous for the rest. Corolla practically glabrous outside. Leaves acutely attenuate at both 

ends (225. R. javanicum var. teysmannii X R. malayanum) 87. R. wilhelminae 

17. Ovary lepidote, epilose or nearly so; style subdensely lepidote and laxly patent-pubescent in 
the lower 2/3, glabrous for the rest. Corolla laxly lepidote and patent-pubescent outside. Leaves 
shorter or longer acuminate at the apex, rounded to subcordate at the base (136. R. jasmini- 
florum var. punctatum /. R. malayanum) 88. R. hybridogenum 

75. Rhododendron himantodes Sleum. Bot. Jahrb. leaves and flowers; internodes l-4(-5.5) cm. 

71 (1940) 145; Reinwardtia 5 (1960) 97. Leaves in 4-5-merous pseudowhorls at the end 

of the upper 2 or 3 (-4) internodes, linear, elongate, 

var. himantodes. apex subacute, base obtuse, rz coriaceous, green 

Large epiphytic shrub, with lax spreading above, paler grey-green and with a brownish 

branches; branchlets slender, at the upper inter- midrib beneath, initially entirely covered with 

nodes very densely covered with scales of the scales which disappear above with age, but remain 

R. malayanum type, which are also found on beneath, margin strongly revolute to nearly the 



Jan. 1966] 



Ericaceae (Sleumer) 



529 



top, 5-9(-10) cm by 2-4 mm, midrib deeply im- 
pressed, very prominent beneath, nerves obscure; 
petiole densely lepidote, stoutish, 2-3 mm. 
Flowers 4-6 per umbel. Ferulae ovate to obovate- 
spathulate, the upper dorsal part and/or the 
mediane lepidote, ciliolate, up to 10 by 6 mm. 
Bracteoles narrow-linear, glabrous or laxly lepi- 
dote, up to 8 mm. Pedicels slender, densely 
lepidote, 1.5-2 cm at anthesis. Calyx minute, 
oblique, obsoletely 5-lobulate or -denticulate, 
densely lepidote. Corolla campanulate to infundi- 
buliform-campanulate, deeply 5-lobed, 1.2-1.4 cm 
in all, snow-white with brown dots, membranous, 
tube broadly tubular-campanulate, 3-6 mm long, 
at the base with 5 small pouches c. 2.5 mm 0, 
rather densely lepidote outside, ^ densely hairy in- 
side, lobes obovate-spathulate, — spreading, 
glabrous outside in the upper part as well as inside, 
lepidote at the base outside, 8-10 by 7-9 mm. 
Siamens 10; filaments linear, 4—5 mm, white- 
hairy especially at the very base; anthers c. 2.5 
by 0.3 mm. Disk glabrous. Ovary conical, entirely 
covered by scales, 4-5 by 2 mm, gradually tapering 
to the glabrous style, the latter slightly curved, 
c. 4 mm; stigma capitate. Capsule narrow-oblong, 
lepidote, c. 1.5 cm. Seeds 7-8 mm including the 
tail-like appendages. 

Distr. Malesia: North Borneo (Mt Kinabalu). 

Ecol. In mossy forest, crest of ridge, 1525-2135 
m. Fl. Feb.-Oct. 

var. lavandulifolium Sleum. Reinwardtia 5 (1960) 
97. 

Leaves 3-4.5 cm by c. 2 mm. Flowers only seen 
in bud, very light yellow. 

Distr. Malesia: E. Borneo (Mt Kemul), epi- 
phyte in mountain forest, 1800 m. Fl. Oct. 

76. Rhododendron vinicolor Sleum. Reinwardtia 

5 (I960) 98. 

Epiphytic shrub. Branchlets elongate, terete, 
slender, completely covered with scales of the 
R. malayanum type as they occur also on the 
inflorescences and the leaves. Leaves rather laxly 
scattered along the branchlets, more densely so 
distally, linear or narrow-lanceolate, apex gradu- 
ally attenuate, obtuse, base narrowed into the 
petiole, subcoriaceous or coriaceous, still at age 
very densely covered with scales which are pallid 
on the upper, red-brown on the undersurface of 
the leaves, strongly revolute, 3-6(-7) by (2-)3-4 
mm. midrib impressed above over its entire length, 
very prominent beneath, nerves obsolete; petiole 
densely lepidote, 2-3 by 0.5 mm. Umbels 1-3 (very 
rarely -4)-nowered. Pcrulac ovate, subapiculate, 
densely lepidote dorsally, rather long white-ciliate, 
up to 5 by 3 mm. Bracteoles narrow-subulate, 
glabrous, up to 1 cm long, base c. I mm wide. 
Pedicels very slender, densely lepidote, 6-8(-l2) 
mm. Calyx minute, disk-like, shallowly or not 
lobcd. ♦: wavy, densely lepidote. Corolla winc-rcd 
or reddish purple, tubular. 2-2.4 cm in all, tube 
cylindric, f curved, gradually attenuate towards 
the slightly 5-saccatc base. 2-2.5 mm at base, 
c. 4 mm at throat, laxly and .-. caducously lepidote 



outside, glabrous inside, lobes somewhat spread- 
ing, obovate, lepidote on the dorsal midline, 
glabrous elsewhere, 5-6(-7) by 3^(-5) mm. 
Stamens 10, 1.7-1.9 cm; filaments narrow-linear, 
filiform distally, slightly unequal, glabrous; 
anthers broadly obovate, c. 1.5 mm. D/i/c glabrous. 
Ovary obconical-cylindric, densely lepidote, c. 
4 by 1.5 mm, gradually passing into the style, the 
latter slender, lepidote in the lower third or only 
at the base, (8-) 10-1 2 mm; stigma shortly club- 
shaped and obscurely lobed. 

Distr. Malesia: Sumatra (Atjeh: Mt Lembuh 
region; Tapanuli: near Sidikalang). 

Ecol. In flat forest ridges with cold solfatara 
fields and on narrow cricoid ridge, 1100-1500 m. 
Fl. Feb. -March. 

77. Rhododendron durionifolium Becc. Malesia 1 
(1878) 202; Ridl. J. Str. Br. R. As. Soc. n. 63 
(1912) 61; Merr. En. Born. (1921) 461; Sleum. 
Reinwardtia 5 (1960) 98; Blumea 11 (1961) 116; 
ibid. 12 (1963) 100. — /?. mjobergii Merr. Sarawak 
Mus. J. 3 (1928) 542. 

Treelet or shrub, terrestrial or mostly epiphytic. 
Branchlets terete, stout, entirely covered in the 
younger parts with initially chestnut to dark 
golden squamular scales of the R. malayanum type, 
as they are found also in the inflorescences and 
the leaves; internodes 3-16 cm by (2-)3-4 mm. 
Leaves 3-4(-5) in each of the spaced pseudowhorls, 
or only 2-4 not properly whorled in the apical 
part of the upper internode, sessile or nearly so, 
lanceolate to ovate-lanceolate, apex ± long- 
acuminate to short-caudate, base rounded or most- 
ly subcordate-auriculate, coriaceous, showing very 
thin silvery dots or small pits above, entirely and 
persistently covered with red-brown scales beneath, 
margin repand-denticulate to nearly entire, slightly 
revolute, (8-)ll-l5 (-17) by (2-)2.5-4(-6.5) cm, 
midrib distinctly impressed above, very stout and 
prominent beneath, nerves 6-9 pairs, rather 
straight and spreading below, ± curved and 
anastomosing towards the edge, slightly impressed 
above in mature leaves only, r indistinct beneath, 
veins laxly reticulate and mostly minutely im- 
pressed above only; petiole densely lepidote, up 
to 4 by 3 mm, but mostly very short or almost 0. 
Flowers 8-10(-14) per umbel; floral buds ovoid, 
large, 2-2.5 by 1.2-1.5 cm, their outer pcrulae 
ovate, obtuse to subapiculate, base truncate, 
subcoriaceous, subpuberulous outside, lepidote 
along the upper part of the dorsal mediane, 
finely whitish ciliatc, up to 2 by 1 .2 cm, inner ones 
± spathulatc and smaller. Bracteoles filiform, 
papillose-pubescent, often subspalhulate and 
barbulate-ciliatc at the apex, up to 1.5 cm long. 
Pedicels densely lepidote, 1-1.5 cm by c. I mm 
at anthesis. Calyx very small, formed by the upper- 
most thickened cup-shaped part of the pedicel. 
Corolla yellow to orange, or scarlet red, funnel- 
form, 2.4-2.7(-3.2) cm, mostly completely gla- 
brous, rarely with a few (apparently rather early 
caducous) scales on the lower part of the tube 
outside, glabrous or slightly pubescent at 
the base of the tube inside, tube ± straight, 



530 



Flora Malesiana 



[ser. I, vol. 6* 







F\g. n. Rhododendron fallacinum Slevm. a. Habit, X 2/3, 6. flower, x 1, c. ovary and style, x 2, ^.stigma, 

X 5, e. stamen, x 3,/. fruit, x Vs' S- seed, x 10, h-k. subsequent outer to inner perulae, x *js {a and 

f-g Griswold 122, b-e Darnton 585, h-k Clemens 27106, 27800, 32714). 



Jan. 1966] 



Ericaceae (Sleumer) 



531 



1.2-1.5 cm long, c. 3 mm o at the slightly 
5-pouched base, c. 4—5 mm at throat, lobes ± 
spreading, obovate to nearly rounded, 4- as long 
as the corolla tube. 8-11 mm wide. Stamens 10, 
slightly unequal, c. 2 cm; filaments linear, flattened 
and rather densely whitish hairy in the lower part, 
filiform and glabrous distally; anthers oblong, 
c. 3.5 by 0.5-0.8 mm, not rarely somewhat 
curved. Disk glabrous. Ovary elongate-conical, 
5-6(-8) by 1.5 mm, densely lepidote, gradually 
tapering to the style which is rather stout and 
lepidote in the lower half, 7-8(-10) mm; stigma 
capitate at anthesis, slightly 5-lobed, lobes accres- 
cent into separate club-shaped spreading parts 
afterwards. Capsule cylindric, often somewhat 
curved, 2.5-3.5(^.5) cm by 3-4 mni, densely 
lepidote, slightly 5-ribbed. Seeds 4-5 mm includ- 
ing the filiform tails; fruit-pedicel 1-1.5 mm o, up 
to 2.5 cm long. 

Distr. Males'ur. Borneo (Sarawak, Central and 
SE. Borneo). 

Ecol. Generally in moss forest, also found in 
low scrubby vegetation on bare sandstone rocks 
of B. Tanggoi summit, apparently but locally 
common, 800-2400 m. Fl. May, Aug. -Nov. 

78. Rhododendron fallacinum Sleum. Reinwardtia 
5(1960)99;Blumea 12 (1963) 100; Jahrb. Deutsch. 
Rhodod. Ges. Bremen (1965) f.7 & 8. — /?. 
duriunifolium (nan Becc.) Staff, Trans. Linn. Soc. 
II, Bot. 4 (1894) 197. — Fig. 17. 

Treelet or shrub, sometimes a tree, generally 
terrestrial, more rarely epiphytic, up to 6 m. 
Branchlets terete, robust, densely covered with 
the dark brown scales of the R. malayanum type 
which equally are found in the inflorescences and 
on the leaves; internodes 2-9 (sometimes up to 
30) cm. Leaves 3-4(-6) per pseudowhorl at the 
upper nodes, sessile or subsessile, ovate-oblong 
or subovate-lanceolate, apex shortly to rather 
long acuminate or subcaudate, subacute, base 
mostly slightly cordate-auriculate, (7-) 10-16 by 
(2-)2.5-6(-7) cm; petiole up to 3 by 3 mm. 
Inflorescences IO-30-flowered. Outer perulae ovate, 
inner ones obovatc-spathulatc, : caducously 
lepidote in the upper part of the mediane outside, 
otherwise papillate-puberulous, densely whitish 
ciliatc, up to 2 by 1 cm. Bractcolcs filiform and 
papillous below, spathulate and subvillous-ciliate 
above, up to 1.5 cm. Pedicels rather stoutish, 
mostly 2-3(-4) cm already at anthesis, rarely a 
little shorter then, densely lepidote. Calyx disk- 
like, minute, obsolctcly lobcd, c. 2 mm 0. Corolla 
funnel-shaped, light red, rose-purplish or red- 
orange, deep orange pink with a deep pink centre, 
also bright red, rarely dark red or pale scar'et, 
scentless, c. 2 cm long, subdensely brownish 
lepidote at the tube and the lower part of the 
lobes outside, otherwise completely glabrous, 
tube straight. 7-10 by c. 3 mm at the slightly 
5-saccatc base, lobes obovatc-spathulate to almost 
rounded, slightly cmarginatc, 1-1.6 by 0.8-1.1 
(-1.3) cm. Stamens 10, ; unequal, c. 1.8 cm; 
filaments linear, slightly dilated towards the base 
and rather densely and ; patently hairy there. 



filiform and glabrescent upwards; anthers elongate- 
oblong, c. 3 by 0.3 mm. Disk glabrous. Ovary 
elongate-conical, densely lepidote, 6-7 by 1.5 mm, 
gradually extending into the glabrous style (c. 
10 mm); stigma shortly turbinate-subcapitate, 
shortly 5-Iobed initially, ultimately 5-clubbed. 
Capsule golden brown or purple, cylindric, not 
rarely a little curved, 2.5-3 cm by c. 3 mm; 
fruit-pedicel rather slender, 2.5-3.5(-4) cm. 

Distr. Malesia: Borneo (North Borneo: Mt 
Kinabalu; Mt Trus Madi). 

Ecol. In mossy forest in damp shaded places, 
but also on ridge crests or open exposed places, 
among ferns and scrub rather frequent, 1220-2600 
m. Fl. fr. Jan. -Dec. 

Note. Sterile specimens hardly distinguishable 
from those of R. durionifolium Becc. 

79. Rhododendron apoanum Stein, Gartenflora 34 
(1885) 194, t. 1196; Jahresber. Schles. Ges. 
Vaterl. Cult. f. d. Jahr 1885 (1886) 414-416; 
ViDAL, Rev. PI. Vase. Filip. (1886) 172; Merr. 
Publ. Gov. Lab. Philip. /;. 29 (1905) 43; Philip. 
J. Sc. 3 (1908) Bot. 380; Elm. Leafl. Philip. Bot. 
3 (1911) 1107; MiLLAis, Rhodod. (1917) 115; 
Merr. En. Philip. 3 (1923) 243; Millais, Rhodod. 
ser. 2 (1924) 83; Corel./. Philip. J. Sc. 40 (1929) 
146; Sleum. Reinwardtia 5 (1960) 99; Blumea II 
(1961) 116. — Rhododendron sp. Vidal, Sinopsis 
Atl. (1883) 30, t. 60, f.E. — R. jasminiflorum {nan 
Hook.) F.-Vill. Nov. App. (1883) 353. 

Lax erect shrub, rarely subscandent, mostly 
epiphytic, c. 1 m, stems usually several fron\ the 
same root, profusely branched above the middle, 
the suberect twigs greyish brown. Branchlets 
terete, the upper internodes (2-7 cm) densely 
covered with dark rusty scales of the R. malayanum 
type, which are also found on inflorescences and 
leaves. Leaves close together towards the end of 
the upper internodes in a 3-5-merous pseudowhorl, 
lanceolate to elliptic-lanceolate, sometimes rather 
obovate-lanceolate or oblong, variable in shape 
and size, apex gradually or subabruptly acuminate, 
subacute or obtuse, base broadly to narrowly 
attenuate into the petiole, J coriaceous, dull 
green above, initially densely lepidote on both 
faces, glabrescent only above with age, sn\all pits 
remaining visible for a long time, entire to feebly 
dentate towards the apex, shortly revolute at the 
edge especially towards the base, (5-)6-ll by 
(|.5_)2-4 cm, midrib slightly impressed above, 
very prominent beneath in the lower part of the 
lamina, nerves 6-7 pairs, irregularly spreading, 
upper ones rather straight, all anastomosing, very 
slightly depressed above in fully mature leaves, 
somewhat raised or almost faintly visible beneath, 
reticulation 0; petiole scmiterctc, lepidote (0.9-) 
1-1.5 cm by 2-3 mm, grooved above, Flowers C)-\ 5 
per umbel. Perulae ovate, obtuse, outer ones 
densely lepidote in the dorsal mediane outside, 
inner ones progressively larger and less lepidote, 
all finely greyish sericeous the margin excepted, 
densely whitc-ciliate, up to I by 0.8 cm. Bractcolcs 
filiform, short-hairy, apex slightly spathulate 
and barbate, up to I cm. Pedicels rather stoutish, 



532 



Flora Malesiana 



[ser. I, vol. 6* 



densely lepidote, 7-1 1 mm. Calyx c. 2 mm 0, 
shortly 5-lobed, all over lepidote. Corolla tubular 
1.6-2 cm, purple to blood red, sometimes shiny 
bright red and a bit waxy, very laxly lepidote out- 
side at least at the sinuses of the lobes, tube 
straight, 1.1-1.5 cm long, 2-3 mm at the slightly 
5-pouched base, lobes subpatent, broadly obovate 
to nearly rounded, 5-7 by 4—6 mm. Stamens 10, 
about 2/3 as long as the corolla; filaments linear, 
flat, glabrous; anthers broad-oblong, c. 1.7 mm. 
Disk glabrous. Ovary subovoid-conical, 4-5 by 
2 mm, densely lepidote; style as long as or shorter 
than the ovary, rather thick, lepidote at the base, 
glabrous in the upper half; stigma thick-capitate. 
Capsule fusiform, 2-2.2 by c. 3 mm. Seeds c. 
2 mm including the filiform appendages. 

Distr. Malesia: Philippines (Mindanao: Mt 
Apo and other higher peaks). 

Ecol. Moss forest on peaks and ridges, 1380- 
2930 m, apparently not rare on Mt Apo. Fl. March, 
May-Oct. 

Vern. Kaliimping-busait, malagos, malagos- 
tabusa. Bag., saring-siiigon, Mbo. 

80. Rhododendron malayanum Jack, Mai. Misc. 

2 (1822) 17, reimpr. Hook. Bot. Misc. 1 (1834) 
369; DC. Prod. 7 (1839) 725; Planch. Fl. Serres 
5 (1849) 477-480 h; MiQ. Fl. Ind. Bat. 2 (1859) 
1059;Suppl. 1 (1860) 251; Hook. Bot. Mag. (1873) 
t. 6045; Becc. Malesia 1 (1878) 205; Clarke in 
Hook. /. Fl. Br. Ind. 3 (1882) 462; Henslow, 
J. R. Hort. Soc. 13 (1891) 262, f. 42 (Jol.), 43g 
{fl.)\ Ridl. J. Str. Br. R. As. Soc. n. 23 (1891) 145; 
ibid. n. 39 (1903) 15; K. & G. J. As. Soc. Beng. 
74, ii (1905) 78; Ridl. J. Linn. Soc. Bot. 38 (1908) 
313, reimpr. J. Fed. Mai. St. Mus. 2 (1908) 120; 
ibid. 4 (1909) 44; Koord.-Schum. Syst. Verz. 
(1912) fam. 233, p. 106; Koord. Exk. Fl. Java 3 
(1912) 6; Ridl. J. Fed. Mai. St. Mus. 5 (1914) 39; 
J. J. S. in K. & V. Bijdr. 13 (1914) 103; Ridl. J. 
Fed. Mai. St. Mus. 6 (1915) 157;MiLLAis,Rhodod. 
(1917)206;Merr. En. Born. (1921) 462; Burk. & 
Holtt. Card. Bull. S. S. 3 (1923) 57; Ridl. Fl. 
Mai. Pen. 2 (1923) 219; J. Mai. Br. R. As. Soc. 87 
(1923) 75; Burk. & Henders. Gard. Bull. S.S. 

3 (1925) 390; Sp. Moore, J. Bot. 63 (1925) Suppl. 
57; Hochr. Candollea 2 (1925) 492; Henders. 
J. Mai. Br. R. As. Soc. 5 (1927) 256; L Fed. Mai. 
St. Mus. 13 (1927) 5; J. J. S. in Merr. Contr. Arn. 
Arb. 8 (1934) 122; Burk. Diet. (1935) 1903; 
Symington, J. Mai. Br. R. As. Soc. 14 (1936) 
355; J. J. S. Bot. Jahrb. 68 (1937) 201; Hutch. 
in Craib, Fl. Siam. En. 2 (1938) 317; Steen. 
Bull. Jard. Bot. Btzg lO, 17 (1948) 387; Amsh. in 
Back. Bekn. Fl. Java (em. ed.) 7 (1948) fam. 162, 
p. 3; Henders. Mai. Nat. J. 6 (1950) 262; Merr. 
J. Arn. Arb. 33 (1952) 243; Sleum. Blumea Suppl. 

4 (1958) 48; Reinwardtia 5 (1960) 100; Blumea 11 
(1961) 117; Steen. Endeavour 21 (1962) 189, t. 3; 
Sleum. Blumea 12 (1963) 101; Jahrb. Deutsch. 
Rhodod. Ges. Bremen (1965) f. 2. — R. tubiflorum 
R(einwardt) in Bl. Cat. Gew. PI. Btzg (1823) 
72, nom.nud.; DC. Prod. 7 (1839) 725; Hassk. 
Cat. Hort. Bog. (1844) 162; Moritzi, Syst. Verz. 
(1846) 42; Planch. Fl. Serres 5 (1849) 477-480 h; 



ZoLL. Syst. Verz. 2 (1854) 137; Mig. Fl. Ind. Bat. 
2 (1859) 1059; Ann. Mus. Bot. Lugd.-Bat. 1 
(1863) 44, t. II A; Bl. Fl. Jav. Pl. ined. (1863-83) 
t. 5. — Vireya tubiflora Bl. Bijdr. (1826) 855; 
Don, Gen. Syst. 3 (1834) 848. — Azalea tubiflora 
Bl. ex DC. Prod. 7 (1839) 725 in syn. — R.fuscum 
Bl. pi. Jav. PI. in6d. (1863-83) t. 7 C, f.l, cf. 
Steen. Bull. Jard. Bot. Btzg III, 17 (1948) 388. — 
Azalea malayana (Jack) O. Ktze, Rev. Gen. PI. 2 
(1891) 387. 



key to varieties and forms 

1. Inflorescences exclusively terminal. 
2. Filaments glabrous. . . 1. var. malayanum 
2. Filaments ± densely short-pubescent in basal 

3. Corolla besides some scales laxly short- 
pubescent outside 2. var. pubens 

3. Corolla besides some scales glabrous. 

3. var pUosifilum 
1. Inflorescences both terminal and axillary. 
4. Filaments glabrous. 
5. Leaves oblong-lanceolate, ± acute at apex 
and base . . . 5a. var. axillare /. axillare 
5. Leaves elliptic or ovate. 
6. Leaves broadly elliptic, apex shortly ob- 
tusely attenuate, base i truncate. 

5b. var. axillare/. latifolium 
6. Leaves ovate, apex shortly obtusely acumi- 
nate, base rounded 5c. var. axillare /. ovatum 
4. Filaments ± densely short-pubescent in the 
lower V3-V2 4. var. infrapilosum 

1. var. malayanum. 

Erect, straggling shrub, not rarely crooked, 
terrestrial or frequently epiphytic, sometimes 
treelet up to 5 m. Branchlets terete, slender, tips as 
all parts of the plant very densely covered with 
flat, ± roundish scales of two different sizes, very 
manifest to the naked eye; scales very close, 
touching or partly overlapping each other, the 
marginal zone thin, brownish, becoming ± silvery 
with age, ± irregularly and mostly deeply crenate 
or lobed or cleft, the centre dark chestnut-brown 
to sooty, large in relation to the marginal zone, 
leaving a small dark pit when the scale has gone; 
internodes 2-10 cm. Leaves subopposite or in a 
3-6-merous pseudowhorl at the end of the upper 
1-3 internodes, ± ascending, oblong- to lanceolate- 
elliptic, sometimes lanceolate or oblanceolate, 
variable in shape and size, always attenuate at 
both ends, apex mostly gradually short-acuminate, 
the very apex obtuse by a small gland, base 
acutely attenuate into the petiole, coriaceous, dark 
green and shining above, rubiginose to copper- 
coloured beneath, entirely covered with scales on 
both faces initially, glabrescent above with age, 
persistently lepidote beneath, entire or indistinctly 
repand-denticulate in the upper half, not rarely 
slightly revolute especially towards the base, (4-) 
5-10(-15) by (l-)1.5-3(-4) cm, midrib somewhat 
depressed above, bold and prominent beneath 
in the lower part, gradually less so distally, nerves 
in (5-)6-8 pairs, ± irregular, mostly rather 



Jan. 1966] 



Ericaceae (Sleiimer) 



533 



straight, anastomosing before the edge, a little 
sunk in mature leaves, not rarely rather obscure 
above, slightly or not raised beneath, reticulation 
mostly not visible; petiole stout, grooved above, ± 
semiterete, densely lepidote, (l-)1.2-1.8(-2.2) cm. 
Umbels exclusively terminal, (3-)4-8-flowered; 
flowers pendulous. Ferulae ovate-triangular or 
ovate to spathulate, densely lepidote dorsally in 
their middle, otherwise glabrous or finely sericeous, 
densely whitish ciliate, 0.3-0.8(-l) by 0.2-0.4 
(-0.6) cm. Bracteoles filiform, slightly dilated at 
the ape.x and barbate there, otherwise glabrous, 
up to 1.5 cm. Pedicels rather slender, — densely 
pubescent to nearly short-velutinous, the hairs 
mostly completely covered by the scales, 6-9(-15) 
mm. Calyx — oblique, small, 5-dentate or -lobed, 
densely lepidote, lobes — obtuse, sometimes up to 
1 mm. Corolla tubular in general, somewhat dilated 
towards the lobes which are — spreading in full 
anthesis, waxy and glossy, bright red to scarlet- 
crimson or purplish, rarely salmon-red or rose- 
pink, sometimes yellowish at the tube, subdensely 
or mostly sparsely lepidote outside, at least at the 
sinuses of the lobes, glabrous inside, (l.5)-1.7-2.2 
f-2.4, rarely up to 2.8) cm, tube ± straight or 
slightly curved, somewhat laterally compressed, 
slightly 5-saccate and 3 (-4) mm at base, 4—5 mm 
at throat, lobes broadly obovate to nearly rounded, 
5-7 by c. 4 mm. Stamens 10, unequal, as long as or 
slightly longer than the corolla tube; filaments 
linear, nearly filiform above, glabrous, i papillose; 
anthers broad-oblong, 1.2-1.6 mm. Disk glabrous 
where not covered with scales. Ovary oblong- 
conical, densely lepidote, 4-5 by 1.5-2 mm, 
tapering to the style, the latter slender, equalling 
or mostly slightly surpassing the stamens in 
length, or slightly e.xserted from the throat of the 
corolla, l-l.5(-2) cm, mostly lepidote in the lower 
y^ to '/j, rarely so to 1/2, or glabrous except the 
very base; stigma shortly club-shaped. Capsule 
subcylindric, 2.2-2.6(-2.8) by 3(-4) mm. Seeds 
very narrow, 5 mm including the tails. 

Distr. S. Siam, in Malesia: Sumatra, Malay 
Peninsula, W. Java, Borneo, Celebes. 

Ecol. Usually epiphytic in primary and second- 
ary rain-forest from (550-) 1 000-2400 (in Celebes 
and .Sumatra up to 3000) m, but also terrestrial 
on peaty soil, stony or sandy ground, quartzite 
cliffs, hard sandstone ridges or exposed rocks or 
crests, on slopes or margins of craters, where sub- 
alpine scrub vegetation dominates, in mossy 
heath forest on Mt Kinabalu, not rarely common 
locally. Fl. Jan. -Dec. 

Vcrn. Api-api, kaju si taltom liunik, M, Tapa- 
nuli, hunffoi rorun^, I ban. 

Note. The flowers of R. malayanum are appar- 
ently constantly somewhat larger in Celebes, 
compared with those of the other islands, attain- 
ing the largest size given above. 

2. var. pubcas Sleum. Reinwardtia 5 (1960) 102. — 
R. malayanum; J. J. S. in f-"cddc. Rep. 30 (1932) 
164, p.p. 

Corolla distinctly short-pubcsccnt in- and out- 
side, 3 cm. Filaments pubescent below. Leaves 



8-12 by 2.5-5 cm. Inflorescences terminal. 

Distr. Malesia: Moluccas (W. Ceram: Ora 
Mts), 1000 m, rare on steep limestone ridge, once 
found. Fl. May. 

3. var. pilosifilum Sleum. Reinwardtia 5 (1960) 102; 
Blumea 11 (1961) 117. — /?. malayanum; J. J. S. 
in Fedde, Rep. 30 (1932) 164, p.p. 

Pedicels finely patent-hairy. Corolla rose, gla- 
brous. Filaments laxly to subdensely hairy in the 
lower half. Inflorescences terminal. 

Distr. Malesia: Moluccas (Central Ceram and 
Buru), 1075-1700 m, on peat moor respectively 
light summit forest on limestone, and in N. 
Borneo on Mt Kinabalu rare among bracken, 
scrub and grasses at c. 2440 m. Fl. Dec, Feb., 
March. 

4. var. infrapilosum Sleum. Reinwardtia 5 (1960) 
103; Blumea 12 (1963) 101. 

Leaves subovate- or subelliptic-lanceolate, not 
rarely obtuse to subcordate at the base, 9-17 by 
3-4.5 cm. Filaments laxly to subdensely hairy in 
the lower Vi-Vi- Inflorescences both terminal and 
axillary. 

Distr. Malesia: Borneo (Sarawak and Central 
Borneo at Sarawak border), 1700-1830 m. Fl. 
Oct.-Dec. 

5a. var. axiUare J. J. S. Ic. Bog. 4 (1919)74, note, 
/. axillare; Sleum. Reinwardtia 5 (1960) 103; 
Blumea 11 (1961) 117. 

Inflorescences both terminal and axillary on the 
same specimen. Leaves oblong-lanceolate, 12-20 
by 4-6 cm. 

Distr. Malesia: Borneo; here and there, 
possibly together with the typical variety. Such 
forms with both terminal and axillary inflore- 
scences are limited to Borneo. 

5b. var. axillare J. J. S. /. latifolium Sleum. 
Reinwardtia 5 (I960) 103. 

Leaves (broadly) elliptic, apex shortly obtusely 
attenuate, base ± truncate-obtuse, 9-14 by (4-) 
5-7.5 cm. Inflorescences both terminal and 
axillary. 

Distr. Malesia: Borneo (Sarawak and SE. 
Borneo). Fl. Oct. 

5c. var. axillare J. J. S./. ovatum Sleum. Reinward- 
tia 5 (I960) 103. 

Leaves ovate, base rounded, 9-15 by 5-8 cm. 
Inflorescences both terminal and axillary. 

Distr. Malesia: Borneo (Sarawak, Brunei and 
Central Borneo). 

Ficol. In Brunei found gregarious as a terres- 
trial plant in extreme heath ('keran^as') forest 
at low level (r. 260 m), in heath forest at 550 m, 
and in mossy forest Ccllin woodland') at 1525 m, 
always on deep peat between hard sandstone 
boulders on sharp narrow ridges. //. April. 

Vcrn. Ihini^ai rorun^, Iban, Brunei. 

81. Rhododendron nortonlae) Mikk. Philip. J. Sc. 
1 (1906) Suppl. 220; (/;/(/. 3 (1908) Bot. 380; 



534 



Flora Malesiana 



[ser. I, vol 6*. 



Elm. Leaf). Philip. Bot. 3 (1911) 1107; Millais, 
Rhodod. (1917) 218; Merr. En. Philip. 3 (1923) 
244; Millais, Rhodod, ser. 2 (1924) 195; Copel./. 
Philip. J. Sc. 40 (1929) 149, t. 2, f. 2-4; Sleum. 
Reinwardtia 5 (1960) 103. — R. catanduanense 
Merr. in Copel. /. Philip. J. Sc. 40 (1929) 149, 
t. 2, f. 5-6. 

Epiphytic shrub. Branches terete, brown; 
branchlets (as inflorescences and leaves) densely 
covered with dark brown scales of the type 
observed in R. malayanum. Leaves in pseudo- 
whorls at the upper nodes, lanceolate to oblong- 
lanceolate or narrow-elliptic, acuminate to the 
rather slender long apex, attenuate to the acute 
base, coriaceous, glabrous and shining above, 
densely lepidote beneath, very similar to those of 
R. malayanum and R. apoaniim, with slightly 
recurved margin, (5-)8-15 by 1.5-3.5 cm, mid- 
rib impressed above, prominent beneath, nerves 
5-6 distant pairs, ascending, slightly impressed 
above and raised beneath, reticulation obscure; 
petiole stoutish, densely lepidote, c. 1 cm. Umbels 
± 12-flowered. Perulae ovate to oblong-ovate, 
subacute, glabrous dorsally, ciliate, 1-1.7 cm. 
Bracteoles filiform. Pedicels rather slender, patent- 
puberulous, not lepidote (seen only in fruit-stage), 
l-1.5(-2) cm. Ca/yA" disk-like, c. 3 imn 0, obscurely 
and obtusely lobed, lobes short-ciliate. Corolla 
tubular, crimson or scarlet, 4-4.5 cm, glabrous 
outside, pubescent inside along the 5 main nerves, 
tube cylindric, c. 3.5 cm, scarcely widened distally, 
slightly curved, 6-7 mm 0, lobes broad-obovate 
to nearly roimded, 8-10 by 6-8 mm. Stamens 10, 
slightly unequal, as long as the corolla or slightly 
exserted; filaments filiform, densely brownish 
pubescent in the lower third; anthers oblong, 
3.5 mm (in the Negros specimen only 2 mm). 
Disk glabrous. Ovary narrow-oblong, densely 
lepidote; style elongated, filiform, densely lepidote 
in the lower half only, nearly as long as the corolla; 
stigma turbinate. Capsule cylindric, slightly 
curved, densely rusty-lepidote, (4-)5-8 by c. 3 
mm. Seeds 4-5 mm long including the filiform 
tails. 

Distr. Malesia: Philippines (Mindanao: near 
Lake Lanao, 800 m; Catanduanes: Mt Mariguidon, 
270 m; S. Negros: Cuernos Mts). 

Vern. Malagas, Catand. 

Note. There are slight diff"erences between the 
3 specimens from the above mentioned localities 
in size of leaves, anthers, and capsules. 

82. Rhododendron acuminatum Hook. /. in Hook. 
Ic. PI. (1852) t. 886; Walp. Ann. 5 (1858) 446; 
Becc. Malesia 1 (1878) 207, p.p.; Stapf, Trans. 
Linn. Soc. H, Bot. 4 (1894) 197; Gibbs, J. Linn. 
Soc. Bot. 42 (1914) 104; Merr. En. Born. (1921) 
460; Sleum. Reinwardtia 5 (1960) 104; Blumea 12 
(1963) 101; Jahrb. Deutsch. Rhodod. Ges. Bremen 
(1965) f. 13. —Fig. 18. 

Small tree or shrub, c. 3, rarely up to 6 m. 
Branches few, robust. Branchlets short, much 
divaricate, upper parts subterete to obtusely 
angular especially towards the end of each inter- 
node; upper internodes 3-7 cm, densely dark 




Fig. 18. Rhododendron acuminatum Hook./. Mt 

Kinabalu, 2600 m, N. Borneo (Sleumer 4691) 

(Sleumer, 1963). 



lepidote at the ends, more silvery so below. 
Leaves 3-5 (-6) close together at the upper nodes 
or condensed in pseudowhorls, ovate or elliptic- 
ovate, sometimes ovate-lanceolate, apex rather 
short-acuminate, ± obtuse, base mostly rounded, 
sometimes very broadly attenuate, stiff coriaceous, 
dark green and glossy above, densely scaly on 
both faces initially, glabrescent above (pits of the 
gone scales remaining visible), permanently cover- 
ed beneath with scales of the type found in R. 
malayanum, edge not or slightly revolute, 6-10 
(-12) by 2.5-5(-6) cm, midrib strongly impressed 
above, very thick and rigid beneath, continuing 
the petiole and gradually narrower apically, 
nerves 6-8 pairs, spreading, straight or slightly 
curved to the edge, distinctly anastomosing, — 
deeply depressed above, ri strongly prominent 
beneath, veins laxly reticulate and iz manifestly 
impressed above, not or slightly raised beneath, 
the whole lamina thus rather coarsely — bullate 
at least in the dry state; petiole subterete, grooved 
above, densely lepidote, (1-) 1.2-2 cm by 2-3 mm. 
Flowers (7-) 10-1 5 per umbel. Flower buds ovoid, 
c. 2.5-1.5 cm. Perulae ovate, truncate at the base, 
obtuse at the apex, greenish, all over covered 
with a very fine brownish or greyish puberulence 
outside, moreover lepidote outside along the 
mediane and/or at the apex, but glabrous at the 
thin margin, up to 2 by 1.3 cm. Bracteoles linear 
to filiform, subspathulate-dilated at the top, 
patent-puberulent, up to 1.5 cm. Pedicels rather 
stoutish, densely scaly, puberulent among the 
scales, 8-12 mm in flower. Calyx cup-shaped, 
shallowly 5-lobed, c. 2.5 mm 0, densely lepidote. 
Corolla tubular-subinfundibuliform, red, scarlet or 
purple to deep pinkish orange, 2.5-2.8 cm, sub- 
densely to sparsely and caducously brownish 
lepidote, finally — glabrescent outside, shortly 
subdensely pubescent at the lower half of the tube 
inside, tube 1.5-1.7(-2) cm, rather straight, 
cylindric, c. 4 mm at the slightly 5-pouched base, 
somewhat widened to the mouth, lobes subpatent, 



Jan. 1966] 



Ericaceae (Sleumer) 



535 



obovate to almost rounded, (6-)8-10 by 5-8 
mm. Stamens 10, unequal, c. 2 cm; filaments linear, 
flattened to nearly the top, rather densely whitish 
patent-hairy in the lower third; anthers broad- 
oblong, 2.5-2.8 by 1 mm, no basal appendages. 
Disk glabrous. Ovary cylindrical, c. 5 by 2 mm, 
densely squamular-lepidote, gradually tapering 
to the rather stoutish style, which is lepidote in its 
lower half; stigma thick, turbinate, with several 
lobes. Capsule cylindric, short-attenuate at both 
ends, densely lepidote. 3-3.5 cm by 4-5 mm. 

Distr. Malesia: North Borneo (Mt Kinabalu). 

Ecol. In mossy forest and scrub forest, on 
ridge top, 1830-3350 m, not rare. Fl. Jan.-Dec. 

83. Rhododendron fortunans J. J. S. Bull. Jard. Bot. 
Btzg III, 1 (1920) 401, t. 49; Sleum. Reinwardtia 5 
(1960) 104; Blumea 12 (1963) 101. 

Shrub. Branchlets stoutish, subangular, upper 
parts (as inflorescences and leaves) densely 
squamular brown-lepidote as in R. malayanum; 
internodes 2-7 cm. Leaves 3-5 crowded near the 
end of the upper node, ±: set in a pseudowhorl, 
sometimes subopposite, elliptic-lanceolate, apex 
rather long subcaudate-acuminate, subacute, base 
acutely attenuate into the petiole or rarely obtuse, 
coriaceous, dark olivaceous to brownish when 
dry above, dark rubiginose beneath, lepidote on 
both faces initially, glabrescent above with age 
(dark pits of the fallen scales left), completely 
covered with dark brown scales beneath, edge 
slightly revolute especially to the base, (6.5-)7-13 
by 2.3-4 cm, midrib very stout beneath, gradually 
decreasing apically, nerves 5-10 pairs, ± patent 
and partly rather straight, manifestly anastomos- 
ing, very impressed above and prominent beneath, 
veins laxly reticulate, lamina i markedly bullate; 
petiole somewhat flattened, grooved above, densely 
lepidote, 5-IO(-12) by 2-3 mm. Umbels 10-13- 
flowered. Flower buds ovoid, large, perulae ovate 
to obovate-spathulate, outer ones all over puber- 
ulent, lepidote in the upper part, inner ones lepi- 
dote only, up to 2.4 by 1.2 cm. Bracteoles linear 
below, spathulate towards the apex, ciliolate, 
up to 1.6 cm. Pedicels rather stoutish, densely 
lepidote, 1.5-2 cm. Calyx suboblique, small, cup- 
shaped, shortly obtusely 5-Iobed, lepidote. Corolla 
white, widely funnel-shaped, c. 1.5 cm long in all, 
c. 2 cm at limb, 5-lobed, tube 3-4 mm long, 
very densely brownish lepidote at the tube and the 
lower half of the lobes outside, densely pubescent 
at the tube inside except the spots where the fila- 
ments arc inserted, glabrous at the lobes inside, 
lobes suborbicular, 9-12 mm 0. Stamens 10, c. 
9 mm; filaments linear, glabrous at the very base, 
short-villous above, more laxly pubescent up- 
wards to the top; anthers elongate, c. 3 mm, 
thccac divergent below, each bearing a basal 
conical obtuse lobe or appcndicc. Disk glabrous. 
Ovary conical, 2.5 by 1.7 mm, densely lepidote, 
gradually attenuate into a rather thick glabrous 
style (2.5 mm); stigma thick, turbinate. Capsule 
short-cylindric, c. 2.3 by c. 0.3 cm; fruit-pedicel c. 
2 cm. 

Distr. Malesia: Central Borneo (top of B. 



Radja in the Schwaner Mts), c. 2000 m, said to be 
common there. Fl. Oct., Dec. 
Vern. Daun tali, kaju tari, Daj. 

84. Rhododendron lineare Merr. J. Str. Br. R. As. 
Soc. n. 76 (1917) 108; En. Born. (1921)462; Sleum. 
Reinwardtia 5 (1960) 104; Blumea 11 (1961) 117. 
Epiphytic or terrestrial shrub, up to 2 m. 
Branchlets slender, terete, irregular; internodes 
1.5-6 cm, the upper 2-4 ones densely lepidote. 
Leaves in 3-5-merous pseudowhorls, mostly 
present only at the end of the uppermost 2 nodes, 
linear to linear-lanceolate, gradually attenuate 
towards the obtuse apex, base very broadly 
attenuate, or mostly subtruncate-rounded, cori- 
aceous, glabrous and shining above in the mature 
state, densely lepidote beneath (scales in general of 
the type which is found in R. malayanum, but 
much less distinctly of two different sizes, although 
larger ones are present, the majority are small 
spaced ones with relatively few ones partly over- 
lapping), margin somewhat cartilagineous, wavy, 
mostly revolute in dry state, (2.8-)3-4(-5) cm by 
(3-)4-5 (very rarely up to 8) mm, midrib strongly 
impressed on the upper surface, sharply prominent 
beneath, nerves obscure; petiole rather thick, 
densely lepidote, c. 2 mm. Umbels 2-3 (rarely up 
to 4)-flowered. Flower buds elongate-ovate, sub- 
acute, outer perulae narrow ovate, inner ones 
obovate-oblongto subspathulate, all short-acumin- 
nate, densely lepidote dorsally in their upper part, 
densely whitish ciliate towards the apex, up to 
1.5 cm by 7 mm. Bracteoles narrow linear, 
barbate-ciliate at the top, up to 1.2 cm. Pedicels 
rather stoutish, covered with squamular scales 
which are easily detached, 8-10 mm. Calyx very 
small, oblique, disk-shaped, c. 2 mm 0, obscurely 
lobed, densely lepidote. Corolla infundibuliform, 
i.e. tubular below, campanulate above, bright 
(primrose) yellow with reddish brown external 
spots (scales!), c. 2 cm, densely lepidote at the 
tube, laxly so on the lower half of the lobes outside, 
glabrous inside, tube straight, cylindrical, 7-8 mm, 
c. 3 mm at the slightly saccate base, lobes 
spreading, oblong-obovate, apex rounded, 10-12 
by 6-7 mm. Stamens 10, slightly unequal, nearly 
as long as the corolla; filaments filiform above, 
dilated towards the base, rather densely whitish 
patent-hairy in the lowest 3 mm; anthers narrow- 
oblong, r. 3 mm by 0.5 mm, thecae somewhat 
pointed at the base. Disk rather thick, glabrous. 
Ovary subcylindric, r. 4 by 1.5 mm, densely 
lepidote, gradually tapering to a rather thick ish 
style, which is almost as long as the corolla and 
lepidote in its lower half; stigma rather thick- 
obconical, distinctly 5-lobcd. Capsule cylindric, 
densely lepidote, 1-1.2 cm by c. 2.5 mm. 

Distr. Malesia: Borneo (Sarawak; Brunei). 
Ecol. Moss forest, epiphytic or on moss covered 
ground, 360-1230 m. Fl. Junc-Oct. 

85. Rhododendron obscurum Sleum. Reinwardtia 

5 (1960) 104. 

Shrub, r. 3 m. Branchlets subdcnsely minutely 
rufou.s lepidote; ultimate internodes c. 3 cm by 



536 



Flora Malesiana 



[ser. I, vol. 6^ 



2 mm. Leaves in 3-4-merous pseudowhorls at the 
upper 2 or 3 nodes, oblong-obovate, apex rounded 
or mostly slightly retuse, base ± broadly cuneate, 
coriaceous, lepidote initially on both faces, at full 
maturity ± glabrescent above, subdensely and 
persistently lepidote beneath (scales minute, the 
thin marginal zone variously dented, the centre 
but little impressed, rather large and of the chest- 
nut-brown colour typical for those of R. mala- 
yanum), fiat, 4.5-7.5 by (2-)2.3-3 cm, midrib 
narrow and immersed above, as thick as the petiole 
beneath at the base and much prominent, gradu- 
ally less so apically, nerves 7-9 pairs, suberect 
below, curved anastomosing towards the edge, 
minutely impressed above, somewhat raised or 
nearly invisible beneath, no reticulation; petiole 
a little flattened, grooved above, densely lepidote, 
6-8 by c. 2 mm. Umbels ± 3 flowered. Flower 
bud scales ovate-acuminate, subsericeous-puber- 
ulous on both sides, white-c'liate, c. 7 by 4 mm. 
Pedicels rather slender, laxly lepidote, 1.4-1.8 cm. 
Calyx oblique, disk-shaped, very shortly or not 
lobed, lepidote dorsally, c. 2.5 mm 0. Corolla 
tubular, slightly curved, gradually widened to the 
mouth, pink, 2.6-3 cm long in all, laxly lepidote 
outside, glabrous inside, tube c. 2 cm, at the base 
3-4, at the apex c. 6 mm 0, lobes obovate, sub- 
erect, 0.7-1 by 0.4—0,5 cm. Stamens 10, unequal, 
the longer ones nearly as long as the corolla; 
filaments linear, glabrous; anthers oblong-obovate, 
c. 1.5 by 0.7 mm. Disk glabrous. Ovary elongate- 
conical, densely lepidote, c. 4 by 1.5 mm, gradually 
tapering to the s'ender, somewhat exserted style; 
stigma subcapitate. Capsule c. 1 cm. 

Distr. Malesia: Malay Peninsula (Pahang: top 
of Mt Benom), 2105 m, once found. Fl. May. 

Note. Apparently a natural hybrid of R. 
malayanum with a species oi subsect. Pseudovireya 
(13. R. perakense, 16. R. scortechinii, or 10. R. 
spathulatum). 

86. Rhododendron variolosum Becc. Malesia 1 
(1878) 206; Merr. En.Born. (1921) 463; Sarawak 
Mus. J. 3 (1928) 543; Sleum. Reinwardtia 5 
(1960) 105; Blumea 12 (1963) 101. 

var. variolosum. 

Terrestrial or epiphytic shrub, 0.5-0.8 m, or 
treelet up to 1.6 m; branchlets subterete, rather 
stout; internodes 2-12 cm, upper ones densely 
squamular-lepidote, lower ones glabrous, minutely 
tubercled. Leaves scattered, rather close together 
in the upper part of the upper nodes, lanceolate to 
elliptic-lanceolate, rarely elliptic, gradually attenu- 
ate towards the apex and base, apex subacute, the 
very base mostly obtuse before running into the 
petiole, coriaceous, glabrous and shining above in 
mature state, some scattered scales excepted (fallen 
scales leaving no distinct pits), densely covered 
beneath with scales (these in general of the type 
found in R. malayanum, but the larger ones sparse 
and less diff"erent in size than in R. malayanum, 
rather small and largely spaced initially, but some 
touching or overlapping, all distinctly impressed), 
narrowly revolute, (7-)8-12(-16) by (2-)2.5-4 



(-5.5) cm, midrib deeply depressed above, strongly 
raised beneath, nerves 10(-12)pairs,ratherstraight, 
curved-anastomosing before the edge, all slightly 
impressed above, prominent beneath as are the 
reticulate veins; petiole flattened, grooved above, 
densely lepidote, (1-) 1.5-2. 5 cm by 2-3 mm. 
Umbels terminal and axillary (the latter apparently 
not well developed), 5-8(-12)-flowered. Flower 
buds oblong-ellipsoid, acuminate, perulae ovate- 
oblong to subspathulate, outer ones acutely 
acuminate, lepidote upwards dorsally, up to 1.5 by 
0.7 cm. Bracteoles filiform, glabrous, up to 1 cm. 
Pedicels nodding, slender, densely lepidote, 1-1.5 
(-2) cm. Calyx disk-shaped, rather oblique, wavy, 
obscurely lobed, densely lepidote, c. 3 mm 0. 
Corolla pink, or scarlet with yellow tube, infundi- 
buliform, 2.5-2.8(-3) cm, very laxly lepidote or 
glabrous outside, entirely glabrous inside, tube 
cylindric, i straight, saccate at the base, 1.2-1.5 
cm, 3-4 mm at the base, 6 mm at throat, lobes 
obovate-spathulate, rounded-obtuse, spreading, 
1.1-1.3(-1.5) by 0.6-0.8 cm. Stamens 10, unequal, 
1.7-2 cm; filaments linear, glabrous; anthers al- 
most rectangular, c. 2 by 0.7 mm. Disk glabrous. 
Ovary elongate-conical, densely lepidote, c. 5 by 
1.5 mm, deeply 5-ribbed, gradually tapering to 
a rather thick style as long as the stamens and 
lepidote in its lower 2/3, glabrous elsewhere; 
stigma short-turbinate or subcapitate, shortly 
5-lobed. 

Distr. Malesia: Borneo (Sarawak: Mt Pueh, 
c. 1500 m; N. Borneo: on crest of Mt Trus Madi, 
c. 2600 m). 

var. andersonii (Ridl.) Sleum. Reinwardtia 5 
(1960) 105; Blumea 11 (1961) 117; ibid. 12 (1963) 
101. — R. andersonii Ridl. Kew BuU. (1914) 209; 
Merr. En. Born. (1921) 460. 

Filaments rather densely patent-hairy in the 
lower third. Corolla sometimes pubescent inside 
at the tube, of a pink to deep red colour, often 
with yellow corolla tube or base. 

Distr. Malesia: Borneo (Sarawak, Central 
Borneo, and Mt Kinabalu in North Borneo). 

Ecol. In open exposed crest of spur, among 
bracken, scrub and grasses, rare on Mt Kinabalu, 
where it grows from 2345 to 2440 m, and almost 
certainly a natural hybrid of R. malayanum with 
a yellow-flowered species oi ser. Javanica. 

Vern. Bunga klangin, I ban. 

87. Rhododendron wilhelminae Hochr. Candollea 
2 (1925) 493; Sleum. Reinwardtia 5 (1960) 106. 
Shrub, c. 2 m. Branchlets somewhat flattened 
at the ends, lepidote; internodes 5-8 cm. Leaves in 
4-5-merous pseudowhorls at the upper 1 or 2 
nodes (one leaf mostly inserted a little below the 
node), broadly lanceolate, apex subacutely acumi- 
nate, base cuneate, coriaceous, lepidote on both 
faces initially, ± glabrescent and shining with 
age, persistently lepidote beneath (scales rather 
flaky, dense but spaced 1-3 times their own 
diameter, light brown marginal zone rather 
narrow, irregularly dented or lobed, centre rela- 
tively large, dark chestnut-brown as in R. mala- 



Jan. 1966] 



Ericaceae (Sleumer) 



537 



yanum), edge slightly recurved especially towards 
the base of the lamina, 8-10 by 2.6-3.7 cm, midrib 
slightly impressed above, much prominent be- 
neath, nerves 8-9(-10) pairs, slightly raised 
beneath only, reticulation obsolete; petiole — 
flattened, lepidote, 8-15 by c. 2 mm. Umbels — 
4-flowered. Flower buds elongate-ovoid, c. 1.7 by 
0.5 cm. Ferulae oblong-ovate, subobtuse, glabrous 
in- and externally, white-ciliate towards the apex, 
up to 1.5 by 0.7 cm. Bracteoles linear. Pedicels 
rather stoutish, — densely lepidote and finely 
patent-pubescent, 1.2-1.6 cm. Calyx oblique, very 
shortly subacutely lobed, or occasionally one lobe 
elongated up to 2 mm. Corolla tubular-funnelform, 
c. 3.5 cm long in all, c. 2.5 cm at the limb, 
carmine, practically glabrous outside, tube c. 
1.8 cm, base somewhat pouched and 4-5 mm 0, 
widened to c. 6 mm at throat, lobes obovate- 
spathulate, c. 1.5 by 1 cm. Stamens 10, very un- 
equal, the longest c. 2.6 cm; filaments linear 
below, somewhat broadened above the base, — 
densely patently hairy in the lower third, narrower 
and glabrous towards the top; anthers elongate- 
obovate, c. 2.5 by 0.5 mm, containing pollen. Disk 
hairy on the upper margin, glabrous below. Ovary 
subcylindric, densely lepidote and laxly finely 
patent-pubescent, c. 4 by 1 .5 mm; style continuous, 
subdensely patent-pubescent for c. 2 mm at the 
base, glabrous for the rest, c. 2.2 cm; stigma 
distinctly 5-lobed. 

Distr. Malesia: W.Java (MtSalak), c. 1350 m, 
once found in shrubbery near the crater, very 
rare. Fl. Oct. 

Note. The characters of R. wilhelminae show a 
combination of those of 225. R. javanicum var. 
feysmannii and /?. malayanum, which both are 
found on Mt Sa'ak, and suggest a natural hybrid 
of these species. 

A garden-hybrid "Monarch" (of the formula 
jasminiflorum > Javanicum / longiflorum / broo- 
keanum) has been crossed with R. malayanum; the 
flower of this hybrid, figured by Henslow, J. R. 
Hort. Soc. 13 (189!) 276, f. 44 n, reminds much 
of that of R. wilhelminae. 

88. Rhododendron hybridogcnum Sleum. Rein- 
wardtia 5 (I960) 106. 

Shrub. Branchlets rather slender, densely lepi- 
dote; intcrnodes 4-7.5 cm by c. 2 mm. Leaves in 
4-merous pseudowhorls at the upper 2 nodes. 



subovate-elliptic, apex shorter or longer sub- 
acutely acuminate, base rounded or mostly 
slightly cordate, coriaceous, rather densely set 
with small scales or their relics above at maturity, 
very densely and persistently lepidote beneath 
(scales rather small, often contiguous, narrow 
marginal zone dented or lobed to various degree, 
centre slightly impressed, relatively large and of 
the chestnut or russet-brown colour found in 
R. malayanum), slightly revolute towards the base, 
4.5-6 by (2-)2.5-2.9 cm, midrib much dilated at 
the base only, further up abruptly narrowed 
and slightly impressed above, as thick as the 
petiole at the base, gradually narrower and less 
prominent distally beneath, nerves (6-J8-10 pairs, 
slightly curved, inarching near the edge, faintly 
depressed above, slightly raised beneath, no 
reticulation; petiole densely lepidote, somewhat 
flattened, 4-6 by c. 2 mm. Umbels — 8-flowered. 
Ferulae ovate-acuminate, apiculate, subsericeous 
dorsally, lepidote towards the top, obscurely 
ciliolate, up to 8 by 6 mm. Bracteoles linear, 
subglabrous, 5 mm. Fedicels subdensely lepidote, 
more laxly short-patent-pubescent, 1-2 cm by 0.5 
mm at anthesis. Calyx obliquely disk-shaped, 
indistinctly lobed, c. 3 mm 0. Corolla tubular, 
said to be yellowish, c. 3 cm long in all, outside 
laxly lepidote and especially in the lower half 
laxly patent-pubescent, inside finely hairy, tube 
but a little widened upwards, c. 2 by 0.3 cm, lobes 
obovate-spathulate, :•: expanded, c. 1 by 0.5-0.6 
cm. Stamens 10, unequal, c. 2.5 cm; filaments 
linear, densely hairy below, less so towards the 
middle, narrower and glabrous above; anthers 
obovate-oblong, c. 1.5 by 0.7 mm. Disk somewhat 
prominent, hairy at the upper margin, glabrous 
otherwise. Ovary elongate ovoid, continuous with 
the style, very densely lepidote, with a hair here 
and there between the scales, r. 5 by 2 mm; style 
slender, c. 2.8 cm, subdensely lepidote and laxly 
patent-pubescent to the lower V}, glabrous for 
the rest; stigma subcapitate, distinctly 5-lobed. 

Distr. Malesia: Malay Feninsula (Pahang: 
Cameron Highlands). Once found. Fl. Oct. 

Note. R. liybridogenum most certainly is a 
natural hybrid of R. malayanum and 136. R. 
jasminijlorum var. punctatum, and exactly inter- 
mediate between these species, which (both ter- 
restrial) are not rare in the Cameron Highlands. 



5. Subsection Albovireya 

Sleum. Rcinwardtia 5 (1960) 107. — Rhododendron sect. Zygomorphante S( hltr, Bot. Jahrb. 
55 (19IK) 145,/7./7. - Fig. 20— 21. 

Scales very dense, in general rather large, touching or slightly overlapping each other, forming 
a coherent layer on the undersurface of (sub)mature leaves, which is persistent for a rather 
long time. Corolla of variou.s shape. 

Distr. Malesia: Sumatra, Java, Celebes, Philippines (Luzon), New Guinea. Fig. 19. 

Note. It is not clear whether the Albovireya group is a natural entity, and deserves as such the rank of 
a subsection, or that it should be merged as a series into subsect. Fuvireya. Presumable hybrids ol 
R. malayanum with species oi subsect. Fuvireya show a similar dark red to copper coloured layer ot very 
dense, partly overlapping scales on the undersurface of the leaves. Possibly such apparently rare species 



538 



Flora Malesiana 



[ser. I, vol. 6* 



as R. lampongum and R. cernuum are of hybrid origin, less probably R. album and R. aequabile. As the 
area of R. malayanum does not extend farther than Ceram and Buru in the East, the New Guinean 
species of subsect. Albovireya cannot be hybrids; they form a rather coherent group with those of Celebes 
within the subsection. 




Fig. 19. Distribution of Rhododendron subsect. Albovireya. 



KEY TO THE SPECIES 

1 . Ovary densely exclusively lepidote. 

2. Corolla ± campanulate. (Leaves 4.5-12.5 by 1.3-5 cm; petiole 0.9-3.5 cm). 
3. Leaves lanceolate to narrow-elliptic; nerves and veins distinctly raised above. Filaments rather 
longish and patently hairy in the lower third 89. R. album 

3. Leaves elliptic; nerves and veins slightly impressed above. Filaments glabrous . . 90. R. aequabile 
2. Corolla ± tubular. 

4. Petiole 1.5-2 cm 91. R. proliferum 

4. Petiole 0-6 (rarely up to 8) mm. 

5. Leaves subcoriaceous to coriaceous (nerves mostly visible on both faces or at least on the upper- 
or undersurface). Style ± as long as or shorter than the ovary. 

6. Leaves ± long cuneate-attenuate to a distinct petiole 92. R. zollingeri 

6. Leaves broadly attenuate or mostly subtruncate or obtuse, sometimes rounded at the base, 
petiole (very) short to 0. 
7. Corolla (1. 3-) 1.5-1. 6 cm, but slightly oblique, lobes equal. Ovary obovoid-obconical. Capsule 

c. 6 mm 93. R. lagunculicarpum 

7. Corolla 1-1.2 (rarely up to 1.3) cm, more distinctly oblique, lobes unequal. Ovary ovoid- 
conical. Capsule 4—5 mm 0. 
8. Perulae lepidote and shortly subsericeous-pubescent dorsally, manifestly ciliate. 

94. R. yelliotii 

8. Perulae caducously lepidote along the dorsal median, or quite glabrous and ± lustrous, 

fimbriate, no proper hairs present at all. More densely lepidote forms of 183. R. inconspiciium 

(subsect. Euvireya ser. Buxifolia). 

5. Leaves ± rigidly coriaceous (nerves ± obscure). Style longer than the ovary. 

9. Leaves elliptic or mostly subovate- or oblong-, rarely ovate-elliptic, (1.5)-1.8-2.7(-3.7) by 

(0.6-)0.8-1.4(-1.6) cm, base ± subtruncate-rounded; petiole distinct, subterete. (2-)3-6(-8) by 

1.5-2 mm 95. R. correoides 

9. Leaves oblong to narrowly obovate-oblong, 1.8-2.4 (rarely up to 3) by (0.4-)0.6-0.8(-l ) cm, 
base cuneate into the very short, much flattened petiole (up to 2 by 1-1.5 mm), or almost sessile. 

96. R. comptum var. comptum 
I. Ovary lepidote and ± densely and patently hairy. 
10. Corolla campanulate or funnelform-campanulate, yellow, lobes — as long as the tube. 
11. Leaves ovate-lanceolate to oblong-elliptic, 2-4(-4.5) cm wide. Corolla funnelform-campanulate, 
1.8-2 cm 97. R. lampongum 

11. Leaves lanceolate, 1-1.5 (rarely up to 1.7) cm wide. Corolla broad-campanulate, 1-1.2 cm. 

98. R. cernuum 
10. Corolla tubular or funnelform-tubular, pink to scarlet, lobes (mostly much) shorter than the tube. 

12. Leaves coriaceous with well visible, either prominent or impressed lateral nerves. 

13. Leaves (4.5-)6-9 by (3-) 3. 5-5 (-5. 5) cm. Corolla 3.3-3.8 cm long in all . . 99. R. pudorinum 



Jan. 1966] 



Ericaceae (Sleumer) 



539 



13. Leaves 1.5-4 by 1-2.6 cm. 
14. Corolla 2.3-2.5(-2.6) cm long in all, laxly to subdensely lepidote outside . . 100. R. arenicolum 

14. Corolla (2.8-)3-3.2 cm long in all, glabrous outside 101. R. giulianettii 

12. Leaves thicker and more rigid, the nerves faint or not visible. 

15. Leaves oblong to obovate-oblong 97. R. comptum var. trichodes 

15. Leaves elliptic or more rarely oblong or subovate-elliptic 102. R, versteegii 



89. Rhododendron album Bl. Cat. Hort. Btzg 

(1823) 72; DC. Prod. 7 (1839) 721; Planch. F1. 
Serres 5 (1849) 477^80 h; Hook. Bot. Mag. 
(1857) t. 4972; Planch. Belg. Hort. 8 (1858) 
70, f. 20; MiQ. Fl. Ind. Bat. 2 (1859) 1058; Ann. 
Mus. Bot. Lugd.-Bat. 1 (1863) 43; Bl. Fl. Jav. PI. 
in^d. (1863-83) t. 3; Koord. Exk. Fl. Java 3 
(1912) 4; J. J. S. in K. & V. Bijdr. 13 (1914) 101; 
Koord. Fl. Tjib. 3 (1918) 3; Steen. Bull. Jard. 
Bot. Btzg III, 17 (1948) 387; Amsh. in Back. Bekn. 
Fl. Java (em. ed.) 7 (1948) fam. 162, p. 4; Sleum. 
Reinwardtia 5 (1960) 108; Steen. Endeavour 
21 (1962) 189, t. 7, non Hoffmannsegg, 1824, nee 
Buch.-Ham. ex D. Don, 1825. — Vireya alba 
(Bl.) Bl. Bijdr. (1826) 855; G. Don, Gen. Syst. 
3 (1834) 848. 

Epiphytic shrub, up to 1 m. Branchlets stoutish, 
densely covered with brown-golden scales at the 
upper 3-4 internodes, the latter 4-8 (rarely up 
to 13) cm long, ultimate ones subangular and 
striate lengthwise in dry specimens, lower ones 
terete. Leaves in 4-5-merous pseudowhorls at the 
upper 3-4 nodes, lanceolate to narrow-elliptic, 
acuminate and subacute at the apex and base, 
coriaceous, rather stiff, sparsely lepidote initially, 
becoming entirely glabrous in mature state and 
slightly shining above, dull brown beneath and 
densely rufous- or rubiginose-lepidote (scales 
rather small, flat, very dense, touching or slightly 
overlapping each other, forming a ± continuous 
layer for a rather long time, the marginal zone 
wide in relation to the small darker centre, ± 
deeply and irregularly crenate to moderately in- 
cised, finally caducous, the centre depressed, re- 
maining pits small, rather close to each other), 
entire, the edge markedly revolute in dry specimens, 
5-12.5 by 1.3-3(-3.5) cm, midrib manifestly im- 
pressed the entire length above, robust and promi- 
nent beneath, nerves 7-10 pairs, irregular, ± 
spreading, rather straight below, anastomosing 
towards the edge, slightly though manifestly 
raised above, obscure beneath; petiole stout, 
grooved above, very densely lepidote, l.2-3(-3.5) 
cm. Umbels 7-12-flowcrcd. Flower buds sub- 
globose, abruptly contracted into a long beak 
(5 mm). Pcrulae ovate-triangular, apex subulate- 
acuminate, lepidote dorsally, up to 1.4 by 0.5 cm. 
Bractcolcs linear, lepidote, up to 1 cm. Pedicels 
rather slender, l-2(-2.3) cm, densely lepidote. 
Calyx disk-shaped, lepidote dorsally, lobes broadly 
ovate, obtuse, apprcssed to the corolla, 1-1.5 mm. 
Corolla campanulatc, cream or pale yellow, rarely 
yellowish pinkish, 1.2-1.5 cm (r. 2 cm when fresh) 
in all, 3-4 mm o at base, c. 1.5 cm at limb, laxly 
goldcn-lcpidotc outside, glabrous inside except for 
the lowest 2 mm which arc laxly pubescent, lobes 
transverse-oval to suborbicular, 7-K by 9-10 mm. 



Stamens 10, c. 1.1 cm; filamentslinear, patent-hairy 
in the lower third, glabrous above; anthers broad- 
oblong, c. 1.8 mm. Disk glabrous. Ovary conical, 
exclusively and densely lepidote, c. 3 by 1.5 mm, 
tapering to the style, the latter rather thick, 6-7 
mm, lepidote in the lower half, glabrous for the 
rest; stigma thick, annular, deeply 5-lobed. 
Capsule fusiform, 2 cm by 4 mm, densely lepidote, 
on fruit-pedicel 2-2.3 cm. Seeds 5-6 mm including 
the tails. 

Distr. Malesia: W. Java (Mts Salak, Gedeh, 
and vicinity), reported from Central Java (Kedu: 
Dieng, leg. Junghuhn), but not recollected 
there. 

Ecol. Mountain forest, 1200-1700 m, not rare 
locally. Fl. Sept. -Jan. 

Vern. Tjantigi koneng, J. 

90. Rhododendron aequabile J. J. S. Bull. Jard. 
Bot. Btzg III, 13 (1935)451; Sleum. Reinwardtia 
5 (I960) 109. — R. album (non Bl.) Ridl. J. Fed. 
Mai. St. Mus. 8 (1917) 58. 

Much branched tree or large shrub, 1.5-4 m. 
Branchlets stout, terete, densely dark rufous- 
lepidote at the upper 2 longitudinally striate inter- 
nodes; internodes 3.5-16 cm. Leaves in 6-9- 
merous pseudowhorls at the ultimate 2 or 3 
nodes, elliptic, apex shortly acuminate or apicu- 
late, subacute to rather obtuse, base 4: long- 
attenuate into the petiole, stiff-coriaceous, gla- 
brous at maturity above, very densely dark 
rufous-lepidote all over the undersurface (scales 
very similar to those of R. album), entire, the 
edge ± revolute in dry specimens, 4.5-10(-ll) 
by (1.7-)2.5-5 cm, midrib depressed over its 
entire length, strongly convex-prominent beneath, 
nerves 6-10 pairs, irregularly spreading, rather 
straight below, anastomosing towards the edge, 
slightly but distinctly depressed above, a little 
or not raised beneath, veins rather indistinct, 
faintly impressed above; petiole grooved on both 
sides, densely lepidote, 0.9-2.8 cm by 1.8-3 mm. 
Umbels 6-12-nowered. Outer perulac ovate- 
acuminate, abruptly subulate in the upper half, 
-!- lepidote outside especially at the acumen, up 
to 2.2 cm by I cm, inner ones lincar-spathulate, 
apex crose, glabrous, up to 1.5 cm by 3 nun. 
Bracteoles filiform, up to I cm. Pedicels thickened 
at apex, curved, densely lepidote. cpilosc. 1-1.5 
cm by I mm. Calyx rather small, oblique, disk- 
shaped, angular, 3-4 mm o, margin indistinctly 
lobed. wavy, densely lepidote dorsally. Corolla 
tubular below, widened distally, i.e. campanulatc 
in outline, brick red to carmine, laxly to sub- 
densely lepidote at the tube and the lower part 
of the lobes outside, glabrous inside, 1.7-2(-2.5) 
cm long in all. tube 1-1.2 cm, 4-5 mm at base, 



540 



Flora Malesiana 



[ser. I, vol. 6* 



8-10(-15) mm at throat; lobes obovate-orbicular, 
slightly retuse, repand or wavy at the apex, 
6-9(-ll) by 5-9(-ll) mm. Stamens 10; filaments 
linear, glabrous, 1-1.3 cm; anthers broad-oblong, 
2-2.2 by 1 mm. Disk glabrous. Ovary conical- 
ovoid, densely lepidote, epilose, c. 4 by 3.5 mm, 
abruptly attenuate to the style, the latter rather 
thick, glabrous, 8-10 mm; stigma very thick, 
broad-conical or subcapitate, 2 mm 0, 5-lobulate. 
Capsule elongate-ellipsoid, n.4-)1.6-2(-2.2) cm 
by 6-7 mm, slightly 5-ribbcd, densely lepidote, on 
fruit-pedicel 1-1.7 cm by 1-1.5 mm. Seeds 4 mm 
including the tails. 

Distr. Malesia: Sumatra (Westcoast to Ben- 
kulen: Mts Singgalang, Kerintji, and Pesagi). 

Ecol. Dense (subalpine) forest, also in Vacci- 
niuni forest, (1220-)2300-2870 m. Fl. Jan. -June, 
//-. May-June. 

91. Rhododendron proliferum Sleum. Blumea 12 
(1963) 101. 

Erect, terrestrial, few-branched shrub, 1.5-2 m. 
Branchlets purplish, obtusangular, tips very 
densely and rather persistently lepidote; innova- 
tions not rarely with numerous basal, long- 
persistent thin perulae, as are found in the flower 
buds, due to a median foliar prolification of the 
inflorescence; internodes (3-)6-10 cm by 4—6 mm. 
Leaves in (8-)10-12-merous pseudowhorls at the 
upper 2 or 3 nodes, erect, a little spreading (upper- 
most whorl ± concealing the inflorescence or 
infructescence), subspathulate-obovate orobovate- 
oblong, apex broadly attenuate to almost rounded, 
base cuneate to and somewhat decurrent along 
the petiole, young ones covered by scales on both 
faces, mature ones coriaceous, firm, glabrous 
above, very densely and subpersistently lepidote 
beneath (scales subequal, partly covering each 
other, brown to reddish brown and shining, 
marginal zone thin, rather wide, irregularly sub- 
stellate-angular, -lobed or -dented, centre small, 
impressed, leaving dark pits when the scale is 
gone), entire, slightly convex in dry specimens, 
(6-)7-10.5 by (2-)2.5-4.5(-5) cm, midrib rather 
narrowly impressed above, as thick as the petiole 
and obtusely prominent beneath, nerves 7-9 pairs, 
lower ones in an acute, upper ones in a wide angle 
from the midrib, obscurely inarching before the 
edge, faintly immersed above, a little prominent 
or obscure beneath, no reticulation; petiole 
dorsally compressed and grooved, very densely 
lepidote, 1.5-2 cm by c. 3 mm. Umbels (4-)6-10- 
flowered (to conclude from the young infructes- 
cences known); perulae numerous, thin, purplish, 
persistent for a long time, outer ones elongate- 
deltoid to ovate-acuminate and lepidote dorsally, 
inner ones elongate-oblong to -spathulate, up to 
4 by 2 cm. Pedicels densely lepidote, 2.5-3.2 cm 
by c. 1 mm in fruit. Calyx but slightly oblique, 
patellar, shortly 5-lobed, c. 5 mm across. Corolla 
(only known in withered state on the young fruit) 
tubular below, expanded at the lobes, apparently 
slightly zygomorphous and red when fresh, c. 
3 cm long in all, lepidote at the base of the tube 
outside, otherwise glabrous, tube c. 2 cm by 4-5 



mm, lobes obovate, c. 5 mm 0. Stamens 10; 
filaments linear (1 mm) below, narrower upwards, 
glabrous, c. 2.7 cm; anthers subobovate-elliptic, 
base obtuse, c. 3.5 by 1.5 mm. Disk prominent, 
glabrous. Ovary in a developed state (or young 
fruit) very densely set with rubiginous stellate 
scales, tapering to the style, which is lepidote at 
the base, glabrous for the rest. 2-2.5 cm; stigma 
subpeltate, 2-2.5 mm 0. Fruit thick-fusiform, 
with 5 thick ribs, densely lepidote, finally 2-2.5 cm 
by 6-8 mm. 

Distr. Malesia: W. New Guinea (Arfak Mts: 
on Mt Gwamongga, Anggi Gigi Lake), once 
collected. 

Ecol. Dense submossy forest, on steep slope 
below summit, 2550 m, apparently rare. Fr. Jan. 

92. Rhododendron zollingeri J. J. S. Ic. Bog. 4 
(1910) 73, t. 322; Koord. Exk. Fl. Java 3 (1912) 6; 
Back. Bull. Jard. Bot. Btzg 11, 12 (1913) 28 
Vzollingerianum"); J. J. S. in K. & V. Bijdr. 13 
(1914) 108; MiLLAis, Rhodod. (1917) 265; J. J. S. 
Bull. Jard. Bot. Btzg III, 4 (1922) 240, incl. var. 
latifolium J. J. S.; Hochr. Candollea 2 (1925)492; 
Malm in Fedde, Rep. 34 (1934) 284; J. J. S. 
Bot. Jahrb. 68 (1937) 200; H. J. Lam, Blumea 5 
(1945) 577; Amsh. in Back. Bekn. Fl. Java (em. ed) 
7 (1948) fam. 162, p. 3; Sleum. Reinwardtia 5 
U960) 109; Steen. Endeavour 21 (1962) 189, 
t. 5. — R. tubiflorum (nan Bl.) Mor. Syst. Verz. 
(1846) 42. — R. album (non Bl.) Zoll. Syst. Verz. 
2 (1 854) 1 37. — R. malayanum {non Jack) Koord. 
Junghuhn Gedenkb. (1910) 185. — Fig. 20. 

Much-branched, erect treelet or shrub, (0.2-) 
1-4 m. Branchlets rather slender, terete, rather 
densely reddish lepidote at the upper 1-2, glabres- 
cent at the lower internodes, the latter (1.5-) 
2.5-6(-8, rarely up to 10) cm, finally greyish 
corticate. Leaves in (4-)5-6-merous pseudowhorls 
or at least close together at the upper 2-3 
nodes, not rarely an odd leaf much lower 
inserted than the others, lanceolate to obovate- 
or elliptic-lanceolate, apex shortly acuminate, 
subacute or sometimes rather obtuse, not rarely 
minutely glandular-apiculate, base ± narrowly 
attenuate into the petiole, :+: coriaceous, initially 
lepidote on both faces, glabrescent or glabrous 
above at maturity (scales leaving small pits), 
densely lepidote beneath (scales rather small, dense 
to very dense, similar to those of R. album, but 
paler and a little smaller, marginal zone finally 
disappearing, remaining pits minute and dense), 
somewhat revolute at the edge, (3-)3.5-5.5(-7) 
by (l-)1.2-1.6(-2.2, very rarely up to 3) cm, 
midrib impressed above, prominent beneath, 
reddish there in fresh specimens, nerves 5-6(-7) 
pairs, lower ones steeply ascending, upper ones 
more spreading, curved, indistinctly anastomosing, 
faintly impressed above, minutely but mostly well 
visibly raised beneath, veins obscure on both 
sides; petiole semiterete, grooved above, lepidote, 
2.5-5 by 1-1.5 mm. Inflorescence terminal (if also 
axillary ones are present, these reduced). Umbels 
(3-)5-6(-8)-flowered. Perulae ovate to obovate, 
obtuse, thin, lepidote dorsally towards the apex. 



Jan. 1966] 



Ericaceae (Sleumer) 



541 




Fig. 20. Rhododendron zollinxcri J. J. S.. y Vi- In debris of solfatara up to the indicated line. Jang 
plateau, Wclirang summit, 2950 m. I-. Java (van Stef.nis 10968). 



542 



Flora Malesiana 



[ser. I, vol. 6* 



glabrous otherwise, up to 8 by 4 mm. Bracteoles 
filiform, glabrous, up to 6 mm. Pedicels slender, 
densely lepidote, 1-1.5 cm, accrescent in fruit up 
to 2.2 cm. Calyx cup-shaped, oblique, c. 3 mm 0, 
irregularly 5-lobed or -dented, 2 lobes narrow- 
triangular, subacute, up to 2 mm, the other ones 
smaller, all densely lepidote. Corolla tubular, 
1.4—1.5 cm long in all, brick red to scarlet, sub- 
densely lepidote outside, margin of the lobes 
excluded, glabrous inside, tube c. 1 cm long, 
3-4 mm at base, slightly dilated towards the 
limb, lobes suborbicular, c. 4 mm 0, straight. 
Stamens 10, nearly as long as the corolla tube; 
filaments linear, glabrous; anthers broadly ob- 
tusely obovate, c. 1 by 1 mm. Disk glabrous. Ovary 
conical, 5-ribbed, densely rufescent-lepidote, 3-4 
by 2-2.5 mm, rather abruptly attenuate to the 
style which is glabrous, only 4-5 mm long and 
bears a subcapitate 5-lobed stigma. Capsule thick- 
fusiform or subcylindric, sometimes slightly curved 
in the upper part, densely lepidote, 1.3-1.5(-1.8) 
by (0.5-)0.6 cm, crowned by the short style. Seeds 
c. 3 mm including the tails. 

Distr. Malesia: Central and E. Java (Kedu, 
Besuki, Madiun), Lesser Sunda Is (Bali, Lombok), 
Central and SW. Celebes, Philippines (Luzon, 
Prov. Benguet: Mt Bandschan, not recollected 
there). 

Ecol. Bare open slopes or ridges, exposed 
summits, volcanic rocks or crater wails, in fern-, 
grass- and scrub-vegetation or open forest, in 
Casuarina forest, on dry places, on sandy, stony, 
or ashy soil, also near solfataras, common locally, 
(1200-) 1800-3090 m. Fl.fr. predominantly in the 
dry season April-Oct. 

Vern. KSmbang geni lanang, mentigi, rang- 
kapelSt, J, tasi, Toradja: Tai, kajukaleleng, lassa- 
-lassa, Makassar: Kondje, kenjeri, Bali. 

93. Rhododendron lagunculicarpum J. J. S. Bot. 

Jahrb. 68 (1937) 200; Sleum. Reinwardtia 5 (1960) 
110. 

Much-branched shrub or treelet, up to 2 m. 
Branchlets slender, erect or erecto-patent, tips 
rather densely covered with both subsessile and 
stalked star-shaped scales, lower parts verruculose 
and rough to the touch by the remaining stalks 
or rib-like and i longitudinally arranged emer- 
gences; internodes 4-11 cm, upper ones bearing 
7-12 leaves which are rather close together 
in the upper third or half. Leaves suborbicular to 
elliptic, rarely subovate elliptic, broadly obtusely 
attenuate to rounded at the apex and base, the 
apex with a small glandular tip, coriaceous, ± 
glabrous above at full maturity, but rather densely 
pitted, not rough to the touch, undersurface all 
over rufous-lepidote (scales small, sessile, very 
dense and touching each other, marginal zone ± 
irregularly incised, centre rather deeply impressed), 
very densely pitted in later glabrescent stages, 
margin slightly or not revolute, finally crenulate 
by the impressed scales, l-2(-2.5) by (0.6-)0.9-1.6 
cm, midrib slightly sunken above, somewhat 
raised beneath, not rarely becoming indistinct 
distally, nerves 4(-6) pairs, irregularly spreading 



and curved-ascending, anastomosing, slightly 
depressed above as are some of the veins, faintly 
raised beneath; petiole semiterete, grooved above, 
lepidote, 1-2 by 1-1.5 mm. Umbels 4-5(-8)- 
flowered. Outer perulae ovate to suborbicular- 
ovate, subacuminate, inneronesspathulate, obtuse, 
thin, lepidote dorsally towards the apex which is 
finely fimbriate or denticulate, not pubescent, 
up to 8 by 5 mm. Bracteoles linear-spathulate, 
glabrous, up to 8 mm. Pedicels densely covered 
with sessile or very shortly stalked scales. 1-1.3 
cm, becoming rather stoutish in fruit and accres- 
cent to 2.2 cm. Calyx ±_ oblique, disk-shaped, 
3-4 mm 0, lepidote dorsally, limb spreading, wavy 
and irregularly minutely dented (sometimes one 
teeth longer than the others). Corolla tubular, 
straight or slightly curved, shortly 5-lobed, 
densely lepidote outside margin of the lobes 
excepted, glabrous inside, (l.3-)1.5-1.6 cm long 
in all, yellow to light salmon, tube 1-1.2 cm long, 
4—5 mm at base, 5-6 mm just below the lobes, 
the latter rather regular, suborbicular, 4-5 mm 0, 
irregularly crenate at the margin, not rarely retuse. 
Stamens 10; filaments narrowly linear, or almost 
filiform, glabrous, 1-1.2 cm; anthers suborbicular, 
c. 1 mm. Disk glabrous. Ovary broadly obovoid- 
obconical, densely lepidote, 3-4 by 2.5-3 mm; 
style abrupt with the ovary, stoutish, glabrous, 
3-4 mm; stigma subcapitate, shortly 5-lobed. 
Capsule obovoid-fusiform, conspicuously 5-ribbed, 
densely rusty-lepidote, 1.3-1.5 by c. 0.6 cm at full 
maturity, style not accrescent. 

Distr. Malesia: SW. Central Celebes (Lati- 
modjong Range: Mt Pokapindjang and its spurs), 
locally not rare in mountain heath, 2800-3000 m. 
Fl.fr. June. 

94. Rhododendron yelliotii Warb. Bot. Jahrb. 16 
(1892) 25 Cyelliottii'); K. Sch. & Laut. F1. 
Schutzgeb. (1901) 487; Schltr, Bot. Jahrb. 55 
(1918) 147, f. 3; Hoogl. Blumea Suppl. 4 (1958) 
233; Sleum. Reinwardtia 5 (1960) 110; Blumea 1 1 
(1961) 117; ibid. 12 (1963) 102. 

Erect, much-branched and mostly compact 
shrub or small tree, l-2.5(-5) m. Branchlets terete, 
rather slender, tips densely covered with shortly 
stalked, stellate, dilutely rusty scales, lower parts 
rather scabrid; internodes not marked. Leaves 
scattered, rather densely set along the branchlets, 
elliptic to ovate-elliptic, rarely partly ovate or 
obovate in the same specimen, apex mostly 
broadly obtusely attenuate, sometimes rounded, 
the terminal gland small but mostly somewhat 
protruding beyond the edge, base rounded or 
truncate-subcordate, coriaceous, greyish dark 
green and shining above, pale brownish silvery 
or not rarely dilutely yellowish beneath, glabrous 
or very laxly lepidote above at full maturity, the 
caducous scales having left but very shallow pits, 
very densely lepidote beneath (scales ~ light 
yellowish brownish, small, ± touching each other, 
forming a coherent layer initially, marginal zone 
lobed or incised to various degree, centre small, 
rather deeply depressed), distinctly minutely 
crenate at the margin by impressed scales, mostly 



Jan. 1966] 



Ericaceae (Sleumer) 



543 



somewhat revolute, l.l-2.5(-3) by 0.6-1.5(-2)cm, 
midrib slightly impressed above, manifestly 
obtusely raised beneath, nerves 4— 6(-8) pairs, 
rather regular, i straight below, curved towards 
the edge and anastomosing distally, very slightly 
impressed above, faintly raised beneath, veins 
obscure; petiole a little flattened, minutely grooved 
above, lepidote, l-2(-3) by c. 1 mm. Umbels 3-5- 
flowered. Outer perulae ovate, subacute, inner 
ones spathulate. obtuse, all lepidote dorsally and 
shortly subsericeous-pubescent, finely though well 
visibly ciliate, up to 8 by 5 mm. Bracteoles 
linear-subspathulate, glabrous or barbate apically 
up to 8 mm. Pedicel slender, 0.8-1.3 cm in anthesis, 
to 1.8 cm in fruit, densely covered with subsessile 
scales. Calyx disk-shaped, ± oblique, 2.5-3.5 
mm o, very densely lepidote outside, irregularly 
obtusely 5-lobed or -dented. Corolla tubular, 
widened towards the limb, ± zygomorphous and 
not rarely somewhat curved, mostly (blood-) red 
or pale red or deep red within, yellow-brown 
externally, sometimes pale yellow-green, densely 
yellowish scaly outside, glabrous inside, 1-1.2 
(rarely up to 1.3) cm in all, tube c. 8 mm long, 
2.5-3 mm o at base, 4—5 mm below the limb, 
lobes suberect, unequal, obliquely suborbicular, 
c. 3 mm 0, often shortly retuse, irregularly crenu- 
late. Stamens 10, — as long as the corolla tube; 
filaments narrow-linear, glabrous; anthers sub- 
globose, c. 1 mm. Disk glabrous. Ovary ovoid- 
conical, c. 5 by 2.5 mm, densely lepidote, broadly 
attenuate into a stoutish glabrous style (3-4 mm); 
stigma capitate, 5-lobulate. Capsule 1.2-1.5 by 
0.4-0.5 cm, crowned by the relatively short style. 
Seeds c. 4 mm including the tails. 

Distr. Malesia: New Guinea (Finisterre Mts, 
Mt Saruwaged region, Mt Wilhelm and vicinity, 
Owen Stanley Range). 

Ecol. Epiphytic in mountain forest, but mostly 
terrestrial in (Podocarpiis-Libocedrus-)foTest fringe 
growth, on steep slopes or exposed rocks or grass- 
land above the forest line, often along streams, 
(1 300-)! 800-3680 m. Fl.fr. Jan. -Dec. 

Vern. Kutengaigai, Chimbu: Waimambuno, 
wtibamp, Minj: Kamang, kitsch, Hagen: Togoba. 

Note. For differences with 183. R.inconspicuum 
(subsect. Euvireya) see under that species. 

95. Rhododendron correoides J. J. S. Med. Rijks- 
hcrb. 25 (1915) 2; Nova Guinea 12 (1917) 500; 
ibid. 12 (1918) t. I95;Sleum. Reinwardtia 5 (I960) 
1 10, — • R. coniferum Wernh. Trans. Linn. Soc. II, 
Bot. 9 (1916) 97. 

.Shrub with erect branches, (0.25-)l-4 m. 
Branchlcts obtusangular, tips densely clothed with 
subsessile or very shortly stalked scales, glabrcs- 
ccnt and early covered with greyish cork below; 
inlcrnodcs (l-)2-8 cm. Leaves c'Mhcr scattered and 
approxintatc in the upper 2/3 of the upper two 
inlcrnodcs, or crowded in more distinct 5-7- 
mcrous pscudowhorls, elliptic or mostly sub- 
obovatc or oblong-, rarely ovatc-clliptic, apex 
broadly obtusely attenuate, sometimes rounded, 
apiculatc by a small terminal gland, base very 
broadly attenuate to subtruncatc-roundcd, firmly 



coriaceous, densely lepidote initially on both 
faces, glabrescent in later stages and hardly pitted 
above, very densely and more persistently lepidote 
beneath (scales similar to those in R. album, but 
slightly smaller and paler, flat, marginal zone 
finally dissolute, centre moderately sunk, leaving 
dense pits), edge thickened and crenulate by 
impressed scales, (l.5-)1.8-2.7(-3.7) by (0.6-) 
0.8-1. 4(-l. 6) cm, midrib slightly depressed above, 
bluntly raised beneath especially in the lower part, 
evanescent distally, or sometimes entirely in- 
conspicuous, nerves 3-4(-6) pairs, curved-ascend- 
ing, faintly impressed or nearly obscure above, 
indistinct beneath; petiole subterete, slightly 
grooved above, lepidote, (2-)3-6(-8) by 1.5(-2) 
mm. Umbels (2-)3-5-flowered. Outer perulae 
ovate-acuminate, subacute, inner ones oblong to 
obovate-spathulate, membranous and lustrous, 
laxly lepidote on the dorsal midline or keel, up to 
2 by 1 cm. Bracteoles linear-subspathulate, 
glabrous, 1-1.5 cm. Pedicels very densely lepidote, 
epilose, l-1.5(-2) cm by c. 1 mm, forming an 
angle with the calyx. Calyx very shortly cup- 
shaped, oblique, densely lepidote dorsally, 2.5-3 
mm 0, ± obscurely lobed. Corolla tubular below, 
dilated towards the limb, (dark) yellow with 
orange base, or orange-red, or reddish brown, 
rarely brownish yellow tinged with green, (1.5-) 
1.8-2.6 (rarely up to 3) cm long in all, subdensely 
or laxly lepidote at the tube and the middle of the 
lobes outside, glabrous otherwise and inside, 
tube (1 -)1.2-1.8(-2.2) cm long, 3-4 mm at the 
subsaccate base, 5-6 mm across the throat, lobes 
broad-elliptic or suborbicular, suberect or ± 
spreading, 5-8(-10, rarely up to II) mm 0. 
Stamens 10, unequal, rfc as long as the corolla 
tube; filaments linear, glabrous; anthers broadly 
obovate-oblong, 2-2.5 by 1.3-2 mm. Disk slightly 
prominent, glabrous. Ovary conical, densely 
lepidote, c. 4 by 2.5-3 mm (at the base), rather 
abrupt with the style initially, less so in later 
stages; style stoutish, lepidote at the very base, 
glabrous otherwise, as long as the corolla or 
finely exserted; stigma thick, subturbinate- 
capitate, 5-lobed. Capsule subcylindric, apex long- 
attenuate, base short-attenuate, c. 1.5 by 0.4 cm, 
on fruit-pedicel up to 3.5 cm. 

Distr. Malesia: New Guinea (Main Range 
from Mt Carstcns, Hubrecht and Orange Mts 
to Mt Wilhclmina). 

Ecol. Subalpine forest, marginal forest shrub- 
beries and in open vegetation, also on ridge crests, 
not rarely on mossy or peaty soil, locally common, 
3100-3800 (possibly up to 4.300) m. I'l. fr. Feb., 
Aug.- Dec. 

96. Rhododendron comptum C. H. Wuicirr, Kcw 
Bull. (1899) 103;Sleum. Reinwardtia 5 (I960) 111. 
— I-ig. 21. 

var. comptum. 

Low shrub. Branchlcts slender, firm, obtus- 
angular and longitudinally striate, densely covered 
with sessile scales initially, glabrescent and smooth 
with age; inlcrnodcs 2-6(8) cm. Leaves alternale, 



544 



Flora Malesiana 



[ser. I, vol. 6^ 



scattered over the whole length or at least the 
upper half of the upper 1-2 internodes, oblong 
to (narrowly) obovate-oblong, coriaceous, stiff, 
apex shortly attenuate, subapiculate, or sometimes 
nearly rounded, base cuneate into the very short 
petiole, densely lepidote on both faces initially, 
glabrescent and ± shining above with age, under- 
surface more persistently lepidote (scales of the 
type found in R. album, very dense, touching each 
other, flat, thin, rather small, rufescent initially, 
becoming silvery with age, centre moderately 




Fig. 21. Rhododendron comptum C. H. Wright 
var. comptum. a. Branchlet with flowers, X 2/3, b. 
undersurface of leaf, x 6, c. ovary and style, x 2, 
d. stamen, x 2 {a — d MacGregor anno 1889). 

impressed), tardily glabrescent, densely pitted then, 
1.8-2.4 (rarely up to 3) by (0.4-)0.6-0.8(-l) cm, 
edge densely crenulate by impressed scales, not 
revolute, midrib a little sunk above, but sUghtly 
and obtusely raised beneath, ± obscure apically, 
nerves obscure; p>etiole much flattened, 0-2 by c. 
1.5 mm, grooved dorsally, lepidote. Umbels 3-4- 
flowered. Outer perulae ovate-oblong, subacumi- 
nate, inner ones oblong to oblong-obovate, lepidote 
at a broad midline dorsally, glabrous for the rest, 
up to 2 by 1 cm. Bracteoles linear, glabrous, c. 
1.5 cm. Pedicels very densely lepidote, epilose, 



1.2-1.8 cm by 1-1.5 mm. Ca/yxshortly cup-shaped, 
oblique, forming an angle with the pedicel, c. 
3.5 mm 0, densely lepidote outside, obsoletely and 
bluntly lobed, but one lobe may be more distinct 
sometimes. Corolla tubular below, campanulate 
distally, 2-2.4 cm long in all, presumably red, 
laxly lepidote outside, glabrous inside, tube 1.2- 
1.5 cm long, c. 4 mm at base, c. 5 mm at the 
mouth, lobes suborbicular, — spreading, 7-9 mm 
0. Stamens 10, — as long as the corolla tube; 
filaments linear, glabrous; anthers obovate to 
suborbicular, c. 1.3 mm 0. Disk g'abrous. Ovary- 
very densely lepidote, conical, 4-5 mm long, c. 3 
mm at base, tapering to or subabrupt with the 
style, the latter r as long as the corolla, stoutish, 
lepidote, in its lower !4-'/3, glabrous for the rest; 
stigma large, capitate, 5-lobed. 

Distr. Malesia: New Guinea (Owen Stanley 
Range: Mts Scratchley and Victoria), 3050-3960 
m. 

var. trichodes Sleum. Reinwardtia 5 (1960) 111. 

Ovary densely lepidote and patently hairy, other- 
wise not different from var. comptum. 

Distr. Malesia: E. New Guinea (Mt Albert 
Edward), twice found on exposed, rocky bluff's in 
open grassland, 3680 m. Fl. May-July. 

97. Rhododendron lampongum MiQ. Fl. Ind. Bat. 
Suppl. 1 (1860) 251, 585 (descr.); Sleum. Rein- 
wardtia 5 (1960) 111. — Azalea lamponga (MlQ.) 
O. Ktze, Rev. Gen. PI. 2 (1891) 387. 

Shrub. Branchlets rather robust, terete, tips 
densely fuscous-lepidote, glabrescent below; inter- 
nodes (l-)3-l 1 cm. Leaves in 5-6-merous pseudo- 
whorls at the upper 2 nodes, ovate-lanceolate to 
oblong-elliptic, apex shortly to subcaudate- 
acuminate, subacute, base attenuate to the petiole, 
coriaceous, stiff", dark green above, brownish 
greenish underneath, practically glabrous above 
at maturity, dark fuscous- or rubiginose-lepidote 
beneath (scales exactly of the same type as in R. 
album, very dense, leaving small pits), (4-)5-9.5 
by 2-4 (-4.5) cm, margin — revolute in dry 
specimens, midrib strongly impressed above, very 
prominent beneath, nerves c. 8 pairs, irregular, ± 
spreading, straight below, anastomosing towards 
the edge, slightly sunk above in quite mature 
leaves, minutely or not raised beneath, veins ± 
perceptible above only; petiole subterete, grooved 
above, densely lepidote, 1-2.5 cm by 1.5-3 mm. 
Umbels 5-6-flowered; flower buds ovate-apiculate 
as in R. album. Outer perulae ovate, abruptly and 
rather longish apiculate or subulate, inner ones 
oblong-spathulate, obtuse, all lepidote dorsally, up 
to 1.6 by 0.6 cm. Bracteoles linear, laxly lepidote, 
up to 1.5 cm. Pedicels stoutish, n densely rufous- 
lepidote and laxly short-hairy, 1-1.5 cm. Calyx 
small, obtusely shortly 5-lobed, densely lepidote, 
c. 4 mm 0. Corolla campanulate from a basal tube, 
1.8-2 cm long in all, yellow, laxly lepidote on the 
tube and near the sinuses to the lobes, base of the 
tube 3-4 mm and somewhat pouched, lobes 
broadly obovate-spathulate, spreading, 7-9 by 
6-8 mm. Stamens 10; filaments linear, patently 



Jan. 1966] 



Ericaceae (Sleumer) 



545 



long-hairy in the lower third, glabrous elsewhere, 
1.4-1.6 cm; anthers broadly oblong, 2 by 1 mm. 
Disk glabrous at the base, densely whitish hairy 
upwards. Ovary obovoid-cylindric, i.e. attenuate 
at the base, c. 4 by 2 mm, densely lepidote and 
patently hairy, gradually narrowed to the style, 
the latter nearly as long as the corolla, slender, 
lepidote and patently hairy in the lower, con- 
spicuously hairy in the middle, glabrous in the 
upper third; stigma subcapitate, indistinctly lobed. 
Submature capsule obovoid-fusiform, long-attenu- 
ate to the base, broadest in the upper third, c. 1.8 
by 0.3 cm, subdensely lepidote and patently hairy, 
pedicel c. 1.5 cm. 

Distr. Malesia: Sumatra (Padang Highlands: 
Mt Gombak; Lampongs: Mt Radja Bassa), in 
low forest, 1370-2330 m. Fl. Nov. 

98. Rhododendron cernuum Sleum. Reinwardtia 5 
(1960) 111. 

Shrub, 1 m. Branchlets divaricate, nodose, tips 
rufescent-lepidote; internodes 1.5-8 cm. Leaves in 
ir 4-merous pseudowhorls at the upper 1 or 2 
nodes, lanceolate, shortly or long acuminate- 
attenuate at both ends, coriaceous, glabrescent 
above with age, the remaining scales flat and not 
impressed, densely and t: persistently lepidote 
beneath (scales similar to those of R. album, 
centre sunk), edge slightly revolute in dry speci- 
mens, (4-)5-9 by 1-1.5 (rarely up to 1.7) cm, 
midrib impressed above, rather thick-prominent 
beneath, nerves 6-8 pairs, straight and ± spread- 
ing, whether or not minutely depressed above, 
obscure beneath; petiole subterete, lepidote, (6-) 
10-15 by c. 1.5 mm. Umbels ± 5-flowered. Outer 
perulae ovate to subrotundate, abruptly subulate- 
attenuate at the apex, densely lepidote dorsally, 
inner ones ovate to subspathulate, obtuse, up to 
8 by 5 mm. Bracteoles filiform, laxly lepidote, up 
to 6 mm. Pedicels slender, nodding, densely lepi- 
dote, 1-1.5 cm. Calyx suboblique, almost cup- 
shaped, c. 2.5 mm o, margin spreading or reflexed, 
irregularly shortly and obtusely 5-lobed. Corolla 
broadly campanulate, yellow, 1-1.2 cm long in all, 
2-3 mm at base, c. 10 mm at limb, subdensely 
lepidote outside margin of the lobes excepted, 
glabrous inside, lobes broad-obovate, 5-6 by 4-5 
mm, shortly rctusc or irregularly crenate distally. 
Stamens 10, nearly as long as the corolla; filaments 
linear, long patently hairy in the lower third, 
glabrous above; anthers 1.7 mm. Disk prominent, 
very shortly hairy at the upper margin, glabrous 
below. Ovary ovoid-conical, densely patently 
hairy and lepidote, c. 2.5 by 1.7 mm; style rather 
abrupt, lepidote at the very base, glabrous other- 
wise, 4-5 mm; stigma rather large, subcapitate, 
S-lobulate. 

Distr. Malesia: S. Sumatra (Bcnkulcn: Mt 
Belirang and Mt Kaba). 

Ecol. On Mt Belirang on dry sulphur-stone 
slope, I5(X) m, rare, //. July-Aug. 

99. Rhododendron pudorinum Sleum. Rcinwardtiu 
5 (I9«)) 112. 

Trcclct, 4 m. Hrunchlcts robust, 5-7 mm 0, 



obtusangular in the new, densely rufous-lepidote 
shoots, subterete, early glabrescent and greyish 
corticate in the older parts; internodes 12 cm long 
or more. Leaves laxly scattered along the upper 
1 or 2 internodes, broad-elliptic, sometimes nearly 
rounded or obovate-elliptic, apex shortly (0.5 cm) 
acuminate or apiculate, not rarely blunt, base 
broadly cuneate to the petiole, or more rarely 
subtruncate-rounded, coriaceous, glabrous and 
impressed-punctulate above at full age in dry 
specimens, very densely copper-coloured or 
rubiginose-lepidote beneath (scales rather small, 
of the type in R. album), entire or obscurely 
crenulate, edge slightly revolute, (4.5-)6-9 by 
(3-)3.5-5(-5.5) cm, midrib thick and somewhat 
prominent at the base, gradually flattened and 
finally grooved distally above, rather flat and but 
slightly raised beneath, nerves c. 8 pairs, rather 
steeply ascending, slightly prominent (but a little 
depressed in the middle) above, slightly raised 
beneath, veins laxly reticulate, nearly obscure 
above, more visible beneath; petiole flattened and 
grooved dorsally, initially lepidote, (0.8-)1.5(-2) 
cm by 2-3(-4) mm. Umbels 8-1 5-flowered; 
terminal flower buds (i.e. the normal ones) broad- 
ovoid, c. 2 by 1 .5 cm, lateral ones in the axils of the 
upper leaves much reduced in size. Outer perulae 
suborbicular, inner ones broadly spathulate, 
obtuse, sericeous on both sides margin excepted, 
moreover lepidote dorsally, up to 1.3 cm. Bracte- 
oles linear-spathulate, c. 1 cm by 2 mm, densely 
long-hairy. Pedicels densely lepidote, epilose, 2- 
2.5 cm by c. 0.8 mm. Calyx disk-shaped, thick- 
membranous, c. 4 mm 0, wavy and indistinctly 
lobulate at the margin, densely lepidote dorsally, 
almost fimbriate by the scales which project 
beyond the margin. Corolla elongate-funnelform, 
pink, 3.3-3.8 cm long in all, laxly to very laxly 
lepidote outside at the tube and the middle of the 
lobes, glabrous inside, tube 2.2-2.5 cm, straight, 
c. 4 mm at base, 6-7 mm at the mouth, lobes 
broadly obovate to nearly rounded, not rarely 
slightly retuse, 1.1-1.4 by 1-1.2 cm. Stamens 10, 
somewhat longer than the corolla tube, unequal; 
filaments linear below, filiform distally, quite 
glabrous; anthers broadly subobovate-oblong, 
2.5 mm. Disk thick, blackish when dry, glabrous. 
Ovary obconical, c. 5 mm long, 3 mm in the 
upper part, slightly less below, very densely clothed 
with subpatent, short yellowish hairs and scales 
(the latter covered by the hairs), shortly contracted 
into the style, the latter hairy and lepidote at the 
very base, glabrous otherwise, stoutish, c. 2 cm 
long at anthesis; stigma shortly turbinate-capitate, 
but faintly lobed. Capsule subcylindric, apex 
acuminate, base somewhat attenuate, slightly 
thicker in the upper third, densely hairy and 
lepidote, 2.5-3 cm by c. 6 mm; style c. 2.5 cm; 
fruit-pcdiccI 2.5^ cm, glabrescent. 

Distr. Malesia: SW. Central Celebes (Lati- 
modjong Range: Mt Pokapindjang and its spur to 
Tin^bang). 

licol. In low forest, 2500-3000 m. /"/. June. 



546 



Flora Malesiana 



[ser. I, vol. 6* 



100. Rhododendron arenicolum Sleum. Rein- 
wardtia 5 (1960) 113. 

Shrub. Branchlets slender, terete, tips more 
densely lepidote than lower parts; internodes 
3-8 cm. Leaves scattered in the upper half of the 
upper 1-2 internodes, elliptic, rarely subovate- 
elliptic, apex very shortly acuminate or nearly 
apiculate, with a blunt terminal gland, base sub- 
truncate to rounded, coriaceous, glabrous and 
somewhat rugose above with full age, very densely 
reddish brown-lepidote beneath at least initially 
(scales similar to those in R. album and forming 
the same coherent layer, centre but slightly im- 
pressed), finally glabrescent beneath too, densely 
pitted then, margin not or but very slightly revo- 
lute, minutely crenulate, (2-)2.5-4 by 1.5-2.6 cm, 
midrib flat above, slightly prominent beneath, 
nerves 5-7 pairs, lower 2-3 pairs from or from 
somewhat above the base, the other higher from 
the midrib, all straight below, curved distally and 
obscurely anastomosing, faintly sunk above, a 
little raised beneath, veins subobscure; petiole 
lepidote, 3-4 by c. 1.5 mm, flattened dorsally. 
Umbels 7-9-flowered; flower buds broadly ovoid, 
c. 1.5 by 1 cm. Outer perulae ovate, inner ones 
subspathulate, blunt, whitish sericeous and lepi- 
dote along the dorsal midline, up to 1.5 by 1 cm. 
Bracteoles filiform below, up to 2 mm sub- 
spathulate-dilated distally, pubescent, up to 1 cm. 
Pedicels densely rusty-lepidote, epilose, 1.5-2.7 
cm by 0.5 mm. Calyx disk-shaped, c. 3.5 mm 0, 
densely lepidote dorsally, margin wavy and patent, 
indistinctly bluntly 5-lobed. Corolla tubular below, 
dilated towards the limb, pink, 2.3-2.5(-2.6) cm 
long in all, laxly to subdensely lepidote outside 
at the tube and the middle of the lobes, tube 1.2- 
1.5 cm, a little pouched and 3-4 mm at base, 
6-7 mm below the lobes, the latter broadly obovate 
or nearly rounded, retuse or irregularly crenulate, 
spreading, 8-11 mm. Stamens 10, slightly exceed- 
ing the corolla tube in length; filaments filiform, 
glabrous or with some sparse hairs; anthers 
oblong, 1.2 by 0.8 mm. Disk glabrous. Ovary 
thick-conical, 4-5 by 2-2.5 mm, very densely 
lepidote and densely shortly patently hairy; style 
abrupt with the ovary, rather thick, glabrous, 5-6 
mm; stigma large, almost turbinate, manifestly 
5-lobed. 

Distr. Malesia: SW. Central Celebes (Latimod- 
jong Range). 

Ecol. On open sandy ridges, 2600-3000 m. 
Fl. June. 

101. Rhododendron giulianettii Laut. in K. Sch. & 
Laut. Nachtr. (1905) 338; Sleum. Reinwardtia 5 
(1960) 114. — R. papuanum {nort Becc.) C. H. 
Wright, Kew Bull. (1899) 104. 

Small shrub. Branchlets rather slender, ± 
densely covered with shortly stalked scales in the 
younger, scabrous verruculose in the lower parts. 
Leaves scattered, mostly at the upper third of the 
upper internodes (2-8 cm), elliptic to ovate- 
elliptic, sometimes nearly rounded, apex broadly 
attenuate, obtuse to rounded, not apiculate, base 
± rounded, abruptly contracted into the petiole. 



which is somewhat winged by the decurrent blade in 
its upper half, coriaceous, densely lepidote on both 
faces initially, early glabrescent above, the rather 
flat scales leaving but small and shallow pits on 
the upper surface, very densely rusty-lepidote 
beneath (scales rather small, of the type as in R. 
album, but some of the scales not rarely darker in 
colour, the undersurface thus becoming ±spotted), 
margin slightly revolute, (1 .5-)l. 7-2.8 by 1-1.8 
cm, midrib a little impressed above, faintly raised 
beneath, nerves 4-5 pairs, lower ones steeply 
curved upwards, upper ones more spreading, all 
rather irregular, indistinctly anastomosing, slightly 
depressed above, obscure below; petiole flattened, 
grooved above, lepidote, c. 3 by 1-2 mm. Umbels 
± 4-flowered. Perulae ovate, obtuse, outer ones 
lepidote dorsally, inner ones larger, glabrous, not 
hairy or sericeous at all, up to 1 by 0.7 cm. 
Bracteoles filiform, glabrous, up to 1 cm. Pedicels 
stoutish, densely rufous-squamular-lepidote,l-1.4 
cm. Calyx ± obliquely disk-shaped, 3-4 mm 0, 
margin wavy and obscurely dented, densely 
lepidote dorsally. Corolla tubular below, dilated at 
the lobes, apparently red, completely glabrous in- 
and outside, (2.8-)3-3.2 cm long in all, tube 
cylindric, straight, slightly dilated upwards, (1.8-) 
2-2.2 cm, 4—5 mm at base, 6-7 mm at the mouth, 
lobes nearly rounded, ± spreading, c. 1 cm 0, 
mostly somewhat retuse. Stamens 10, unequal, 
2.5-3 cm; filaments linear, dilated below (1 mm), 
nearly filiform distally, glabrous; anthers 2.6-3 by 
1 mm. Disk conspicuous, glabrous. Ovary conical, 
5 by 2.5 mm, densely shortly patently hairy and 
lepidote, rather gradually attenuate to the style, 
the latter stoutish, glabrous the lepidote base 
excepted, c. lA cm; stigma broad-conical, lobes 
inconspicuous. 

Distr. Malesia: E. New Guinea (Owen Stanley 
Range: Mt Scratchley), 3050-3960 m. Once found. 

102. Rhododendron versteegii J. J. S. Med. Rijks- 

herb. 25 (1915) 2; Nova Guinea 12 (1917) 501; 
ibid. 12 (1918) t. 196; Sleum. Reinwardtia 5 (1960) 
114;Blumeall (1961) 117. 

Erect shrub, 0.2-0.35 (sometimes up to 1) m, 
sparsely branched. Branchlets stoutish, terete, 
densely covered with sessile scales, early glabres- 
cent and smooth then; internodes 2-8(-12) cm. 
Leaves scattered, ± dense but distant from each 
other in the upper Vi-Vi of the upper internode, 
obovate or elliptic-obovate, elliptic or more rarely 
oblong- or subovate-elliptic, apex broadly attenu- 
ate and bluntish or rounded, base subtruncate- 
rounded, or broadly attenuate, thick-coriaceous, 
flat or but slightly convex, very densely lepidote on 
both faces initially, tardily glabrescent above, 
lepidote for a longer time beneath (scales small, 
very dense, of the type of R. album, first rusty, 
then silvery, marginal zone finally dissoluted, 
centre leaving shallow pits), marginate, mani- 
festly densely crenulate by impressed scales, 
(l-)1.5-3(-4) by (0.6-)0.8-1.4(-2) cm, midrib 
narrowly grooved above, thick and obtusely 
prominent in the lower, gradually obscure towards 
the upper part of the lamina beneath, nerves 3-4 



Jan. 1966] 



Ericaceae (Sleumer) 



547 



pairs, faintly impressed above, or mostly i 
inconspicuous on both faces; petiole — flattened, 
grooved above, lepidote, 2-3 (-4) by 1-2 mm. 
Umbels (2-)3-5-flowered. Outer perulae ovate- 
oblong, subacuminate or obtuse, inner ones oblong 
to oblong-spathulate, membranous, lepidote on a 
broad dorsal midline, glabrous otherwise, up to 
1.9 by 0.8 cm. Bracteoles linear to linear-sub- 
spathulatfc, glabrous, up to 1.6 cm. Pedicels erect, 
forming an angle with the calyx (the flower 
nodding), very densely rusty-lepidote, epilose, 
0.9-2 cm by 0.8 mm. Calyx very oblique, disk- 
shaped, densely lepidote, 4-5 mm 0, margin wavy, 
obtusely 5-lobed, one or the other lobe some- 
times elongate. Corolla tubular below, widened 
towards the limb, (2.5-)3-3.5 cm long in all, 
laxly to subdensely lepidote outside at the tube 
and the dorsal midline of the lobes, glabrous 
inside, tube (1.8-)2-2.2 cm, somewhat angular, 
red, 4-5 mm o at the slightly pouched base, 6-8 
mm at the mouth, lobes red, yellow, or partly 
yellow and partly red (tinged red outside), obovate 



to suborbicular, somewhat spreading, (1-) 1.2-1. 4 
by 1-1.3 cm. Stamens 10, unequal, the longest 
nearly as long as the corolla; filaments linear, 
glabrous; anthers obovate-oblong, c. 2.5 by 1 mm. 
Disk prominent, glabrous. Ovary shortly conical, 
very densely lepidote and hairy (the spreading 
hairs covering ± the scales), 5-6 by 2.5-3 mm, 
rather abrupt with the style initially, less so in 
later stages; style slender, ± as long as the corolla, 
densely lepidote and laxly to sparsely hairy in the 
lower ^/s to ^/g, glabrous for the rest; stigma 
capitate, distinctly 5-lobed. Capsule oblong- 
conical, 5-ribbed, densely lepidote and hairy, 
(1. 3-) 1.5-2 by 0.5-0.6 cm, broadest below the 
middle, pedicel 2-3 cm by 1-1.5 mm. 

Distr. Malesia: W. New Guinea (Quarks 
Valley to Mt Wilhelmina and Habbema Lake). 

Ecol. Grassy edges oi Podocarpus forest, alpine 
grassland or boggy slopes, scattered through open 
mossy thickets of exposed ridges, 3200-3950 m, 
sporadic, but locally common, in wet shallow 
sandy or peaty soil. Fl. Feb., Sept. 



6. Subsection Solenovireya 

CoPEL./. Philip. J. Sc. 40 (1929) 136, 158; Sleum. Bot. Jahrb. 74 (1949) 5^1, p.p.; Reinwardtia 
5 (1960) 114. — FJwdodendron sect. Hadranthe Schltr, Bot. Jahrb. 55 (1918) 157, p.p. — 
Fig. 23—25. 

Scales star-shaped to various degree, mostly flat, rarely subdendroid, rarely each on top of a 
low, hardly permanent epidermal tubercle, ± equal in size, distant from each other. Corolla 
tubular, salver-shaped, tube straight or slightly curved, long and narrow, lobes relatively short 
(at most 1/4 as long as the total corolla), spreading ± horizontally. 

Distr. Malesia (excluding Java and Lesser Sunda Is). Fig. 22. 




Fig. 22. Distribution of Rhododendron subsect. Solenovireya. 



KEY TO THE SPECIES 

I . Ovary densely and exclusively (or at least seemingly so) lepidote. 
2. Style lepidote or glabrous. 
3. Style glabrous. Leaves 3.5-6 cm wide. 

4. Leaves h broadly elliptic or obovate-, rarely oblong-elliptic; nerves I0-I2(-I4) pairs, rather close 

to each other. I'crulac lepidote in the upper part dorsally, not hairy at all . . 103. R. miiltincTvium 

4. Leaves oblong-elliptic; nerves 6-8 ±, distant pairs. I'crulac densely sericeous on both faces, 

otherwise very laxly or not lepidote 104. R. natuliciuin 

3. Style lepidote to the top. Leaves 1.8-2.7(-3) cm wide 105. R. rultenii 



548 Flora Malesiana [ser. I, vol. 6* 

2. Style laxly hairy 106. R. rhodosalpinx 

1. Ovary manifestly (mostly densely) set with shorter or longer hairs, or ovary both hairy a//i/iepidote. 
5. Leaves ± persistently soft-pubescent beneath, or very tardily glabrescent . . . 107. R. stapfianum 
5. Leaves initially with some scattered hairs at the base and early glabrescent, or mostly not hairy at all. 

6. Leaves linear, 3-5 cm by 3-5 mm 108. R. cbamaepitys 

6. Leaves of other shape, much wider. 
7. Anthers (5-8 mm) with a distinct, elongate apiculus or caudate appendix (0.5-1.5 mm) at the 
base of each cell. 
8. Anthers 7-8 mm. Corolla tube slightly gradually dilated towards the lobes, the latter 2.5-3 cm 0. 

109. R. macrosiphon 
8. Anthers 5-6(-7) mm. Corolla tube rather equally wide from the base to the lobes, the latter 
1-1.5 cm 0. 
9. Corolla subdensely lepidote outside. Leaves ± densely reticulate on both faces, veinlets ± 

distinct 110. R. carringtoniae 

9. Corolla glabrous outside. Leaves much less or hardly reticulate, veinlets obscure. 

10. Corolla c. 7 cm long in all 111. R. carstensense 

10. Corolla (8.5-)9-10(-10.5) cmlongin all 112. R. syringoideum 

7. Anthers (2-6 mm) provided with a very short obtuse knob-like apiculus (0.3 mm) or truncate- 
obtuse at the base. 
11. Leaves manifestly, i.e. ± densely and prominently or depressedly reticulate on both faces or 
at least underneath (well visible both in younger and older leaves). 
12. Corolla tube (7-)8-8.8 cm. Leaves cuneate to obtuse at the base, manifestly petioled. 

113. R. maius 
12. Corolla tube generally up to 6.5 cm (if attaining 7 cm, leaves cordate at the base, subsessile 
or very shortly petioled). 
13. Leaves ± acutely attenuate or cuneate at the base. Style hairy at the base, glabrous for the rest. 

114. R. cruttwellii 
13. Leaves broadly attenuate, obtuse or truncate-rounded or (sub)cordate at the base. 
14. Style hairy and lepidote nearly to the top. Leaves ovate or elliptic-ovate, apex acuminate. 

Umbels 8-13 (-15)-flowered 115. R. archboldianum 

14. Style hairy and lepidote in the lower 2/3 or less. 
15. Umbels (8-) 1 5-20-flowered. Style hairy in the lower ^/a-V*- Leaves elliptic or obovate- 

elliptic, apex ± obtuse 116. R. pleianthum 

15. Umbels up to 7-flowered. 
16. Flowers solitary or in twos. Style hairy to the middle. Leaves subovate-elliptic, subsessile. 

117. R. oliganthum 
16. Umbels 3-7-flowered. 
17. Style hairy but for c. 5 mm above the base. Leaves ovate or subovate-rounded, sessile or 

nearly so 118. R. carrii 

17. Style hairy up to the middle, or sometimes even to the lower 2/3. Leaves broad-, oblong- 
or subobovate-elliptic. 

18. Leaves 7-9(-10) by 4-5(-6) cm. Petiole (6-)8-12 mm 119. R. armitii 

18. Leaves 4-7 by 2.5-3. 5(-5, rarely up to 6) cm. 
19. Leaf base subtruncate-rounded, or sometimes subcordate; petiole 2-4 mm. Corolla 
tube very slightly or not widened upwards. Anthers c. 2.5 mm .... 120. R. tuba 
19. Leaf base distinctly (rather deeply) cordate; petiole 0-1 mm. Corolla tube distinctly 

gradually dilated upwards. Anthers 3.5-4 mm 121. R. rhodoleucum 

11. Leaves but indistinctly and laxly (prominently or impressedly) reticulate when young, less 

so when fully mature, or without a trace of reticulation. 
20. Corolla tube laxly to densely short-hairy, otherwise whether or not lepidote outside. 

21. Corolla tube (5-)5.5-6.5 cm 122. R. pubitubum 

21. Corolla tube 3^.5 cm. 

22. Leaves 8-1 2 by 3-5 cm 123. R. pseudotrichanthum 

22. Leaves 2.5-6 (rarely up to 7.5) by (1-) 1.5-3 cm. 

23. Corolla lobes ciliate. Disk glabrous 124. R. amabile 

23. Corolla lobes not ciliate. Disk pubescent 136. R. jasminiflorum 

20. Corolla epilose outside (except some scattered hairs which may occur at the very base), 
otherwise whether or not lepidote. 
24. Corolla tube at least 4.5 cm, mostly longer. 
25. Leaves sessile or nearly so (petiole 0-2, rarely up to 4 mm), in general ± obtuse, rounded 
or subcordate at the base. 
26. Anthers c. 1.8 mm 125. R. radians 

26. Anthers 4.5-5 mm 116. R. pleianthum 

25. Leaves distinctly petioled, in general attenuate to cuneate at the base. 

27. Leaves acutely attenuate at both ends. Pedicels stoutish, c. 2 mm . . 126. R. oreadum 



Jan. 1966] 



Ericaceae (Sleumer) 



549 



. . 127. R. cinerascens 
128. R. brachypodarium 



27. Leaves obtusely acuminate (or attenuate) to rounded at apex. 
28. Pedicels rather slender, c. 0.5 mm 0. 
29. Pedicels 9-20 nun. Umbels 5-6(-7)-flowered 

29. Pedicels4-8 mm. Umbels (8-)10-12(-14)-flowered .... 
28. Pedicels stoutish, 1-2 mm o, 3-7 mm long. 

30. Corolla 5.7-7 cm long in all 129. R. goodenoughii 

30. Corolla (8.5-)9-10(-10.5) cmlongin all 112. R. syringoideum 

24. Corolla tube 2-4 (rarely in part up to 4.5) cm. 
3 1 . Corolla tube — manifestly and gradually narrowed from the base upwards. Leaves subsessile. 
32. Pedicels all over densely lepidote and laxly short-hairy. Corolla i densely lepidote outside 
at least initially. Leaves elliptic-oblong or -obovate 130. R. pneumonanthum 

32. Pedicels lepidote in the distal part only, otherwise densely hairy. Corolla laxly lepidote or 
glabrous outside from the beginning. Leaves broadly elliptic, subovate- to suborbicular- 
elliptic, or orbicular 131. R. orbiculatum 

3 1 . Corolla tube equally wide all over or slightly and gradually widened from the base upwards. 
Leaves distinctly petioled, but short-petioled to subsessile in R. jasminiflorum p.p. and 
R. orbiculatum. 

33. Corolla tube subdensely to densely lepidote outside. 

34. Scales on the undersurface of the leaves subdendroid, i.e. the centre i deepened into a 
kind of stalk, which itself rests on a low epidermal tubercle. Corolla tube 2-2.5 cm, its 
scales stellate 132. R. edanoi 

34. Scales on the undersurface of the leaves sessile in minute epidermal depressions. Corolla 
tube 2-2.5 cm, its scales thick and roundish 133. R. loranthiflorum 

33. Corolla tube laxly lepidote to glabrous outside. 

35. Corolla tube 2-2.3 cm 134. R. retrorsipilum 

35. Corolla tube (3-)3.5-^(-4.3) cm. 

36. Pedicels very slender, practically glabrous. Style laxly hairy and lepidote in the lower 

half, otherwise exclusively lepidote to the top 135. R. subpacificum 

36. Pedicels rather stoutish, lepidote and/or hairy to various degree. 
37. Style laxly lepidote and hairy in the lower third, otherwise exclusively hairy to almost 

the top. Leaves petioled (r. 8 mm) 136. R. jasminiflorum 

37. Style laxly hairy except the glabrous uppermost part, sparsely or not lepidote. Leaves 
subsessile 131. R. orbiculatum 



103. Rhododendron multinervium Sleum. Rein- 
wardtia 5 (1960) 117; Blumea 11 (1961) 111; ibid. 
12 (1963) 102. — Fig. 23. 

Shrub or small tree, 1.5-3 m, terrestrial or 
epiphytic. Branchlets divaricate, tips rather densely 
lepidote initially; internodes 4—6 cm. Leaves 
arranged in 3-merous pseudowhorls, obovate- 
clliptic or — broadly, rarely oblong-elliptic, apex 
very shortly and ± abruptly obtusely acuminate, 
base broadly attenuate into the petiole, lepidote 
initially, glabresccnt with age above, rather densely 
and — persistently lepidote beneath (scales small, 
marginal zone substellate-dentale, centre becoming 
blackish), entire, slightly rcvolute towards the base 
of the blade, (5-)6-8 by 3.5-6 cm, midrib very 
slightly impressed above, robust and strongly 
prominent beneath, nerves I0-12(-I4) pairs 
rather close and ; parallel to each other, • 
straight below, obscurely anastomosing before 
the edge, with some other less distinct nerves 
between them, all a little prtjmincnt on both 
faces, reticulation lax and faintly raised under- 
neath only; petiole grooved above, somewhat 
flattened, lepidote, 6-8 by c. 1.5 mm. Umbels 
t5-flowcrcd. Outer pcrulae broadly ovale, inner 
ones spathulatc, all : densely lepidote in the 
upper part outside, not hairy at all, lepidotc- 
ciliatc, 1-2 by 0.8-1.4 cm. Bractcolcs fililorm. 
Pcdicclj rather slender, densely lepidote, 6-8 mm. 
Calyx oblique, disk-shaped, very shortly lobcd and 



wavy, c. 2.5 mm 0. Coro//a salver-shaped, white, 
sweetly scented, tube straight, glabrous or laxly 
lepidote outside, scales more dense at the sinuses 
of the lobes, subdensely hairy in the lower half, 
glabrescent upwards inside, 3.5-4 cm long, at the 
base c. 2, in the middle 2.5-3 mm 0, very slightly 
or not dilated at the mouth, lobes spreading, 
obovate-elliptic, 8-12 by 6-8 mm. Stamens 10, 
slightly exserted from the throat; filaments linear 
below and rather densely patent-hairy to the 
lower 2/3, filiform and glabrous above, unequal; 
anthers 2.5 by 0.9 mm. Disk rather densely and 
shortly white-hairy. Ovary elongate columnar, 
shallowly 5-ribbed, very densely covered with 
thickish, almost round scales, 5-6 by 1.5 mm, ± 
gradually tapering to the slender and glabrous 
style (3 cm); stigma short-obconical, crcnulate. 
Capsule elongate-fusiform, subdensely lepidote, 
3-3.5 by 0.4-0.5 cm. 

Distr. Malesia: New Guinea (Western High- 
lands; Sepik Distr.). 

Ecol. On grassy slopes or in Castaiiopsis- or 
Nofhofagus-forcst, 1370-2050 m. //. April, Aug.- 
Ocl. 

Vern. Aretna, Anona. 

104. Kh<»dodindron nutalicium Sleum. Reinwardtia 

5 (I9f)()) I IS. 

Shrub, possibly epiphytic. Branchlets flattened, 
tips subdensely suhstcllate-lcpidole; internodes 



550 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 23. Rhododendron multinervium Sleum. a. Branch with flowers, x yj, b. ovary and style, X 
c. stamen x */3 (a-c Hoogland & Pullen 5863). 



7-15 cm. Leaves in ± 4-merous pseudowhorls, 
oblong- or lanceolate-elliptic, apex shortly grad- 
ually acuminate, subacute, base broadly attenu- 
ate, subcoriaceous or coriaceous, entire, sub- 
revolute, entirely glabrescent above at maturity, 
laxly lepidote beneath (scales very small, fuscous, 
irregularly incised in the marginal zone, the centre 
slightly impressed), 7-10 by 3.5-4.5(-5) cm, 
midrib slightly but distinctly impressed above, 
much prominent beneath to 3^^ of its lower part, 
less so distally, more densely lepidote than the 
intervenium beneath, nerves 6-8 pairs, rather 
distant from each other, at an acute angle from 
the midrib, ± straight below, rather obscurely 
curved-anastomosing near the edge, somewhat 
raised above (but impressed in the very middle), ± 
prominent beneath, reticulation obscure; petiole 
subdensely lepidote, grooved above, 1-1.5 cm 
by 1-1.5 mm. Umbels ± 8-flowered. Outer perulae 
ovate, inner ones spathulate, all subsericeous on 
both faces, otherwise not or but very laxly lepidote, 
long-ciliate, up to 2.5 cm by 6-8 mm. Bracteoles 
nearly filiform and glabrous below, narrow- 
spathulate and hairy distally, up to 2 cm. Pedicels 
stoutish, rather densely substeliate-lepidote, 10-12 
mm. Calyx very oblique, irregularly shortly and 
obtusely 5-lobed, lepidote, c. 2.5 mm 0. Corolla 
salver-shaped, snow-white, tube straight or slightly 
curved, c. 4.5 cm by 3-4 mm, faintly or not dilated 



towards the mouth, subdensely substeliate-lepidote 
outside, rather densely, shortly and patently 
whitish hairy in the lower third, gradually glabres- 
cent distally" inside, lobes eliiptic-obovate to 
obovate-spathulate, spreading, laxly lepidote out- 
side, glabrous inside, 1-1.2 by 0.8-1 cm. Stamens 
10, unequal, exserted, c. 5.5 cm; filaments linear 
and densely patently hairy below, less hairy and 
finally glabrous to the top; anthers oblong, obtuse 
base, c. 3 by 0.8 mm. Z)wA: subdensely short-hairy. 
Ovary elongate-conical, densely set with almost 
roundish scales, c. 8 by 1.5 mm, tapering to the 
style, which is lepidote at the very base, glabrous 
otherwise, and ± equals the corolla tube in length; 
stigma narrowly short-obconical. Capsule elon- 
gate-cylindric, curved, shortly attenuate at both 
ends, laxly lepidote, 4-4.5 by 0.4 cm. 

Distr. Malesia: New Guinea (Sattelberg 
region, Morobe Distr.: Ogeramnang; Mt Sam- 
waged), in forest hills at 1770-2440 m. Fl. March, 
fr. May. 

105. Rhododendron ruttenii J. J. S. in Fedde, Rep. 
30 (1932) 170; Sleum. Reinwardtia 5 (1960) 119. 
Shrub. Branchlets terete, tips only sparsely 
lepidote; internodes 1.7-4.5 cm. Leaves 4-5 per 
pseudowhorl, cuneate-obovate, rarely oblong- or 
eliiptic-obovate, apex broadly attenuate, obtuse to 
rounded, sometimes slightly retuse, base cuneate- 



Jan. 1966] 



Ericaceae (Sleumer) 



551 



attenuate, mostly rather acute, coriaceous, laxly 
lepidote initially, glabrescent above, more densely 
and persistently lepidote beneath (scales very 
small, the marginal zone narrow and ± coarsely 
dentate, each scale on top of a minute epidermal 
elevation), margin somewhat revolute in dry 
specimens, (3-)4-5.5(-6.5) by 1.8-2.7(-3) cm, 
midrib impressed above, manifestly raised beneath, 
nerves 6-8 pairs, rather straight below, curved and 
anastomosing before the edge, slightly impressed 
above, rather inconspicuously raised beneath, 
reticulation obscure; petiole semiterete, grooved 
above, lepidote, 3-7 by c. 1.5 mm. Umbels 6-12 
(-14)-flowered. Well developed perulae and 
bracteoles not known. Pedicels rather slender, 
± densely lepidote, 1-1.8 cm. Calyx small, 
obliquely disk-shaped, lobes obsolete. Corolla 
salver-shaped, white to light red, tube cylindric, 
straight, 3-3.5 cm by c. 2 mm, laxly lepidote out- 
side, laxly hairy in the lower part inside, lobes 
spreading, subrotundate-obovate, laxly lepidote 
outside, glabrous inside, 1-1.3 by 0.6-0.8 cm. 
Stamens 10, slightly exserted; filaments nearly 
filiform, laxly hairy above the base only; anthers 
2 by 0.9 mm. Disk sparsely hairy in the upper half. 
Ovary elongate-cylindric, c. 5 by 2 mm, gradually 
attenuate into the slender style (c. 3 cm), the 
latter rather densely lepidote up to almost the top; 
stigma shortly broad-turbinate, subcrenulate. Cap- 
sule lepidote, c. 2 cm by 3 mm. Seeds c. 2 cm by 
0.03 nrni. 

Distr. Malesia: Moluccas (Central Ceram: Mt 
Murkele), twice collected. 

Ecol. Primary forest, abundant on limestone, 
1900-2500 m. Fl. July. 

106. Rhododendron rhodosalpinx Sleum. Blumea 1 1 
(1961) 121. 

Shrub. Branchlets rather slender, c. 2 rrun 0, tips 
densely lepidote; internodes l-2(-4) cm. Leaves in 
(3-)4-merous pseudowhorls at the 2(-3) upper 
nodes, elliptic or subobovate-elliptic, apex broadly 
attenuate and blunt, or almost rounded, some- 
times faintly retuse, base broadly attenuate to the 
petiole or subcuneate, coriaceous, firm, glabrous 
with age above, subdensely lepidote beneath 
(scales small, marginal zone substellate-dented, 
minute centre rather deeply impressed), distinctly 
revolute at the edge, (3-)4-5 by (1.5-)1. 8-2.6 cm, 
midrib very narrowly impressed above, as thick as 
the petiole and much prominent below, gradually 
decreasing distally beneath, nerves c. 3 pairs, a 
little raised beneath only, or not rarely inconspicu- 
ous, no reticulation; petiole slightly flattened, 
grooved above, lepidote, 4-6 by 1 mm. Umbels 
3-flowcrcd. Pedicels subdensely lepidote, minutely 
(lens!) patcntly-pubcrulous, 1.7-2.1 cm by c. 0.8 
mm. Bracteoles linear, c. \ cm. Ca/vjc subobliquely 
disk-shaped, very shortly and irregularly 5-lobcd, 
c. 2.5 mm 0, lepidote dorsally. Corolla salvcr- 
shapcd, apparently red, subdensely substcllatc- 
lepidotc at the lube and the lower part of the lobes 
outside, laxly hairy at the tube below inside, other- 
wise glabrous, tube straight, 3-3.5 cm long, 
(3-)4 mm at base, 5-6 (-7> mm at the base of the 



lobes, the latter ± spreading, obovate, 1-1.2 by 
c. 0.8 cm. Stamens 10, equalling or in part exceed- 
ing the corolla; filaments narrow-linear, patently 
laxly hairy in the lower half, glabrous above; 
anthers narrow-oblong, base obtuse, 2.5 by 1 mm. 
Disk glabrous. Ovary subcylindric, closely covered 
with scales, which touch or even overlap each other 
(and possibly bide some short hairs), c. 6 by 1.7 
mm, gradually tapering to the style, the latter 
rather slender, ir as long as the corolla, lepidote 
in the lower third, moreover laxly patently short- 
hairy in the lower 2/3, glabrous above; stigma 
capitate, shortly 5-lobed. 

Distr. Malesia: W. New Guinea (Swart R. 
Valley, c. 138° 15'-3= 30'), once found at 1600- 
2000 m. Fl. April. 

107. Rhododendron stapfianum Hemsl. ex Prain, 
Bot. Mag. (1911) sub t. 8372; Millais, Rhodod. 
(1917) 247; Sleum. Reinwardtia 5 (1960) 119; 
Blumea 11 (1961) 117; ibid. 12 (1963) 102. — 
R. lacteum Staff, Trans. Linn. Soc. II, Bot. 4 
(1 894) 197, t. 1 5, f.A 1-2; Merr. En. Born. (1921) 
461, non Franch. 1886. 

Epiphytic shrub. Branchlets terete, densely 
softly greyish tomentellous and laxly caducously 
stellate-lepidote at the upper 3-5 internodes, 
glabrescent below; internodes 1.5-3(-4) cm. 
Leaves scattered, mostly close together towards the 
apex of the internodes, not rarely in (3-)4-7- 
merous pseudowhorls at the very nodes, linear- 
oblong, sessile or nearly so, apex — obtuse, base 
truncate, coriaceous or subcoriaceous, shortly 
soft-hirtellous on both faces, longer and more 
densely patent-hairy at the margin and along the 
midrib beneath, never completely glabrescent, 
furthermore initially covered with rusty scales 
beneath (scales star-shaped, minute, on epidermal, 
finally ± disappearing protuberances, very deeply 
incised similar to a stellate hair, rather early 
caducous for their greater number), margin 
strongly revolute. (2.5-)3-5(-5.6) by (0.6-)l-1.5 
(-1.7) cm, midrib impressed above, manifestly 
obtusely raised beneath the full length, nerves 
obsolete; petiole 1-2 by 1-1.5 mm. Umbels 5-7- 
flowered. Perulae ovate to spathulate-obovate, 
sericeous outside, up to 2 by 1.5 cm, forming a 
thick obovate bud (3 by 1 .5 cm) before the anthesis. 
Bracteoles linear to linear-subspathulate, laxly 
hairy up to 3 cm by 0.5-1.5 mm. Pedicels rather 
stoutish, densely whitish tomentulose and cadu- 
cously lepidote, (3-)4-6(-8) mm, thickened to- 
wards the calyx. Calyx minute, shortly cup-shaped, 
densely hairy, lobes rather obsolete. Corolla 
salver-shaped, white or very lightly sulTuscd with 
pale violet, or pale pink, tube ± straight, sub- 
densely white-hairy and very laxly stcllate-lcpidotc 
outside, glabrous inside, (4-)4.5-5.5 cm by c. 2.5 
mm, lobes obovate to obiong-clliplic, i: spreading 
horizontally, glabrous on both faces. 1-1.5 by 
0.6-0.7 cm. Stamens 10, cxscrtcd for 5-6 mm; 
filaments linear, glabrous or practically so; 
anthers broad-oblong, (l-)1.5 by 0.8 mm. Disk 
hairy. Ovary cylindric, densely clad with fulvous 
or whitish greyish apprcsscd hairs which completely 



552 



Flora Malesiana 



[ser. I, vol. 6^ 



cover the scales, (6-)7-8 by 1.5-2 mm, gradually 
tapering to the style, the latter rather slender, 
exserted, subdensely hairy and laxly lepidote to 
nearly the top, 4(-5) cm; stigma short-obconical, 
lobuiate. Capsule reddish, 3-3.5 by 0.4 cm, oblong- 
fusiform, slightly 10-ribbcd, crowned by the rest 
of the style, subdensely rufous-tomentellous and 
stellate-lepidote. 

Distr. Malesia: Borneo (Mt Kinabalu; and 
Mt Palimasan in W. Kutai). 

Ecol. Mossy ridge forest, (800-) 1065-1 525 m. 
Fl. Jan. -Dec. 

108. Rhododendron chamaepitys Sleum. Bot. Jahrb. 
71 (1940) 144; Reinwardtia 5 (1960) 119; Blumea 
12 (1963) 112. 

Shrub. Branchlets terete, tips rather densely 
papillose-puberulous, glabrous elsewhere; inter- 
nodes 3-5 cm. Leaves in 7-10-merous, spaced 
pseudowhorls, linear or oblong-linear, apex ob- 
tuse, base broadly attenuate, coriaceous, glabrous 
above at full age, subdensely stellate-lepidote 
beneath (scales incised at the marginal zone 
almost to the small centre, each on top of a very 
minute epidermal elevation, ± early caducous), 
strongly revolute parallel to the margin, 3-5 cm 
by 3.5-5(-7) mm in revolute state, midrib im- 
pressed above, somewhat raised in its lower part 
beneath, nerves obscure; petiole 1-1.5 by 1 mm. 
Umbels 4-6-flowered. Ferulae ovate to ovate- 
lanceolate, long-acuminate, acute, glabrous, ciliate, 
5-7 mm. Bracteoles linear, up to 1 cm. Pedicels 
slender, densely rusty-lepidote and very shortly 
pubescent, 8-10 mm. Calyx obliquely disk-shaped, 
wavy and indistinctly 5-lobed, c. 2.5 mm 0. 
Corolla salver-shaped, colour not known, tube 
(3-)3.5-4 cm, 2-3 mm at base, a little wider at 
the throat, laxly rufescent-lepidote outside, pubes- 
cent at the base inside, lobes rounded, very laxly 
lepidote outside, glabrous inside, wavy, c. 8-10 
by 5-7 mm. Stamens 10, slightly exserted from the 
throat; filaments filiform, glabrous; anthers 
broadly oblong, 1.5 by 1 mm. Disk glabrous. Ovary 
elongate-conical, 5-6 by 1.5 mm, densely lepidote 
and short-pubescent; style slender, laxly pubescent 
and lepidote to almost the top, 4-4.5 cm; stigma 
capitate. Capsule oblong-fusiform, densely lepidote 
sparsely hairy, 2.5 by 0.4 cm. 

Distr. Malesia: Borneo (Sarawak: Mt Lambia 
or Lambir), 300^50 m. Fl. May, fr. June. 

109. Rhododendron macrosiphon Sleum. Blumea 1 1 
(1961) 118. 

Subscandent shrub, c. 4 m. Branchlets robust, 
caducously lepidote, pruinose; internodes 6-9 cm. 
Leaves in 4— 5-merous pseudowhorls at the upper 
nodes, obovate-elliptic, broadly to very broadly 
attenuate towards the apiculate apex, or apex 
obtuse, rarely rounded, base cuneate and slightly 
decurrent into the petiole, very coriaceous, rigid, 
similar to those of R. scabridibracteum in form and 
nervation, glabrous above with age, densely 
lepidote beneath (scales minute, variously stellate- 
dented in the hyaline, early caducous, marginal 
zone, centre thick, rather small, deeply impressed), 



± flat, 7-10 by (4— )4.5-6 cm, midrib narrowly 
impressed above, as stout as the petiole and much 
prominent in the lowest part, decreasing upwards 
beneath, nerves 6-7 irregular pairs, with shorter 
ones (or veins) between them, all rh spreading 
and anastomosing, strongly impressed above, not 
much raised beneath, reticulation faintly sunk 
above, slightly prominent or sometimes rather 
obscure beneath; petiole scmiterete, grooved 
above, densely lepidote initially, (2-)2.5 by c. 0.3 
cm. Umbels 8-10-flowered. Outer perulae not 
known, inner ones elongate-spathulate, hirsutulous 
on both faces, up to 3 cm. Pedicels very densely 
lepidote, (4^)5-8 by c. 1.5 mm. Bracteoles linear, 
glabrous, up to 2.5 cm. Calyx obliquely disk- 
shaped, irregularly 5-lobed, densely lepidote 
dorsally, 5-6 mm 0. Corolla tubular below, rather 
abruptly expanded at the lobes, white, subdensely 
or more laxly lepidote at the tube and the median 
part of the lobes outside, with dense appressed 
white retrorse hairs (c. 1 mm) at the tube inside, 
glabrous otherwise, tube 6-7 cm long, c. 5 mm 
at base, gradually dilated to 1-1.5 cm across the 
throat, lobes subspathulate-obovate or sub- 
orbicular, 2.5-3 by 2.5(-3) cm. Stamens 10, 
exserted for 1-1.5 cm from the throat; filaments 
linear (1 mm) and densely clothed with retrorse 
hairs below, more laxly hairy and slender upwards, 
glabrous and filiform for the ultimate 1.5 cm; 
anthers elongate-oblong, 7-8 by 1.5 mm; each 
theca narrowed into a distinct basal appendix 
(c. 1 mm). Disk thick, 10-lobed, more densely 
hairy at than between the lobes. Ovary cylindric, 
c. 1.5 by 0.3 cm at anthesis, densely covered with 
rather coarse yellowish, forward directed hairs 
which cover the scales, gradually tapering to the 
style, the latter 1.5-2 mm below, gradually 
decreasing upwards, hirsute for the lower 2/3, more 
laxly hairy and ± caducously lepidote above, 
glabrous for the ultimate 10 mm; stigma capitate, 
c. 2 mm 0, lobes thick. 

Distr. Malesia: Central New Guinea (Star Mts: 
near the mouth of the Minam R. into the Bon R.), 
one collection. 

Ecol. In a depression on peaty ground in 
primary forest, 1500 m, rather common locally. 
Fl. July. 

110. Rhododendron carringtoniae F. v. M. Vict. 
Nat. 4 (1887) 110, reimpr. Bot. Centralbl. 33 
(1888) 55; Descr. Not. 9 (1890) 63; Sleum. 
Reinwardtia 5 (1960) 120. 

Tall shrub, up to 5 m. Branchlets terete, stoutish, 
tips rather densely covered with thin scales; upper 
internodes 2-3 cm. Leaves scattered or mostly in 
3-4-merous pseudowhorls, obovate to elliptic- 
obovate, apex very broadly obtusely attenuate or 
mostly rounded, broadly attenuate towards the 
base, the very base rounded or mostly truncate, 
rarely broadly subcuneate, stiff-coriaceous, practi- 
cally glabrous above at full age, subdensely to laxly 
lepidote beneath (scales thin, their marginal zone 
± persistent, ± shallowly and irregularly dentate, 
centre small, impressed), slightly or not recurved 
at the margin, (3.5-)4-7 by (2.5-)3-4.5 cm, midrib 



Jan. 1966] 



Ericaceae (Sleumer) 



553 



somewhat impressed above in its lower half, 
strongly prominent beneath, but less so towards 
the apex of the blade, nerves 6-8 pairs, often 
forked before the edge, both anastomosing among 
each other and with the stronger veins, the latter 
laxly reticulate, prominent as are the nerves on 
both faces, veinlets reticulate, less manifestly, but 
well visibly raised on both faces; petiole somewhat 
flattened and grooved above, lepidote, 2-5 (-6) 
by 2-2.5 mm. Umbels about 6-flowered. Ferulae 
roundish to obovate-spathulate, dorsaliy lepidote, 
not or only laxly short-pubescent, up to 2.5 by 
1.5 cm. Bracteoles linear. Pedicels stoutish, sub- 
densely lepidote, 5-8 mm. Calyx rim-like, membra- 
nous, irregularly and very shortly 5-dentate, 
fimbriate, c. 2 mm o. Corolla salver-shaped, white, 
tube cylindric, slightly curved, 5-6(-7) cm long, 
rather equally 4.5-5.5 mm across, subdensely 
lepidote outside, copiously set from the base to 
the middle with short ± spreading hairs, glabres- 
cent upwards inside, lobes obovate to broadly 
elliptic-orbicular, spreading, 1-1.5 by (0.6-)l-1.3 
cm, very laxly lepidote outside, glabrous inside. 
Stamens 10, in part slightly exserted; filaments 
linear, densely clad with spreading hairs below, 
gradually glabrescent distally; anthers subobovate- 
oblong, 5-6 (rarely -7) by 1-1.2 mm, base of 
each theca with a distinct spur-like subobtuse or 
acute appendix (0.5-1 mm). Disk densely hairy. 
Ovary narrow-cyiindric, slightly attenuate towards 
the base, densely covered with yellowish whitish, 
ascending hairs which cover numerous scales, 
c. 1 cm by 1.5 mm, gradually attenuate to the 
rather thick style (5.5-6.5 cm), which is hairy and 
i densely lepidote except the glabrous 6 mm 
below the shortly turbinate stigma. Capsule 
narrow-cyiindric, short-attenuate at both ends, 
with 5 thick ribs, subdensely hairy and lepidote, 
c. 5 cm by 6 mm. Seeds with very narrow appen- 
dages. 

Distr. Malesia: SE. New Guinea (Owen 
Stanley Range: Mts Obree and Victoria; Mt 
Dayman). 

Ecol. In open hillside, declivities, along rocky 
river banks, 1830- 2440 m. Ft. Jan., June. 

111. Rhododendron carstcasense Wernh. Trans. 
Linn. Soc. Bot. II, Bot. 9 (1916) 96; Sleum. 
Reinwardtia 5 (1960) 132. 

Shrub. Branchlctssubangular,pruinose, smooth, 
early glabrescent; internodes 7-10 cm. Leaves in 
3-4-mcrous pseudowhorls, elliptic to obovate- 
elliptic, apex obtusely and very broadly attenuate 
to rounded, base short-attenuate into the petiole, 
glabrous above at maturity, subdensely lepidote 
beneath (scales small, marginal zone irregularly 
dentate, centre a little impressed), 6.5-10 by 
4-5.5 cm. midrib somewhat sunk above, rather 
strongly prominent beneath, nerves 8-10 pairs, 
rather irregular, straight below, curved anastomos- 
ing before the edge, slightly impressed above, not 
much prominent beneath, veins laxly reticulate, 
raised beneath only, veinlets • obscure; petiole 
grooved above. 1-1.5 cm by c. 2 mm. Umhets 
6-9-flowercd. Ferulae obovate-lanceolate, dorsaliy 



densely sericeous in the middle and with longer 
and i appressed hairs in the lower part on both 
faces, up to 4 by 1.8 cm. Bracteoles filiform, 
pubescent, 3-5 cm. Fedicels stoutish, lepidote, c. 
5 mm. Calyx minute, oblique, annular, obscurely 
5-lobed. Corolla salver-shaped, c. 1 cm long in 
all, tube cylindrical, hardly widened upwards, c. 
5.5 by 0.7 cm, glabrous outside, densely hairy 
inside in the lower half, lobes obovate-subrotun- 
date, c. 1.7 by 1.4 cm, glabrous. Stamens 10, some- 
what exserted from the throat; filaments linear, 
densely hairy to *l^, glabrous distally; anthers 
oblong, c. 6 by 1 mm, each cell caudate-appendicu- 
late (1-1.5 mm) at base. Disk subglabrous. Ovary 
elongate-cylindric, c. 7 by 1.8 mm, very densely 
clothed with yellowish hairs, gradually tapering 
to the style, the latter yellowish pubescent in the 
lower third, glabrous otherwise; stigma thick- 
obconical. 

Distr. Malesia:'^. New Guinea (MtCarstensz), 
once found at 1675-2040 m. 

112. Rhododendron syringoideum Sleum. Blumea 
12 (1963) 104. 

Terrestrial or sometimes epiphytic shrub, c. 2 m. 
Branches robust, erecto-patent; branchlets obtus- 
angular or laterally compressed, upper 2 internodes 
laxly stellate-lepidote, (5-)8-l2 cm by 3-5(-6) 
mm. Leaves in 5-6-merous pseudowhorls at the 
upper 3-6 nodes or mostly laxly arranged in the 
upper third of the internodes, broadly elliptic or 
subovate- or subobovate-elliptic, apex broadly 
attenuate to rounded, sometimes slightly retuse, 
base broadly attenuate to rounded, or rarely even 
subcordate, coriaceous and stiff, laxly set with 
rufescent scales on both faces initially (scales 
angular or dented to various degree, slightly 
impressed with the small centre), 6-9 by 4-5.5 
(-6) cm, a little revolutc at the very edge, especi- 
ally towards the base, midrib thick and prominent, 
gradually becoming flattish distally above, much 
raised below, less so towards the apex of the blade 
below, nerves 7-8 pairs, rather irregular, in part 
deeply forked, rather indistinctly anastomosing 
before the edge, slightly impressed above, promi- 
nent below, veins obscurely reticulate below; 
petiole 1-2 cm by c. 2 mm. Umbels 8-l2(-14)- 
flowered. Outer perulae ovate to roundish, r. 2 cm 
across, inner ones spathulate, up to 3.5 by 1.8 cm, 
innermost ones clongatc-spathulate, up to 5 by 
1 cm, all dorsaliy laxly short-hairy to glabrescent, 
internally more densely and longer appressedly 
substrigose. Bracteoles subspathulate-lincar, hairy 
especially along the margin, 4-5 cm by 2-5 mm. 
Fedicels laxly lepidote, 5-7 by 1.5-2 mm. Calyx 
much oblique, patellar or only rim-likc, obscurely 
lobed, c. 5 mm across. Corolla trumpct-shapcd, 
known only in a rather withered state, apparently 
fleshy and white when fresh, a little zygomorphous, 
scantily lepidote or glabrous outside, subdensely 
hairy for V* of the tube inside, (8.5-)9-IO( -10.5) 
cm long in all, tube slightly curved, 5-6 mm o at 
base, gradually dilated to 6-8 mm at the mouth. 
lobes \ spreading, obiong-obovatc. 1-1.5 by c. 
0.8 cm. Stamens 10, almost equalling the corolla; 



554 



Flora Malesiana 



[ser. I, vol. 6* 



filaments linear and hairy for c. y^ below, narrower 
and glabrous above; anthers narrow-oblong, wider 
distally, 5-6 by 1-1.3 mm, base of each cell pro- 
tracted into a subacute 0.3-0.4 mm long apiculus. 
Disk hardly prominent, glabrous. Ovary elongate- 
cylindric, 12-15 by 3 mm in a developed state, 
densely set with whitish or yellowish appressed 
hairs, which cover the scales, tapering to the 
rather slender style, which is almost as long as the 
corolla, and densely covered with hairy and scales 
below, less hairy and more lepidote upwards, 
glabrous for the ultimate centimetre; stigma 5- 
lobed, c. 1 .5 mm 0. Fruit apparently up to 5 cm by 
7 mm. 

Distr. Malesia: W. New Guinea (Arfak Mts: 
Mt Saru-mot, Anggi Gigi Lake). 

Ecol. Both epiphytic in rather dense forest and 
terrestrial in ± open devastated Nothofagus 
summit forest or scrub, 2200-2350 m. Fr. Jan. 

113. Rhododendron mains (J. J. S.) Sleum. Rein- 
wardtia 5 (1960) 120; Blumea 11 (1961) 119. — 
R. carringtoniae F. v. M. var. maius J. J. S. Nova 
Guinea 12 (1917) 511; ibid. 12 (1918) t. 205. 

Terrestrial or epiphytic shrub, up to 2 m. 
Branchlets robust, terete, tips laxly lepidote; the 
only internode known 14 cm by 4.5 mm. Leaves 
6-10 in a pseudowhorl, oblong to elliptic-oblong, 
apex broad-attenuate, obtuse, base broadly cuneate 
to subtruncate-obtuse, coriaceous, margin slightly 
revolute, subdensely minutely impressed-punctu- 
late above with age, subdensely and i persistently 
lepidote beneath (scales dark, marginal zone 
coarsely dentate or incised, centre somewhat im- 
pressed), (4.7-)6-8.7 by (2.2-)2.5-3.4(-4) cm, 
midrib slightly impressed above, strongly promi- 
nent beneath, though more slender towards the 
apex of the lamina, nerves 9-10 pairs, lower 4-5 
ones slightly curved, upper ones more straight, 
all ± branched distally and anastomosing with 
each other and the rather laxly arranged veins, 
nerves and veins slightly impressed above, rather 
prominent beneath, the finer reticulation ± dense 
and visibly prominent beneath; petiole lepidote, 
(4-)6-10byc. 1.5mm. f/w6e/5(5-)12-15-flowered. 
Ferulae ovate to obovate, short-pubescent and 
lepidote outside, up to 2.5 by 1.5 cm. Bracteoles 
linear to spathulate, 1-2 cm by 0.5-2.5 mm, 
sparsely pubescent or glabrous. Pedicels thickened 
below the oblique calyx, subdensely covered with 
thin rufescent scales, 2-2.5 cm by 1 mm. Calyx 
small, c. 6 mm 0, 5 -dentate, teeth irregularly 
triangular, obtuse or acute, one of them mostly 
longer (up to 2 mm). Corolla salver-shaped, 
white at the lobes, pinkish at the tube, or even 
darkening to red at the base, carnation-scented, 
tube slightly curved, subcylindric, 8-8.8 cm long, 
3-4 mm at base, gradually dilated to 6.5 mm at 
the throat, laxly appressed-lepidote outside (scales 
thin, irregularly dentate), inside rather densely 
hairy in the lower, more laxly so in the upper 
part, lobes obovate to orbicular, rounded, glabrous 
except some scales which may be found at the 
sinuses or the base of the lobes, 1.4-1.7 by 1.1-1.7 
cm. Stamens 10, somewhat exserted from the 



throat; filaments linear, laxly hairy in the lower 
half, gradually glabrescent upwards, at last com- 
pletely glabrous; anthers narrow-oblong, 3.5 by 
1 mm, base obtuse. Disk laxly appressed-puberu- 
lous or nearly glabrous. Ovary cylindrical or 
elongate-conical, 11-12 by 2 mm, densely sub- 
appressed-tomentose by coarse yellowish hairs, 
which cover the scales, tapering gradually to the 
style, the latter 5-5.5 cm, rather densely patent- 
hairy below, laxly hairy and lepidote in the middle, 
glabrous in the upper third; stigma capitate- 
turbinate, crenulate. 

Distr. Malesia: W. New Guinea (Hubrecht 
Mts and Habbema Lake). A form with but 5- 
flowered umbels on Mt Wilhelm. 

Ecol. In open places or on open banks of a 
stream within the mossy forest, 2770-3000 m. 
Fl. Feb.-Oct. 

114. Rhododendron cruttwellii Sleum. Reinwardtia 
5 (1960) 120; Blumea 12 (1964) 339. — Fig. 24. 

Small tree, 4.5-6 m. Branchlets subterete; 
internodes (2.5-)4-14 cm, upper ones laxly 
lepidote, lower ones greyish corticate. Leaves in 
5-7-merous pseudowhorls, part of them reduced 
in size, normal ones obovate-elliptic or -lanceolate, 
sometimes elliptic-lanceolate, apex shortly ± 
gradually acuminate, subacute, with a small 
terminal gland, base ± acutely attenuate or 
cuneate to the petiole, subcoriaceous or coriaceous, 
dark green and shining above, paler and rather dull 
beneath, glabrous above, laxly to subdensely 
lepidote beneath (scales in the rather broad and 
very thin marginal zone variously dentate or 
lacerate, the centre small, somewhat sunk and 
longer persistent), entire, subrevolute, (5.5-)6-12.5 
by 3-6 cm, midrib slightly depressed above, raised 
beneath, nerves 8-10 pairs, lower ones irregular, 
upper ones more straight and subparallel, curved- 
anastomosing with each other in a certain distance 
from the edge, prominent on both faces, reticula- 
tion dense and slightly but well visibly raised on 
both faces; petiole ± flattened, grooved above, 
lepidote, 6-12 by 1.5-2 mm. Umbels 4-7 (-9)- 
flowered. Ferulae ovate-acuminate, laxly lepidote 
in the middle dorsally, furthermore sericeous all 
over the upper- and undersurface, glandular- 
ciliate, up to 1.5 by 1 cm. Bracteoles narrow- 
linear, glabrous, up to 1.5 cm. Fedicels stoutish, 
laxly patent-hairy, lepidote especially in the upper 
third, l-1.5(-2) cm, obliquely thickened below 
the calyx. Calyx small, margin membranaceous, 
indistinctly 5-lobed, rather glabrous, very laxly 
ciliate, c. 2.5 mm 0. Corolla salver-shaped, pure 
white, tube slightly curved, a little 5-saccate at 
the very base, but slightly dilated towards the 
mouth, (5-)5.5-6 cm, at the base 3-4 mm 0, in 
the middle c. 5 mm, sparsely lepidote outside, 
± densely hairy in the lower half, glabrous 
distally inside, lobes spreading, broadly obovate 
to suborbicular, 1-1.3 by 0.9-1.3 cm, glabrous. 
Stamens 10, somewhat exserted from the throat; 
filaments linear, hairy in the lower half, glabrous 
above; anthers oblong, base obtuse, 2-2.3 by 1 
mm. Disk very low, glabrous. Ovary cylindric. 



Jan. 1966] 



Ericaceae (Sleumer) 



555 



s^l'j'ij 



0^ 




Fig. 24. Rhododendron cruttwellii Sleum. a. Branch with flowers, X 2/3, b. flower, X 1, c. stamen, x 1, 
f/. ovary and style, x 1,^. anther, x 4,/, ovary, x 2 (a-/CRUTTW^ELL 1020). 



gradually narrowed to the slender style, densely 
covered with subpatent whitish or greyish hairs, 
no scales, 6-7 by 1.5 mm; style exserted, c. 6 cm, 
at the base laxly hairy for c. 6 mm, completely 
glabrous otherwise; stigma turbinate-capitate, 
crcnulatc. 

Distr. Malesia: SE. New Guinea (Nfaneau 
Range: Mt Maneao, Mt Yauyama; Mt Simpson). 

Ecol. In thin evergreen forest, edge of small 
patch of wood in gully, among non-calcareous 
rocks in shades, on moss-covered ground, 1860- 
2600 m. Fl. Junc-Aug. 

115. Rhododendron archboldianum Slbum. Rcin- 

wardtia 5 (\')U)) 121. 

Tall shrub, epiphytic or rarely terrestrial. 
Branchlcts terete, stout; intcrnodes 6-16 cm, 
ultimate ones subdenscly Icpidolc. lower ones 
glabrcsccnt. Leaves in 5 -T-mcrous pscudowhorls, 
in which 3-4 leaves arc normal, the rest smaller 
(reduced), apparently with some very small and 
early caducous leaflets along the intcrnodes, their 



scars only seen; normal leaves ovate to elliptic- 
ovate, not rarely somewhat unequal, apex short- 
acuminate, subacute, base broadly attenuate or 
mostly subtruncate-rounded, sometimes sub- 
cordate, coriaceous, subdensely lepidote on both 
faces (scales tardily caducous above, persistent for 
a long time beneath, flat, small, the margin 
irregularly and rfc deeply dentate, the centre not 
immersed), entire, subrevolute below, (4~)5-ll 
by (2-)3-6 cm, midrib slightly depressed above, 
thick-prominent beneath in the lower, less so in 
the upper part, nerves 8-10(-12) pairs, slightly 
curved, i; irregular, anastomosing with each 
other and the veins before the edge, faintly im- 
pressed or elevated above, raised beneath, reticula- 
tion dense, more visible beneath only; petiole a 
little flattened, grooved above, subdensely lepidote, 
(4-)6-9 (sometimes up to 13) by 1.5-2 mm. 
Umbels 8-1 3(-l 5)-flowcred. Ferulae ovate to 
broadly obovate, laxly lepidote and sericeous 
dorsally. up to I by 0.7 cm. Bractcoles narrow- 
linear, glabrous or laxly lepidote, up to 1.5 cm. 



556 



Flora Malesiana 



[ser. I, vol. 6* 



Pedicels stoutish, rather densely lepidote, oblique- 
ly thickened below the calyx, 8-1 1 mm. Calyx 
very small, rim-like, wavy, indistinctly 5-lobed. 
Corolla salver-shaped, rose, tube cylindric, 
slightly curved, gradually dilated from the base 
to the limb, 4.5-5 cm long, c. 2.5 mm at base, 
3.5-4.5 mm higher, a little constricted at the 
throat, laxly to subdensely lepidote outside, it 
densely hairy inside to nearly the throat, lobes 
broadly spathulate to nearly rounded, 1-1.2 by 
0.8-1.2 cm, laxly lepidote outside at the base and 
median, glabrous inside. Stamens 10, exserted; 
filaments linear below, filiform above, short-hairy 
to nearly the top; anthers oblong, base obtuse, 
3.5-4 by 1 mm. Disk glabrous below, densely 
hairy at the upper margin. Ovary cylindric, 
gradually tapering to the style, 7-8 by c. 2 mm, 
densely appressed-hairy and lepidote (scales in 
covered by the tomentum and not well visible); 
style rb as long as the corolla tube, covered with 
subpatent hairs and scales to nearly the top; 
stigma thick, short-obconical. 

Distr. Malesia: SE. New Guinea (Mts Tafa 
and Suckling). 

Ecol. Usually epiphytic in ridge forests, 
occasionally terrestrial in clearings, 2400 m. 

116. Rhododendron pleianthum Sleum. Reinwardtia 
5 (1960) 122; Blumea 11 (1961) 119; ibid. 12 
(1963) 103. 

Small tree or shrub, terrestrial, 2^(-6) m. 
Branchlets very robust, terete, tips laxly stellate- 
lepidote, 5-7 mm 0; internodes 9-16 cm. Leaves at 
the upper 1 or 2 nodes in 5-7-merous pseudo- 
whorls, elliptic or obovate-elliptic, apex rounded 
or very broadly attenuate and blunt, base slightly 
but distinctly cordate, coriaceous, ± rigid, 
glabrous above at full age, laxly lepidote beneath 
(scales stellate-incised at the thin, early dissolute 
marginal zone, centre small, but slightly im- 
pressed), finally minutely impressed-punctulate 
beneath, flat, (6.5-)7-10(-14) by (4-)4.5-6.5(-9) 
cm, midrib dilated below, gradually narrowed 
upwards, impressed for the whole length above, 
thick and obtusely prominent beneath, 2-3 mm 
broad at the base, nerves c. 8 pairs, irregular, 
± spreading, divided and anastomosing before 
the edge, impressed above, prominent beneath, 
veins and veinlets forming an impressed lax 
reticulation above, which is but faintly raised 
or ~ obscure beneath; petiole flattened, glabrous 
2^(-6) by 2-3 mm. Umbels (8-)15-20-flowered. 
Outer perulae ovate, inner ones oblong-spathulate, 
obtuse, at the back densely, inside more laxly 
shortly substrigose or almost sericeous, up to 2 by 

1 cm. Bracteoles linear-subspathulate, more 
densely short-hairy at the dorsal middle line, c. 

2 cm by up to 2 mm. Pedicels densely lepidote and 
very shortly pubescent, c. 2 cm by 1 mm at 
anthesis. Calyx disk-shaped, slightly oblique, c. 
4 mm 0, subglabrous dorsally, shortly subacutely 
5-lobed, or occasionally with irregular 2-9 mm 
long teeth or laciniae, which are glabrous or very 
shortly pubescent, c. 1 mm wide at base. Corolla 
long-tubular below, ± expanded at the lobes, 



8-9 cm long in all, initially very laxly and cadu- 
cously lepidote outside, laxly hairy at the tube 
inside, white or cream, suff'used with pink distally, 
or dark pink, tub€ rather straight, (5.5-)6-7 
cm long, 0.4-0.5 cm at base, gradually dilated 
upwards to 0.7-0.9 cm, lobes obovate-spathulate, 
c. 2 cm across. Stamens 10, slightly longer than 
the corolla tube, unequal; filaments linear, sub- 
densely or towards the apex more laxly hairy in 
the lower ^^ , glabrous for the rest ; anthers elongate- 
oblong, base obtuse, 4.5-5 by 1-1.2 mm, a little 
curved. Disk prominent, glabrous. Ovarvelongate- 
cylindric, gradually tapering to the style, very 
densely yellowish-hairy or -tomentose, rather laxly 
lepidote (scales — covered by the hairs), c. 8 by 
2-2.5 mm; style nearly as long as the corolla tube, 
for about 10 mm subdensely to laxly hairy and 
lepidote above the base (the scales well visible 
among the hairs), glabrous otherwise; stigma tur- 
binate-capitate. Cap^w/f elongate-fusiform, densely 
hairy and lepidote, (5-)6-8 by c. 0.7-0.8 cm. 

Distr. Malesia: New Guinea (Western, 
Eastern and Southern Highlands: Mts Hagen, 
Kerigomna, Wilhelm, Otto, and Giluwe). 

Ecol. On edge of mossy forest-grassland, in 
ridge crest scrub or on high bank above creek, 
locally common, 2680-3260 m. Fl. July-Oct. 

Vern. Wambu, Mendi. 

117. Rhododendron oliganthum SLEtJM. Reinwardtia 

5 (1960) 123. 

Shrub, c. 1.2 m. Branchlets terete, slender, tips 
densely rufous-lepidote; upper internodes 1.5-3 
cm. Leaves nearly opposite or in a pseudowhorl 
of 3 at the upper node only, subsessile, subovate- 
elliptic, apex shortly acuminate, obtuse, base 
truncate to rounded or subcordate, coriaceous, 
glabrous above at maturity, laxly lepidote 
beneath (scales thin, marginal zone irregularly 
dentate or incised, centre slightly impressed), 
margin somewhat recurved, (3-)3.5-5 by (1.5-) 
2-3 cm, midrib faintly impressed or nearly flat 
above, strongly raised beneath in the lower part, 
less so towards the apex of the lamina, nerves c. 
8 pairs, rather straight, curved-anastomosing near 
the edge, prominent on both faces, veins laxly 
reticulate, raised specially beneath together with 
the veinlets; petiole 1-2 by 1-1.5 mm. Flowers 
solitary or in twos, terminal. Pedicels stoutish, 
oblique thickened below the calyx, rather densely 
lepidote and very shortly pubescent, 2.8-3 cm at 
anthesis. Calyx minute, shortly 5-lobed and wavy, 
c. 2.5 mm 0. Corolla broadly salver-shaped, 
intensely rose when fresh, tube cylindric, straight, 
3.5-3.7 cm long, at base c. 4 mm 0, slightly dilated 
to 5-6 mm at the mouth, laxly lepidote and very 
laxly short-hairy outside, subdensely hairy inside 
in the basal part, less so upwards, glabrous at 
the throat, lobes ± spreading, broadly obovate to 
nearly rounded, 1-1.2 cm 0, lepidote outside at 
the base, otherwise glabrous. Stamens 10, some- 
what exserted from the throat; filaments linear, 
subdensely to laxly hairy to the lower 2/3, glabrous 
distally; anthers oblong, base obtuse, c. 2.6 by 1 
mm. Disk pubescent especially at the upper margin. 



Jan. 1966] 



Ericaceae (Sleumer) 



557 



Ovary elongate-conical, densely hairy and lepi- 
dote, c. 7 by 2 mm, with 5 longitudinal furrows, 
gradually tapering to the style, the latter slightly 
exserted. c. 3.5 cm, laxly hairy and very laxly 
lepidote up to the middle, glabrous above; stigma 
thickly turbinate-capitate. 

Distr. .V/a/e5/a: SE.New Guinea (Mt Victoria: 
NW of the "Gap"), in crest forest, at 2895 m. 
Fl. Jan. 

118. Rhododendron carrii Sleum. Reinwardtia 5 
(1960) 124. 

Epiphytic shrub. Branchlets terete, tips glabrous 
or very laxly lepidote; internodes (2.5)-4-9 cm. 
Leaves in 3-4-merous pseudowhorls at the upper 
3-4 nodes, ovate to subovate-rounded, apex 
broadly and shortly acuminate, rather obtuse, 
base manifestly subauriculate-cordate, sessile or 
nearly so, rigidly coriaceous, seemingly glabrous, 
but laxly set with minute irregularly star-shaped 
scales beneath (lens!) (leaving a minute point in 
the slightly impressed centre), flat, (4— )4.5-5.5 
(-6) by (2.'5-)3-5(-5.5) cm, midrib flat or slightly 
impressed above, stout and very prominent 
beneath in the lower part, less so towards the apex 
of the lamina, nerves c. 10 pairs, 2-3 of them 
from the base of the blade, the other from the 
midrib, ail curved-ascending and anastomosing 
near the edge, prominent on both faces, reticula- 
tion rather dense and well visible especially 
beneath; petiole c. 1 by 1.5 mm. Umbels 4—5- 
flowered. Bracteoles linear. Pedicels stoutish, 
subdensely to laxly thin-lepidote, (1.3-)1.5-2.5 
cm, thickened at apex. Calyx small, oblique, 
membranous, shortly (1 mm) obtusely 5-lobed, 
shortly fimbriate, c. 2 mm 0. Corolla salver-shaped, 
white, tube slightly curved, dilated gradually up- 
wards, 5.5-6 cm long in full anthesis, at base c. 
3.5 mm 0, below the lobes c. 7 mm, laxly sub- 
stellate-lepidote outside, densely clad inside with 
retrorse hairs in the lower third or half, upwards 
more laxly so with spreading hairs, glabrous below 
the throat, lobes t. spreading, rounded-obovate, 
1.2-1.5 by 1-1.5 cm, glabrous on both faces. 
Stamens 10, a little exserted; filaments narrow- 
linear, densely hairy at the base, more laxly and 
patently so in the middle, glabrous distally; 
anthers oblong, base obtuse, 2.8-3 by 1 mm. Disk 
densely short-hairy especially above. Ovary 
elongate-conical, densely clothed with appressed 
yellowish hairs and laxly clad with scales (± 
covered or hidden by the hairs), c. 10 by 2 mm, 
gradually extended into the style, the latter ex- 
serted, (5.5-)6-6.5 cm, laxly hairy and lepidote 
at the base for c. 5 mm only, glabrous otherwise; 
stigma thick-turbinatc, crcnulatc. 

Distr. Malesia: SE. New Guinea (Mt Victoria: 
NW of the "Gap"), once found on open hill side, 
2440 m. //. Jan. 

119. Khr>dr>dcndron artnitii F. M. Baiiiy, Bot. 
Bull. Oucc-nsl. Hep. A^r. 10 (1895) 39; Sleum. 
Rcmwardtia 5 (I9«)) 124. 

Shrub, dichotomously branched; bark whitish. 
Branchlets terete, tips sparsely to subdensely clad 



with stellate scales; internodes 3-7 (rarely up to 
12) cm. Leaves in ± 4-merous pseudowhorls at 
the end of the branchlets, broadly elliptic to 
subobovate-elliptic, apex very shortly obtusely 
attenuate, base rounded in general, the very base 
truncate to slightly subcordate, coriaceous, firm, 
subdensely lepidote on both faces initially, early 
glabrescent above (only hair-like flakes, being the 
remains of the scales, present for a longer time), 
more persistently lepidote beneath (scales thin, 
rusty, not rarely on very minute rather flat epi- 
dermal elevations which are still visible when the 
scales have gone being paler than the intervenium, 
marginal zone thin, irregularly deeply incised, 
centre very small), flat, 7-9(-10) by 4-5(-6) cm, 
midrib impressed above, very prominent beneath 
especially towards the base, nerves 8-10 pairs, 
rather straight basally. sometimes bifurcate half 
their length, obscurely curved-anastomosing before 
the edge, somewhat raised above (but impressed 
in the middle), prominent beneath, veins laxly 
reticulate, more visible beneath, veinlets obscure; 
petiole semiterete, grooved above, lepidote, 
(6-)8-12 by c. 2.5 mm. Umbels c. 5-flowered. 
Ferulae ovate to obovate-spathulate, densely 
hairy and laxly lepidote outside, fimbriate, up to 

3 cm by 1.5 cm. Bracteoles linear-spalhulatc, laxly 
hairy, c. 2 cm by up to 2.5 mm. Pedicels subdensely 
lepidote, somewhat hairy only below the oblique 
calyx, 1.8-2.5 cm by c. 1 mm. Calyx distinct, 
lepidote and laxly hairy outside, glabrous inside, 
deeply 5-lobed, lobes deltoid, subacute, 2-3 by c. 
2 mm at base, fimbriate, finally reflexed. Corolla 
salver-shaped, white touched with sea-shell pink, 
tube straight or slightly curved, 5.5-6.5 cm by 
6-7 mm 0, a little wider at the 5-saccate base and 
somewhat expanded below the mouth, very laxly 
(caducously) lepidote or practically glabrous out- 
side, densely long-hairy inside in the lower third, 
less densely so above, glabrous in the upper th'rd, 
lobes broad-obovate to nearly rounded, spreading, 
glabrous, 1.8-2.2 by 1.3-1.7 cm. Stamens 10, 
slightly exserted from the throat; filaments linear 
(c. 1 mm) and densely patently hairy in the lower 
third, glabrescent above, filiform and completely 
glabrous in the upper third; anthers oblong, 

4 by 1 mm, base obtuse. Disk glabrous. Ovary 
elongate-conical, 5-ribbed, c. 1.2 by 0.3 cm, 
densely set with subapprcsscd yellowish or whitish 
hairs, which cover some scales, gradually tapering 
to the style, the latter stoutish and ! densely 
hairy and very laxly lepidote in the lower, glabrous 
and more slender in the upper half; stigma broad- 
obconical, distinctly 5-lobcd. 

Distr. Malesia-.iiE. NewGuinea(Mt Dayman), 
once found on the summit, 2710 m. //. March. 

120. Rhododendron tuba Sii um. Reinwardtia 5 
(I960) 125. 

Terrestrial shrub, 2-5 m. Branchlets somewhat 
flattened, laxly lepidote, pruinosc when dry; inter- 
nodes 3 8( 10) cn\. /.<vyir.v in 4- 5-mcrous pseudo- 
whorls at the upper I or 2 nodes, elliptic to oblong- 
elliptic, apex broadly acuminate, subacute, 
terminal gland not protruding beyond the leaf 



558 



Flora Malesiana 



[ser. I, vol. 6* 



margin, towards the base ± broadly attenuate, 
very base subtruncate or rounded, sometimes sub- 
cordate, coriaceous, subdensely lepidote on both 
faces initially, glabrescent above and dark olivace- 
ous-green at full age, more persistently lepidote 
beneath (scales rather small, stellate-dented in the 
thin and early dissolute marginal zone, centre 
minute and impressed), finally laxly set with fine 
blackish pits beneath, flat, (4-)4.5-7 by 2.5-3.5 
(-5, rarely up to 6) cm, midrib depressed above, 
thick, broad and prominent in the lower part 
beneath, narrowed upwards, nerves 6-8 irregular 
pairs divided and anastomosing before the edge, 
slightly impressed above, a little though distinctly 
raised beneath, reticulation rather obscure above, 
well visible, subdense and prominent beneath; 
petiole flattened, 2-4 by 1.5-2 mm. Umbels (4-) 
5-7-flowered. Outer perulae ovate, inner ones 
obovate, ending in an apiculus or sharp point 
(1-3 mm), membranous, caducously lepidote at 
the tips dorsally, glabrous otherwise, a little 
shining, lepidote-ciliate, up to 1.5 by 1 cm. Brac- 
teoles filiform below, somewhat subspathulate- 
dilated distally, very laxly lepidote or glabrous, 
up to 1.5 cm. Pedicels laxly stellate-lepidote, 
epilose, 1.2-2 cm by c. 1 mm. Calyx \evy obliquely 
disk-shaped, membranous, subglabrous, c. 4 mm 
0, with 5 irregular short teeth (c. 1 mm) or longer 
laciniae (up to 4 mm). Corolla salver-shaped, 
laxly lepidote outside, laxly hairy in the lower ^/^ 
of the tube inside, 6.5-7.5(-8) cm long in all, 
salmon-pink at the tube, white at the lobes, tube 
mostly slightly curved, 5.5-6.5 cm long, c. 5 mm 
at the base, not or very slightly dilated upwards, 
lobes ± spreading, obovate, 1-1.5 by 0.8-1.2 cm. 
Stamens 10, unequal, subequalling the corolla; 
filaments linear, laxly hairy in the lower %, 
glabrous for the rest; anthers subobovate-oblong, 
base contracted to a very short apiculus, c. 2.5 
by 0.7 mm. Disk slightly prominent, hairy in the 
upper, glabrous in the lower part. Ovar>'cylindric, 
gradually tapering to the style, densely whitish 
hairy or -tomentose, minutely lepidote (the small 
scales covered by the hairs), c. 8 by 2.5 mm; style 
rather slender, as long as the stamens, laxly set 
with longish white hairs and scales in the lower 
half; stigma yellow, capitate, shortly 5-lobed. 

Distr. Malesia: SE. New Guinea (N. side of 
summit of Mt Dayman). 

Ecol. In margin of moss forest, at 2500-2750 m. 
Fl. June-Aug. 

Note. R. tuba is in many respects intermediate 
between R. carringtoniae and R. rhodoleucum, 
both also known from the Maneau Range, and 
possibly a local natural hybrid between them. 

121. Rhododendron rhodoleucum Sleum. Blumea 1 1 
(1961) 119. 

Stiff, straggly shrub, (1-) 1.5-3 (-4) m, much 
branched dista'ly. Branchlets rather robust, laxly 
lepidote at the upper internodes (3.5 cm by 2 mm), 
older parts early covered with thin ashy cork. 
Leaves in 4— 6-merous pseudowhorls at the upper 
1 (or rarely 2) nodes, (broadly) elliptic to sub- 
obovate-elliptic, (sub)sessile, apex shortly and 



broadly attenuate, obtuse, more rarely almost 
rounded, base ± deeply cordate, coriaceous, ± 
shining glabrous with age above, laxly lepidote 
beneath (scales flat, rather small, thin marginal 
zone rather wide, variously dented or angled, 
small centre slightly impressed), flat, (3-)4-7 by 
(2-)2.5-5(-6) cm, midrib narrowly impressed 
above, at base as wide as the petiole, decrescent 
distally, not much prominent beneath, nerves 6-8 
irregular pairs, with several equally spreading 
and anastomosing veins between them, all con- 
spicuously raised on both faces, as is the rather 
dense reticulation; petiole 0-1 mm. Umbels (4-) 
5-6-flowered. Bracteoles almost filiform. Pedicels 
stoutish, subdensely or more laxly stellate-lepidote, 
epilose, c. 1.5 cm. Calyx very obliquely disk- 
shaped, indistinctly 5-lobed, 2.5-3 mm 0, ± 
lepidote dorsally. Corolla tubular, rather straight 
or a little curved, laxly lepidote at the tube out- 
side, densely set with patent or retrorse hairs 
below, more laxly so upwards inside, lobes ± 
glabrous, tube red at the base, fading upwards to 
pink or salmon-pink, lobes white, tube 6-7(-7.5) 
cm long, 4-5 mm at base, gradually widened to 
c. 1 cm below the lobes, lobes ± spreading, 
obovate to subrotund, 1.5-2 cm. Stamens 10, a 
little exserted from the throat; filaments linear (c. 
1.2 mm) and subdensely hairy below, narrower 
and more laxly subpatent-hairy upwards, glabrous 
in the upper 14; anthers yellow, narrow-oblong, 
3.5-4 by c. 1 mm, base obtuse. Disk hairy in the 
upper margin. Ovary subcylindric, 7-8 by 2 mm, 
densely appressed-hairy and lepidote, tapering to 
the style, the latter rather slender, densely or 
more laxly hairy and laxly lepidote in the lower 2/3, 
exclusively sparsely lepidote for 3-4 mm above, 
glabrous for the rest; stigma thick-turbinate, 
shortly 5-lobed. Capsule fusiform, c. 3.5 by 0.5 
cm. 

Distr. Malesia: New Guinea (Milne Bay 
Distr.: 3 peaks of the Maneau Range). 

Ecol. On edge of (stunted) mossy forest, some- 
times on very steep escarpments, 2290-2750 m, 
locally abundant. Fl. May-Oct. 

122. Rhododendron pubitubum Sleum. Reinwardtia 

5 (1960) 126. 

Shrub. Branchlets dark brown when dry, ± 
pruinose, tips laxly or very laxly lepidote; inter- 
nodes 4-1 1 cm. Leaves in 3-5-merous pseudo- 
whorls, elliptic or obovate elliptic, sometimes 
obovate, apex very broadly attenuate, obtuse to 
nearly rounded, occasionally very shortly retuse, 
broadly attenuate towards the base, the very base 
rounded, not rarely subcordate, subsessile, 
coriaceous, glabrous above at maturity, laxly 
to subdensely lepidote beneath (scales thin, 
marginal zone irregularly dentate or bitten, ± 
persistent centre blackish and sunk), subrevolute, 
3-6 by 2.2-3.5(-4.2) cm, midrib slightly depressed 
above, much prominent beneath, evanescent 
distally, nerves 6-8 pairs, straight from the midrib 
at an acute angle, subparallel, curved anastomos- 
ing before the edge, very slightly impressed or ± 
obsolete above, prominent beneath, reticulation 



Jan. 1966] 



Ericaceae (Sleumer) 



559 



lax and slightly visible beneath only; petiole 
1-2 by 1-2 mm. Umbels 10-12-flowered. Ferulae 
orbiculate to obovate, densely short-subsericeous 
on both faces, up to 1.5 by 1 cm. Bracteoles linear, 
densely rather longish patent-hairy, up to 1.5 
cm by 0.5 mm. Pedicels stoutish, 8-12 mm, 
densely and patently greyish pubescent. Calyx 
minute, margin narrow, membranaceous, wavy 
and very shortly obtusely 5-lobed, somewhat hairy 
outside, glabrous inside. Corolla salver-shaped, 
white, scented, tube cylindric, straight, somewhat 
5-saccate at the very base, barely dilated towards 
the mouth, (5-)5.5-6.5 cm by c. 2.5 mm, rather 
densely pubescent outside, subdensely to laxly 
and patently hairy inside in the lower ^/j, lobes ± 
spreading, spathulate-obovate, hairy outside along 
the middle line, otherwise glabrous, 1-1.2 cm 
by 5-9 mm. Stamens 10, a little exserted; filaments 
narrow-linear, rather densely hairy below, more 
laxly so upwards, glabrous in the upper ^f^; 
anthers broadly obovate-oblong, base obtuse, 
1.8 mm. Disk glabrous below, pubescent at the 
upper margin. Ovary elongate-conical, densely 
covered with subpatent whitish hairs and scales 
(these covered or hidden by the tomentum), c. 
5 by 1.8 mm, rather abruptly narrowed to the 
style; style as long as or a little shorter than the 
corolla tube, subdensely hairy and laxly lepidote 
below (the scales well visible there), laxly hairy in 
the middle, glabrous for the last 10 mm below 
the short-obconical stigma. 

Distr. Malesia: Central Celebes (Masamba 
Distr.), two collections. 

Ecol. In open places, on whether or not peaty 
soil on granite, 1700-2000 m. Fl. July-Aug. 

123. Rhododendron pseudotrichanthum Sleum. 
Blumea 12 (1964) 340. — R. irichanthum Sleum. 
Reinwardtia 5 (1960) 127, non Rehd. J. Arn. Arb. 
26 (1945) 480. 

Shrub. Branchlets robust, subterete; internodes 
3-6 cm, upper ones rather densely lepidote and 
laxly hairy. Leaves in 3-4-merous pseudowhorls, 
oblong or elliptic-oblong, somewhat inequilateral, 
apex very broadly attenuate to rounded, not 
rarely minutely retuse, base very broadly attenuate 
to nearly rounded, thick-coriaceous, firm, glabrous 
above some hairs excepted, which are found at the 
very base, laxly lepidote beneath (scales small, i 
deeply substcllate-dcntate, flat, centre red brown, 
small, slightly impressed), subrcvolutc, 8-12 by 
(3-)3.5-5 cm, midrib distinctly impressed above, 
stout and prominent beneath in the lower part, 
less so distally, nerves 6-8 pairs, rather spaced, 
± straight below, curved-anastomosing before the 
edge, very slightly impressed above, a little 
prominent beneath, reticulation rather indistinct; 
petiole glabrous or clad with some hairs, rough, 
grooved above, 6-13 by 2-4 mm. Umhcls 7-10- 
flowercd. Pcrulae ovate, glabrous, irregularly 
shortly subscrratc, up to 1.3 by 1 cm. Bracteoles 
linear, laxly hairy, c. I cm. Pedicels rather slender 
initially, finally more stoutish, densely clad with 
i patent greyish hairs or nearly villous, very 
laxly lepidote, 2 3 ^sometimes up to 4) cm at 



anthesis. Calyx armular, inconspicuous, c. 1 mm 
0. Corolla salver-shaped, dilutely red-violet when 
fresh, tube cylindric, c. 3.5 cm long, at base some- 
what inflated and 5-saccate, 3-4 mm 0, hardly 
or not widened towards the mouth, densely clothed 
all over with soft greyish hairs outside, equally 
so inside in the lower half, lobes obovate-rounded, 
spreading, 1-1.2 by 0.5-0.7 cm, rather densely 
hairy outside margin excepted, glabrous inside. 
Stamens 10, unequal, ± as long as the corolla; 
filaments linear, flattened, densely whitish hairy 
to the lower Vi, glabrous for the rest; anthers 
oblong, base obtuse, 2.5 by 0.8 mm. Disk sub- 
glabrous. Ovary cylindric, 5-ribbed, very densely 
covered with subpatent hairs or almost villous, 
4-6 by 1.5-2 mm, tapering to the slender style, 
which attains c. 2/3 of the length of the corolla 
and is laxly set with spreading hairs in its lower 
third; stigma narrow-obconical. 

Distr. Malesia: Central E. Borneo (summit of 
Mt Kemul). 

Ecol. In ridge forest, on humous, peaty soil, 
1800-1850 m. Fl. Oct. 

124. Rhododendron amabile Sleum. Reinwardtia 
5 (1960) 127. 

Shrub. Branchlets subterete and densely lepidote 
at the ultimate, glabrous at the lower internodes 
(2.5-9 cm). Leaves in ± 4-merous pseudowhorls, 
subsessile, oblong or oblong-elliptic, sometimes 
elliptic, apex broadly attenuate, subobtuse, base 
subtruncate-rounded to subcordate, coriaceous, 
caducously lepidote, finally completely glabrous 
above, ± persistently and laxly lepidote beneath 
(scales irregularly dentate at margin, flat, each on 
top of a minute epidermal elevation), subrevolute, 
2.5-6 by (1.2-)1.5-2.3(-2.8) cm, midrib flat or 
faintly impressed above, stout and very prominent 
beneath in the lower 2/3, less so upwards, nerves 
7-8 pairs, rather straight below, curved-anastomos- 
ing near the edge, hardly visible above, somewhat 
raised beneath, reticulation ± inconspicuous; 
petiole lepidote, 2-3 by c. 1.5 mm. Umbels 7-9- 
flowered. Ferulae ovate-acuminate, acute, outer 
ones lepidote and subdensely sericeous dorsally, 
inner ones sericeous only, up to 1.4 by 0.9 cm. 
Bracteoles linear to subspathulate-lincar, laxly 
hairy, up to 1 cm by 1 mm. Pedicels rather densely 
lepidote and shortly subpatently greyish pubescent, 
7-10(-12) by c. 0.5 mm. Cw/va: minute, membra- 
nous, wavy, irregularly fimbriate, c. 2.5 mm 0. 
Corolla salver-shaped, white below, rose coloured 
towards the limb, tube cylindric, straight, 3-3.3 
cm, base rather distinctly 5-gibbous and c. 2.5 mm 
0. slightly dilated towards the mouth, laxly short- 
hairy and caducously lepidote outside (scales 
completely gone at full anthesis), short-hairy in 
the lower half inside, glabrous otherwise, lobes 
slightly spreading, obovate-spathulatc, 1-1.2 by 
0.7-0.8 cm, hairy outside at the base, otherwise 
glabrous, though manifestly ciliolale. Stamens 10, 
slightly exserted; lilaments linear, hairy in the 
lower third, glabrous above; anthers oblong, 
1.8-2 by 0.7 mm. base obtuse. Disk prominently 
10-gibbous. glabrous. Ovary subcylindric. densely 



560 



Flora Malesiana 



[ser. I, vol. 6^ 



hairy and lepidote, 6-7 by 1.6 mm, rather abruptly 
narrowed to the style, the latter almost attaining 
the limb of the corolla, hairy and scaly to nearly 
the top; stigma thick, turbinate-capitate. 

Distr. Malesia: Central Celebes (Masamba 
Distr.: near Limbung), c. 1700 m, rather common 
in secondary forest on stony and peaty soil. One 
collection. Fl. Aug. 

125. Rhododendron radians J. J. S. Bull. Bot. Btzg 
HI, 1 (1920) 403, t. 51; Sleum. Reinwardtia 5 
(1960) 130. 

var. radians. 

Shrub. Branchlets terete, tips laxly pubescent 
and lepidote; internodes 3-12.5 cm. Leaves 5-12 
per pseudowhorl, ovate to ovate-oblong, subsessile, 
apex shortly obtusely attenuate, sometimes nearly 
rounded, base slightly to distinctly cordate, 
coriaceous, glabrous except the very base and the 
midrib above at maturity, laxly lepidote beneath 
(scales ih coarsely dentate or lacerate at the 
marginal zone, each mostly on top of a minute 
epidermal elevation, their ±_ persistent dark centre 
impressed), slightly or not revolute at the base, 
(2-)2.5-5.5 by l-2.2(-2.9) cm, midrib impressed 
above, prominent beneath, nerves 6-8 pairs, 
distinctly anastomosing, slightly impressed above 
and prominent beneath in the oldest leaves, often 
it obscure, reticulation inconspicuous; petiole 
laxly lepidote and pubescent, c. l(-2) by 1-2 mm. 
Umbels 7-20-flowered. Outer perulae ovate- 
acuminate, acute, ± glabrous dorsally, inner ones 
± spathulate, obtuse, rather laxly sericeous 
dorsally and internally, up to 1 by 0.8 cm. Brac- 
teoles linear, patent-hairy, up to 1 .5 cm by 0.5 mm. 
Pedicels rather stoutish, densely greyish-patent- 
pubescent, not lepidote, 7-13 mm. Calyx small, 
oblique, margin wavy and shortly 5-lobed. 
Corolla salver-shaped, white, tube straight- 
cylindric, (5-)5.5-6 cm, equally c. 3 mm wide, 
laxly to very laxly lepidote and epilose outside, 
subdensely to laxly short-hairy to % of the lower 
part inside, glabrous distally, lobes spreading, 
broad-obovate-spathulate, very laxly lepidote out- 
side, glabrous inside, 1-1.3 by Q.1-Q.9 cm. Stamens 
10, slightly exserted from the throat; filaments 
linear, laxly hairy to % of their length, filiform 
and glabrous apically; anthers obovate-oblong, 
base obtuse, c. 1.8 by 1 mm. Disk glabrous at 
the base, but hairy within the upper part. Ovary 
subcylindric, 5 by 1.8 mm, densely ± patently 
hairy (and lepidote, but scales covered or hidden 
by the hairs), rather abruptly narrowed to the 
finally exserted style, which is laxly patent- 
pubescent to the lower ^/g and bears some scales 
in its lower part; stigma turbinate, rather small. 

Distr. Malesia: Central Celebes (Mt Sinadji), 
one collection. 

var. minahasae Sleum. Reinwardtia 5 (1960) 
130. — R. verticillatum {non Low) Koord. Minah. 
(1898) 514; Koord. -Schum. Syst. Verz. 3 (1914) 
100. — R. jasminiflorum {non Hook.) Sarasin, 
Reisen in Celebes 1 (1905) 63. 



Scales more deeply stellate-incised at the 
marginal zone, subdendroid, each on top of a 
more distinct minute epidermal elevation than in 
var. radians. Outer perulae h obtuse, subsericeous 
at the apex or practically glabrous, somewhat 
shining. Style hairy and lepidote up to almost the 
top. 

Distr. Malesia: NE. Celebes (Minahasa, on 
several volcanoes). 

Ecol. In low forest, on volcanic grit or sandy 
soil, 1500-1700 m. 

Vern. Tente-in-talun, Tt., Minahasa. 

Note. The corolla of the type specimen from 
Mt Soputan is given as white and very fragrant by 
the collector, and other collections from that 
region also seem to have had white corollas, as far 
as this can be concluded from the dried material. 
Only Koorders says, that his specimen, also 
collected on Mt Soputan, has dark red corollas, 
but no corollas are preserved in his herbarium. 
Boesveld, the collector of the type specimen of 
var. minahasae, says on the label, that he has found 
"a similar Rhododendron on Mt Klabat with red 
corollas", but I have seen no material from there 
which would agree with that. It remains open 
whether var. minahasae has always white flowers, 
or if, possibly, both white- and red-flowered forms 
exist together. 

126. Rhododendron oreadum Wernh. Trans. Linn. 
Soc. II, Bot. 9 (1916) 98; Sleum. Reinwardtia 5 
(1960) 132. — R. filamentosum Wernh. Trans. 
Linn. Soc. II, Bot. 9 (1916) 97. 

Shrub. Branchlets subangular, rather slender, 
tips subdensely lepidote, glabrescent below. 
Leaves in ± 3-merous pseudowhorls, lanceolate, 
sometimes broadly lanceolate, subinequilateral, 
apex acutely acuminate, base acutely attenuate, 
subcoriaceous, practically glabrous above at 
maturity, rather laxly stellate-lepidote beneath 
(scales thin, irregularly dentate or incised, flat 
centre small, faintly impressed), flat, 8-14 by 
(2.6-)3-4(-5) cm, midrib slightly impressed above, 
prominent beneath, nerves 8-12 pairs, rather dense 
together, curved-anastomosing before the edge, 
obscurely impressed above, raised beneath, veins 
and veinlets subinconspicuous on both faces; 
petioles laxly lepidote, 8-10 by 1.5 mm. Umbels 
compact, 8-12-flowered. Perulae ovate to broadly 
obovate, lepidote outside in the upper half, short- 
hairy otherwise on both faces in the upper, longer 
and appressedly so in the lower part, up to 3 by 
1.8 cm. Bracteoles filiform, laxly hairy, up to 3 cm. 
Pedicels stoutish, lepidote, c. 2 mm. Calyx small, 
obtusely 5-lobed, c. 2.5 mm 0. Corolla salver- 
shaped, probably white when fresh, tube cylindri- 
cal, straight, c. 4.5 cm long, c. 4 mm at the slightly 
swollen base, not much (up to 8 mm) widened 
towards the throat, practically glabrous outside, 
pubescent to nearly the lobes inside, the latter 
broad-elliptic or elliptic-obovate, 1.6-1.8 by 0.9- 
1.3 cm, glabrous. Stamens 10, slightly exceeding 
the corolla tube; filaments linear and densely 
patent-pubescent in the lower half, gradually 
glabrescent above; anthers oblong, base obtuse. 



Jan. 1966] 



Ericaceae (Sleumer) 



561 



c. 4 by 1 mm. Disk short-hairy in the upper part. 
Ovary cylindric, 8-10 by 1.7 mm. densely covered 
with longish, — appressed, forwardly directed 
hairs and some scales between, gradually attenuate 
to the style, which is but laxly covered with the 
type of hairs found on the ovary to 2/3 its lower 
part, glabrous above, c. 4 cm at full anthesis; 
stigma shortly broad-obconical, indistinctly 5- 
lobed. 

D i s t r . Malesia : W. New Guinea (Mt Carstensz), 
1095-1495 m. 

127. Rhododendron cinerascens Sleum. Reinward- 
tia 5 (I960) 130; Blumea 12 (1963) 103. 

Shrub, c. 1 m. Branchlets terete, rather slender, 
ashy, tips laxly lepidote; internodes 2.5-10 cm. 
Leaves 4-6 per pseudowhorl, lanceolate-oblong to 
obovate-oblong, apex shortly obtusely acuminate, 
rarely subrotundate, base cuneate into the petiole, 
coriaceous, subdensely lepidote initially on both 
faces (scales thin, flat, irregularly incised at the 
marginal zone, small centre impressed), finally 
glabrescent above, — persistently lepidote beneath, 
entire, flat, 4-5.5 by (1.2-)1.5-2(-2.3) cm, midrib 
shallowly impressed above, prominent beneath 
especially in the lower part, nerves 6-8 pairs, 
rather inconspicuous on both faces; petiole — 
flattened, grooved above, lepidote initially, 6-10 
by c. 1.3 mm. Umbels 5-6(-7)-flowered. Ferulae 
ovate to obovate-spathulate, laxly lepidote 
dorsally and sericeous apically, glabrous inter- 
nally, ciliate and set with scales at the margin, up 
to 1.5 by 0.7 cm. Bracteoles filiform to sub- 
spathulate-linear, 1-1.5 cm by c. 1 mm. Pedicels 
rather slender, densely lepidote, laxly and very 
shortly hairy, 9-20 by 0.5 mm. Calyx small, very 
oblique, in form of a thin, wavy, irregularly 
5-dentate rim, one or the other teeth elongate to 
2-4 mm. Corolla salver-shaped, white, 5-6 cm 
long in all, tube cylindric, straight or slightly 
curved, (3.7-)4-5 cm, c. 2.5 mm at base, gradu- 
ally dilated to 3-3.5 mm at the throat, rather 
densely lepidote, not hairy outside, laxly pubes- 
cent inside in the lower third only, lobes spreading, 
spathulate-obovate, c. 1 by 0.6-0.7 cm, h laxly 
lepidote outside, glabrous inside. Stamens 10, 
well cxscrted from the corolla tube; filaments 
filiform, hairy in the lower half only; anthers 
oblong, base obtuse, c. 2.5 mm. Disk very shortly 
hairy at the upper margin. Ovary elongate- 
cylindrical, c. 5 by 1.2 mm, subdensely lepidote 
and subapprcsscd-hairy, gradually narrowed to 
the style, the latter exscrtcd for c. I cm, lepidote 
and subpatcntly hairy for 3 cm above the base, 
otherwise lepidote only to nearly the top; stigma 
shortly obconical-capitate. 

Distr. Malesia: W. New Guinea (WisscI Lakes 
area), in secondary forest on Sphagnum, 1750 m. 
Two collections. I'l. April-May. 

Vcrn. Waradc, Kapauku. 

128. Khr»dodendron brachypodarium .Si.kum. Blumea 
12 {\'}(>l) 103; Jahrb. Khoilod. Ges. Bremen 
(1963) 23. in text, f. 27. - li>{. 25. 

Hpiphytic or terrestrial shrub, l-2(-3) m. 



Branchlets densely lepidote, greyish pruinose 
when dry; upper internodes (2-)3-10 cm by 2-3 
mm. Leaves ± reflexed, in 4(-5)-merous pseudo- 
whorls at the 2-3 (-4) upper nodes, elliptic or 
more often obovate-elliptic, rarely obovate, apex 
broadly attenuate, obtuse or rounded, base more 
broadly or narrowly cuneate, coriaceous, sub- 
densely lepidote on both faces initially, rather 
early glabrescent above with age, more tardily 
so beneath (scales flattish, marginal zone angular 
or dentate to various degree, centre rather small, 
slightly impressed), entire, hardly revolute, 
(4-)5-7(-8) by 2-3.5(-4) cm, midrib thick and 
grooved in the lower, becoming flat and slightly 
impressed or even evanescent distally above, as 
thick as the petiole at the base, gradually decres- 
cent upwards beneath, nerves 7-10 pairs, very 
slightly depressed or obscure above, indistinctly 
curved-anastomosing and a little raised beneath, 
reticulation subinconspicuous; petiole very densely 
lepidote initially, tardily glabrescent, 6-10(-12) 
by 1.5-2 mm. Umbels (8-)10-12(-14)-flowered. 
Ferulae greenish when fresh, outer ones ovate, 
firm, lepidote at apex and margin, fimbriate, inner 
ones much thinner, broadly spathulate, up to 2 by 
1.3 cm, very laxly hairy in the lower part on both 
faces, otherwise glabrous. Bracteoles filiform, 
laxly hairy, up to 1.8 cm. Fedicels densely stellate- 
lepidote, 4-8 by 0.5 mm. Calyx much oblique, 
rim-like, obscurely lobed, lepidote dorsally, c. 
2 mm across. Corolla narrowly tubular, pure white, 
4-5.5(-6) cm long in all, ± densely stellate- 
lepidote outside, short-hairy at the tube inside, 
tube straight, 3.5-4(-5) cm by 2-2.5 mm, hardly 
dilated distally, lobes spreading or a little reflexed, 
obovate-oblong, c. 1 by 0.4-0.5 cm. Stamens 10, 
exceeding the corolla tube for c. 1 cm; filaments 
narrow-linear and densely hairy for the lower 3^, 
more slender and glabrous for the rest; anthers 
narrow-oblong, (3-)4 mm, base obtuse. Disk 
slightly prominent, laxly short-hairy. Ovary 
narrowly cylindric, c. 8 by 1.3 mm, whitish or 
yellowish tomentose (the hairs covering the scales), 
tapering to the style, the latter slender, exserted 
from the throat for c. 5 mm, densely hairy below, 
more laxly hairy and densely lepidote above in the 
lower 2/3 part, densely and exclusively lepidote 
for the rest; stigma obcuneatc, glabrous. 

Distr. Malesia: W. New Guinea (Vogclkop 
Peninsula: Tanirau, Tohkiri, and Nettoti Range). 

Ecol. Both in Castanopsis and Nothofagus 
forest, also in mossy crest or riverine forest, 
1200-2000 m. r/. Oct.-Dec. 

Note. The flowers become fragrant towards the 
evening. 

129. Rhododendron goodcnoughii Slf.um. Rein- 
wardtia 5 (i^JW)) 131. 

Shrub. Branchlets 3-6 mm o. terete, pruinose 
in the older parts, glabrous, tips sparsely lepidote; 
internodes 4-10 cm. Leaves in 4-6-mcrous pseudo- 
whorls, elliptic to obovate-elliptic, apex very 
shortly acuminate, sometimes nearly rounded, 
base attenuate to the petiole and a little winged, 
coriaceous, lirm. glabrous above at malurily, 



562 



Flora Malesiana 



[ser. I, vol. 6^ 




Fig. 25. Rhododendron brachypodarium Sleum., Tohkiri Mts East, 1400 m, NW. New Guinea (Van 

RoYEN & Sleumer 7065) (Sleumer, 1961). 



Jan. 1966] 



Ericaceae (Sleumer) 



563 



laxly lepidote beneath (scales small, irregularly 
dentate or incised at the margin, centre very 
small impressed), 7-1 1 by 4-6 cm, midrib but 
slightly impressed above, very prominent below, 
less so towards the apex of the lamina beneath, 
nerves 7-9 pairs, the lowest 2 from the base of 
the blade, all at an acute angle from the midrib, 
rather straight below, curved and obscurely 
anastomosing distally, faintly impressed above, 
prominent beneath, — obscure sometimes towards 
the edge, no reticulation; petiole flattened, 
grooved above, lepidote initially, a little rough, 
7-15 by 2-3 mm. Umbels — 10-flowered. Ferulae 
ovate, rather blunt, outer ones rather densely 
lepidote and very shortly hairy dorsally, longer 
and appressed-pubescent internally and lepidote 
at the apex only, up to 2.5 by 1.8 cm. Bracteoles 
narrow-linear, nearly glabrous, up to 1.5 cm by 
c. 0.5 mm. Pedicels subdensely lepidote, not hairy 
at all, 3-5 (-7) by c. 1 mm. Calyx small, very 
oblique, indistinctly lobed, c. 2.5 mm 0. Corolla 
salver-shaped, probably white when fresh, 5.7-7 
cm long in all, tube cylindric, slightly curved, 
4.5-5(-5.5) cm, c. 4 mm at the 5-pouched 
base, c. 6 mm at the throat, subglabrous outside 
below, very laxly lepidote in the upper part, 
subdensely hairy downwards internally, glabres- 
cent distally, lobes spreading, obovate-rounded, 
1.2-1.5 by 1-1.3 cm, very laxly lepidote outside at 
the base, otherwise glabrous. Stamens 10, ± 
equalling the corolla in length; filaments linear, 
subdensely hairy in the lower 2/3, nearly filiform 
and glabrous above; anthers oblong, 5-6 by 1 mm, 
each theca bearing a very short basal apiculus. 
Disk subglabrous. Ovary elongate-cylindric, 
densely clothed with yellowish, appressed hairs 
and scales (the latter covered or hidden by the 
hairs), 7-8 by c. 2 mm, gradually narrowed to the 
rather stoutish style, the latter c. 5 cm, densely 
clad with scales and forward directed hairs in the 
lower 2/3 only, glabrous for the ultimate 1-1.5 cm; 
stigma thick-obconical. 

Distr. Malesia: SE. New Guinea (Mt Good- 
enough), one collection. 

130. Rhododendron pneumonanthum Sleum. Rein- 
wardtia 5 (I960) 132. — /?. jasminiflorum {nan 
Hook.) Ridl. J. Str. Br. R. As. Soc. n. 63 (1912) 
60; Merr. Sarawak Mus. J. 3 (1928) 541. 

Epiphytic shrub. Branchlcts terete, very densely 
rusty stcllate-lepidotc at the upper 2-3 internodes, 
glabrcscent and a little rough below; internodes 
2.5-6 (rarely up to 10) cm. Leaves 4-6 per pseudo- 
whorl, elliptic-oblong or -obovate, apex blunt, 
nearly rounded, sometimes very shortly retuse, 
base rounded or mostly slightly cordate, sub- 
sessile, coriaceous, rather densely stcllate-lepidote 
initially on both faces (scales rathcrcarlycaducous, 
each on top of a minute epidermal elevation, often 
almost cup-like, margin deeply incised, centre 
faintly impressed), glabrous above at maturity, 
lepidote beneath for a longer time, finally glabrcs- 
cent and rather smooth to the touch, slightly 
revolutc at the margin, (4.5-)5-7.5(-9) by (2-) 
2.5-4(-5) cm, midrib distinctly sunk above, very 



prominent beneath in its lower part, less so to- 
wards the apex of the lamina, nerves in 6-8 pairs, 
rather inconspicuous on both faces, no reticula- 
tion; petiole densely lepidote, 1-2 (rarely up to 3) 
by 1.5-2 mm. Umbels (6-)8-15-flowered. Ferulae 
ovate to obovate-spathulate, obtuse, laxly lepidote 
dorsally, up to 1 by 0.7 cm. Bracteoles linear, 
glabrous, up to 1.5 cm. Pedicels rather stoutish, 
densely stellate-lepidote, furthermore clad with 
very short hairs, 1 .5-2 cm. Calyx minute, obscurely 
wavy or lobulate, hardly 2.5 mm 0. Corolla salver- 
shaped, white or suffused with rose or light 
violet, but faintly scented, tube cylindric, straight, 
3-3.5 cm long, shortly 5-gibbous and c. 3.5 imn 
at the base, somewhat swollen in the lower half 
(c. 4.5 mm 0), narrowed to c. 2.5 mm at the throat, 
i densely lepidote (scales ± caducous), not 
pubescent outside, laxly hairy in the lower third, 
glabrous otherwise internally, lobes spreading, 
broad-obovate, 1-1.4 by 0.6-0.9 cm, sparsely 
lepidote outside at the base and along the midline, 
glabrous inside. Stamens 10, shortly exserted; 
filaments linear, laxly hairy above the base only; 
anthers oblong, truncate at each end, 2.2-2.5 mm. 
Disk glabrous below, short-pubescent above. 
Ovary elongate-cylindric, densely lepidote and 
subdensely very shortly patent-pubescent, c. 6 
by 1.8 mm, gradually narrowed to the style, the 
latter somewhat exserted, rather densely or laxly 
lepidote and hairy in the lower half, glabrous 
above, c. 3 cm; stigma capitate. 

Distr. Malesia: Borneo (Sarawak and SE. 
Borneo). 

Ecol. In montane (mossy) forest, 1700-2400 m, 
once said to have been found at only 35 m. Fl. 
May, Oct. -Nov. 

131. Rhododendron orbiculatum Ridl. J. Str. Br. 
R. As. Soc. /(. 63 (1912) 60; Merr. En. Born. 
(1921) 462; Sleum. Reinwardtia 5 (1960) 206; 
Blumea 11 (1961) 127; ibid. 12 (1963) 112. — 
R. suaveolens Sleum. Bot. Jahrb. 71 (1940) 147; 
Reinwardtia 5 (1960) 133. 

Treelet or shrub, sometimes epiphytic, 0.3-3 m. 
Branchlets terete or subangular, tips laxly lepidote, 
older parts glabrous; internodes 2-6.5(-8) cm. 
Leaves 3-4(-5) per pseudowhorl, or 2 leaves nearly 
opposite, subsessile, orbicular, broad-elliptic, or 
subovate-tosuborbicular-elliptic, apex ± rounded 
or very broadly obtusely attenuate, apiculate by a 
thick, a little protruding gland, base rounded or 
mostly slightly to manifestly cordate, coriaceous, 
firm, at maturity glabrous above, laxly lepidote 
beneath (scales small, thin, margin 1: deeply 
substellatc-incised or -lobed, centre dark, :t 
impressed), flat, (2-)4-lO(-12) by (2-)4-6(-9)cm, 
midrib flat or impressed in the lower part only 
above, slightly raised or mostly i flat beneath, 
nerves (5-)7-8 pairs, slightly curved and rather 
indistinctly inarching before the edge, a little 
raised, though often sunk in the middle, above, ± 
obscure beneath, reticulation obsolete; petiole 
1-2 by (2-)3-4 mm. Umbels (4-)5-7(-9)-flowcred. 
Outer pcrulae ovate-acuminate, laxly lepidote 
dorsally, inner ones larger, ovate to spathulatc, 



564 



Flora Malesiana 



[ser. I, vol. 6* 



blunt, glabrous, up to 2 by 1 cm. Bracteoles 
linear to filiform, glabrous, up to 1.5 cm. Pedicels 
rather stoutish, all over densely clothed with 
dbpatent greyish whitish hairs, lepidote only just 
below the calyx, (1-) 1.5-2 (sometimes up to 3.5) 
cm. Calyx small, margin, thin, laxly lepidote and 
may be pubescent outside, very shortly obtusely 
5-lobed. Corolla ± salver-shaped, white or pale 
pink, scented, (5.5-)6-6.5 cm long in all, tube 
± cylindric, 3.5-4 cm long, at the base 5-pouched 
and 4-5 mm 0, often gradually slightly narrower 
towards the lobes, laxly lepidote or glabrous out- 
side, very laxly hairy in the lower half inside, lobes 
spreading, broadly obovate to suborbicular, 
glabrous, (l-)1.5-2 (rarely up to 2.5) cm 0. 
Stamens 10, hardly exserted; filaments linear, 
laxly hairy above the base, subfiliform and 
glabrous in the upper half; anthers oblong, apex 
truncate, each cell with a short basal apiculus, 
2.5-3.5 by 1 mm. Disk subglabrous below, 
puberulous above. Ovary elongate-conical, 6-8 
by 1.5-2 mm, densely set with short spreading 
hairs and scales, the latter + covered by the hairs 
(more discernible towards the apex of the ovary), 
gradually tapering to the style, the latter laxly 
hairy except the uppermost part, scarcely or not 
lepidote, widely exserted; stigma subturbinate, 
deeply 5-lobed. Capsule narrow-fusiform, 5-ribbed, 
laxly lepidote and hairy, c. 4 by 4 mm, pedicel 
3-5 cm. 

Distr. Malesia: Borneo (Sarawak; Brunei; 
North Borneo: Mt Kinabalu). 

Ecol. On Mt Kinabalu often epiphytic and 
with larger leaves in ridge forest between 1220 
and 2135 m. In N. Sarawak and Brunei rarely 
epiphytic, generally terrestrial and found on 
extreme hill 'kerangas\ i.e. low scrubby vegetation 
on bare sandstone rocks, eroding into white sand, 
or in elfin woodland on rugged hill crest, 800- 
1750 m, locally abundant. Fl. Jan. -Dec. 

Vern. Kajii ala, M. 

132. Rhododendron edanoi Merr. & Quisumb. 
Philip. J. Sc. 83 (1953) 333; Sleum. Reinwardtia 
5 (1960) 134. 

Shrub, c. 1 m. Branchlets terete, very densely 
and ± scurfy rusty-stellate-lepidote at the upper- 
most internodes, glabrescent below; internodes 
2-5.5 cm. Leaves 4-6 in a pseudowhorl, oblong- 
elliptic to obovate-oblong, apex very broadly 
attenuate, obtuse to rounded, base broadly 
attenuate to the petiole, coriaceous, glabrous and 
somewhat lustrous above when old, densely 
lepidote beneath (scales rather early caducous, 
each on top of a minute epidermal elevation, which 
remains for a rather long time underneath, margin 
deeply irregularly incised, centre very small), 
subrevolute, (2.5-)3-5.5 by 1.5-2.5 cm, midrib 
impressed above, strongly raised beneath, nerves 
4—6 pairs, rather straight-ascending, anastomosing, 
slightly impressed above, faintly prominent or 
obscure beneath, reticulation obsolete; petiole 
semiterete, grooved above, lepidote, 5-7 by c. 
1.5 mm. Umbels i 7-flowered. Perulae ovate to 
elliptic or spathulate, densely lepidote outside in 



the upper third, glabrous elsewhere, up to 1.5 by 
0.7 cm. Bracteoles linear to subspathulate, up to 
1.5 cm. Pedicels rather slender, densely stellate- 
lepidote and laxly short-hairy, 9-12(-15) mm. 
Calyx disk-like, small, suboblique, membranous, 
margin narrow and obscurely 5-lobed. Corolla 
salver-shaped, white, tube cylindric, straight or 
but slightly curved, 2-2.5 cm in all, 2.5-3 mm 0, 
densely stellate-lepidote outside, laxly hairy inside 
in the lower third, glabrous upwards, lobes 
spreading, elliptic to broadly obovate, 0.8-1 by 
0.6-0.7 cm, outside laxly lepidote, inside glabrous. 
Stamens 10, slightly exserted; filaments linear, 
subdensely hairy above the base, glabrous for the 
rest; anthers oblong, base obtuse, 3 by 1 mm. 
Disk densely hairy. Ovary cylindric, 5 by 1.8 mm, 
densely lepidote and short-hairy, rather abruptly 
attenuate to the style, the latter c. 2.5 cm, pubescent 
and lepidote in the lower third, laxly pubescent 
in the middle, glabrous in the upper half; stigma 
obconical-subcapitate. 

Distr. Malesia: Philippines (Palawan: Mt 
Mantalingahan, Brooke's Point), in moss forest 
at 1725 m. Fl. May. 

133. Rhododendron loranthiflorum Sleum. Notizbl. 
Berl.-Dahl. 12 (1935) 485; Reinwardtia 5 (1960) 
134; Blumea 11 U961) 120. 

Epiphytic, much branched shrub. Branchlets 
smooth, older parts terete and pruinose, tips 
subangular or flattened and i densely rusty- 
lepidote; internodes 1.5-12 cm. Leaves in 3-5- 
merous pseudowhorls, sometimes only 2 opposite 
on the same node, obovate, apex broadly obtusely 
attenuate or mostly ± rounded, base cuneate, 
subcoriaceous, lepidote on both faces initially, 
glabrous above with age, subdensely and ± 
persistently lepidote beneath (scales thin, irregu- 
larly and rather shallowly dentate or lacerate at 
the margin, centre slightly impressed), subrevolute, 
3.5-5(-6) by 2-2.5 cm, midrib faintly impressed 
above, prominent beneath especially in the lower 
part, nerves 4-5 pairs, indistinct on both faces, 
reticulation obscure; petiole rather slender, 5-7 
mm. Umbels 4-5-flowered. Perulae ovate, thin, 
very laxly lepidote or glabrous, up to 7 by 4 mm. 
Bracteoles filiform, glabrous, 6-7 mm. Pedicels 
slender, laxly lepidote, not hairy at all, c. 10 mm. 
Calyx minute, disk-shaped, indistinctly 5-lobed. 
Corolla salver-shaped, white, tube cylindric, some- 
what curved in anthesis, 2-2.5 cm by 2.5-3 mm, 
densely lepidote outside (the scales thickish, sub- 
entire), laxly papillose hairy in the lower half 
inside, lobes ovate-oblong, spreading in a nearly 
right angle, lepidote outside, glabrous inside, c. 
5 by 2.5 mm. Stamens 10; filaments filiform, shortly 
hairy in the lower 2/3, nearly as long as the corolla 
tube; anthers oblong, base obtuse, 2 mm. Disk 
low, glabrous. Ovarvsubcylindric, slightly 5-ribbed, 
densely lepidote (scales thickish, subentire) and 
very shortly subpatently hairy, gradually tapering 
to the style, the latter c. 2.2 cm, lepidote and hairy 
in the lower part, exclusively hairy above respec- 
tively glabrous below the thick subcapitate stigma. 
Capsule cylindric, slender, curved, attenuate 



Jan. 1966] 



Ericaceae (Sleumer) 



565 



apically, 5-ribbed, ± densely rusty-lepidote and 
short-hairy, 4—4.5 cm by c. 3 mm, on fruit-pedicel 
c. 1.5 cm. 

Distr. Melanesia: New Britain (headwaters of 
Matabunu Creek, epiphyte in high rain-forest, 
180 m) and in the Solomon Is. (Bougainville, 
near Koniguru, Buin, in rain-forest at c. 1000 m). 

134. Rhododendron retrorsipilum Sleum. Blumea 
11 (1961) 120; ibid. 12 (1963) 103. 

Epiphytic shrub. Branchlets rather slender, c. 
2 mm 0, subdensely lepidote initially, very early 
glabrescent and greyish corticate; upper 3-4 inter- 
nodes (l-)2-4 cm, lower ones hardly longer. 
Leaves in ± 3-merous pseudowhorls at the upper 
1 or 2 nodes, elliptic, apex and base broadly 
obtusely attenuate or more often almost rounded, 
coriaceous, subdensely lepidote initially on both 
faces, earlier glabrescent above than beneath, 
not rough to the touch (scales subdendroid, 
each on top of a minute or very minute, finally ± 
evanescent epidermal tubercle, small, marginal 
zone deeply stellate-incised, centre minute, 
deepened and protracted into a shorter or longer 
foot), hardly revolute at the very edge, 4-6.5 by 
2.5-4.2 cm, midrib very narrowly impressed above, 
as thick as the petiole and much raised in the 
basal part beneath, gradually narrowed and less 
prominent upwards and longitudinally striate be- 
neath, nerves c. 5 pairs, spreading, rather straight, 
faintly or not impressed above, a little raised 
beneath, no reticulation; petiole 6-8(-10) by 1-1.5 
mm. Umbels — 6-flowered, glabrous in all outer 
parts. Floriferous buds ovoid-oblongoid, 2.3 by 
0.8 cm, outer perulae ovate, hardly 4 mm, inner 
ones gradually larger, up to 1.5 cm, subdensely 
set with sessile scales in the upper dorsal part, 
fimbriate. Pedicels rather slender, finely short- 
pubescent, 5-8 mm. Bracteoles linear, up to 1 cm. 
Calyx suboblique, disk-shaped, c. 2.5 mm 0, 
irregularly 5-lobed, lobes broadly deltoid, sub- 
acute, rather long whitish fimbriate, ± reflexed 
at anthesis. Corolla salver-shaped, white, tube 
2-2.3 by c. 0.3 cm, subdensely set with retrorse 
white hairs to nearly the mouth inside, lobes 
narrow-spathulate, ::: horizontally spreading, 
8-10 by 3-4 mm. Stamens 10, very unequal, the 
shortest a little, the longest (3.5 cm) much exsert- 
ed from the corolla tube; filaments filiform, 
densely clothed with white retrorse hairs almost 
to the top; anthers narrow-oblong, 3 by c. 0.7 mm; 
cells subapiculate-protractcd at base, obtuse. 
Disk not much prominent, glabrous at the very 
base, hairy above. Ovary elongate-cylindric, 5-6 
by f . 1.5 mm, densely set with apprcsscd, forward 
directed hairs and with scales (which are almost 
hidden by the hairs), gradually narrowed to the 
style, the latter slender and a little longer than the 
corolla tube, densely hairy below, more laxly 
so and more distinctly lepidote upwards, glabrous 
but for the ultimate part below the capitate 5-lobcd 
■tigma. 

Distr. Malesia: E. New Guinea (Morobe 
Distr.: range above Vfarkham Point near Lae), 
at 600-900 m. H. Jan. -Feb. 



135. Rhododendron subpacificum Sleum. Blumea 
12 (1963) 105. 

Epiphytic shrub. Branchlets slender; internodes 
(l-)2-6 cm. Leaves in 3-4-merous pseudowhorls 
at the upper 2-3 nodes, oblong-obovate, apex 
broadly attenuate to rounded, base cuneate to the 
petiole, subcoriaceous, glabrous above, laxly 
lepidote beneath (scales small, flat, thin, margin 
angular or dented to various degree, centre small, 
slightly depressed), hardly revolute, (3.5-)4— 5.5 
by 1.5-2.3 cm, midrib narrowly impressed above, 
prominent above, prominent beneath, nerves 3-4 
pairs, curved-inarching, rather obscure above, a 
little raised beneath, no reticulation; petiole (2-) 
3 (-4) by 1 mm. Umbels 3-flowered. Bracteoles 
linear, glabrous, up to 6 mm. Pedicels slender, 
laxly caducously lepidote, glabrescent, or already 
glabrous in flowering time, 1-1.2 cm. Calyx 
patellar, shortly 5-lobed and wavy, reflexed, c. 
2 mm across. Corolla salver-shaped, white, thin, 
scented, c. 4 cm long in all, tube straight, c. 
3.5 cm by 3 mm, laxly substellate-lepidote out- 
side, papillose-hairy in the lower third inside, 
lobes almost horizontally spreading, oblong- 
obovate, laxly lepidote outside, 6-8 by c. 4 mm. 
Stamens 10, subequalling the corolla; filaments 
filiform, papillose-hairy in the lower half, other- 
wise glabrous; anthers oblong, base obtuse, 1.5 
mm. Disk glabrous below, otherwise densely 
hairy. Ovary cylindric, c. 6 by 1.5 mm, subdensely 
set with forward directed hairs and scales, tapering 
to the slender style, which is lepidote and hairy for 
the lowest 5 mm, and exclusively set with roundish 
scales almost to the slightly thickened stigma. 

Distr. Melanesia: Solomon Is. (New Georgia 
group, Vangunu I.). 

Ecol. On narrow ridge with stunted forest on 
crater rim at c. 600 m. Fl. Dec. 

136. Rhododendron jasminiflorum Hook. Bot. Mag. 
(1850) t. 4524; MiQ. Fl. Ind. Bat. 2 (1859) 1059; 
Clarke in Hook. /. Fl. Br. Ind. 3 (1882) 463; 
Henslow, J. R. Hort. Soc. 13 (1891) 252, 262, f. 
38, f. 42 (fol.), f. 43c (fl.); Ridl. J. Str. Br. R. As. 
Soc. n. 23 (1891) 145; K. & G. J. As. Soc. Beng. 
74, ii (1905) 77; Koord. Exk. Fl. Java 3 (1923) 7; 
Ridl. J. Fed. Mai. St. Mus. 7 (1916) 45; Millais, 
Rhodod. (1917) 195; Ridl. Fl. Mai. Pen. 2 (1923) 
218; BuRK. & HoLTT. Gard. Bull. S.S. 3 (1923) 57; 
Henders. J. Mai. Br. R. As. Soc. 5 (1927) 256; 
Mai. Nat. J. 6 (1950) 251, f. 244; Sleum. Rein- 
wardtia 5 (1960) 128. — R. elongatum Bl. Fl. 
Jav. PI. indd. (1863-83) t. 7 B, f. 1; Steen. Bull. 
Jard. Bot. Btzg 111, 17 (1948) 388. — Azalea 
jasminiflora (Hook.) O. Ktze, Rev. Gen. PI. 2 
(1891) 387. 

key TO THE varieties 

1. Corolla tube laxly to densely short-hairy. 

2. Leaves rather narrow to ovatc-oblong, 
cuneate to subcordatc at the base. Branchlets 
pubcrulous 3. var. hcusscri 

2. Leaves wider, elliptic to broadly subovatc- 
clliptic, rounded to cordate at the base. 



566 



Flora Malesiana 



[ser. I, vol. 6* 



Branchlets not puberulous 2. var. punctatum 
1. Corolla epilose outside, except some scattered 

hairs which may occur at the very base. 
3. Corolla hairy or papillose inside in the lower 

half only 4. var. copelandii 

3. Corolla tube hairy inside for the full length. 

4. Leaves obovate-elliptic to broadly elliptic, 

sometimes suborbicular, base ± rounded to 

cordate 1. var. jasminiflorum 

4. Leaves oblong, base broadly cuneate. 

5. var. oblongifolium 

1. var. jasminiflorum. 

Small, mostly epiphytic shrub, sometimes ter- 
restrial, also scrambling, up to 2.5 m. Branchlets 
subterete, upper internodes 1.5-4 cm, ± densely 
substellate-lepidote, lower ones up to 9 cm and 
glabrescent. Leaves in 3-5-merous pseudowhorls 
at the upper nodes, broadly obovate-elliptic or 
elliptic, sometimes suborbicular, mostly widest in 
the middle, apex broadly obtusely (rarely sub- 
acutely) acuminate, or rounded, base truncate- 
rounded or mostly slightly cordate, subsessile, 
coriaceous, dark green and a little glossy above, 
paler and rather dull beneath, initially lepidote 
but readily glabrescent above, rather laxly sub- 
stellate-lepidote all over the undersurface, which 
may be rather rough to the touch (scales some- 
times subdendroid, with an irregularly and ± 
deeply dentate or incised margin, normally each 
on top of a minute and blunt epidermal elevation, 
faintly impressed in it with the centre), margin 
± revolute in dry specimens, 2.5-5(-6) by (1-) 
1.5-3.2(-3.6) cm, midrib slightly impressed above, 
strongly raised beneath, nerves c. 5 pairs, rather 
straight, anastomosing, mostly ± inconspicuous 
on both faces or faintly prominent only, reticula- 
tion obscure; petiole mostly but 2-3, occasionally 
up to 7 mm long, L5-2 mm 0, lepidote. Umbels 
(3-)5-8(occasionally up to 20)-flowered. Ferulae 
reddish when fresh, ovate to obovate-spathulate, 
obtuse, glabrous or rarely laxly lepidote near the 
top, irregularly ciliate, ± shining, up to 1 by 0.6 
cm. Bracteoles linear, glabrous, up to 1 cm by 0.6 
mm. Pedicels rather slender, densely and patently 
short-hairy and substellate-lepidote, 8-11 mm. 
Calyx obscurely obtusely 5-lobed, c. 2 mm 0. 
Corolla salver-shaped, very fragrant like Narcissus, 
pure white, or flushed pink below the throat, tube 
cylindric, ± straight, base slightly 5-gibbous, 
laxly lepidote (but not hairy) outside, laxly hairy 
from the base to near the mouth inside, 3.5^ 
(-4.3) cm by 3^ mm, lobes spreading horizontally, 
broadly obovate-spathulate, very laxly or not 
lepidote outside, glabrous inside, 0.8-1. 2(-l. 5) by 
0.6-1 (-1.1) cm. Stamens 10, somewhat exserted 
from the throat, unequal; filaments linear, laxly 
hairy to nearly the top; anthers broadly oblong, 
truncate at both ends, c. 1.8 by 1 mm. Disk 
densely covered with short greyish hairs. Ovary 
oblong-cylindric, densely lepidote and short-hairy, 
c. 5 by 1.8 mm, rather abruptly tapering to the 
style, the latter nearly as long as the corolla, laxly 
lepidote in the lower third, and laxly hairy to 
nearly the top; stigma short-obconical or sub- 



capitate. Capsule narrow-ellipsoid, short attenuate 
at both ends, obtusely 5-ribbed, laxly short-hairy 
and lepidote, 1.8-2.8 cm by 4—5 mm. 

Distr. Malesia: Malay Peninsula (Johore: 
Mt Ophir). 

Ecol. In ridge forest, low scrub, and epiphytic, 
or sometimes terrestrial on rocks in more open 
places, 1160-1525 m. Fl. Dec. 

2. var. punctatum Ridl. J. Fed. Mai. St. Mus. 4 
(1909) 42; Sleum. Reinwardtia 5 (1960) 129. — 
R. jasminiflorum {non Hook. s. str.) Ridl. J. Fed. 
Mai. St. Mus. 6 (1915) 157; ibid. 7 (1916) 45. — 
R. jasminiflorum Hook. var. maculatum Ridl. Fl. 
Mai. Pen. 2 (1923) 218; Burk. & Holtt. Card. 
Bull. S.S. 3 (1925) 390. 

Shrub or treelet, up to 3 m. Leaves elliptic to 
broadly subovate-elliptic, base rounded to sub- 
cordate. Corolla white or flushed pink, with i 
conspicuous bright pink spots in the throat, tube 
laxly short-hairy all over outside, moreover 
whether or not laxly lepidote. Perulae ± sericeous 
dorsally. Pedicels more slender and longer 
(mostly 1.5-2.5 cm), making the umbel less com- 
pact than in var. jasminiflorum. 

Distr. Malesia: Malay Peninsula (Kedah: 
Kedah Peak, in the Main Range from Perak/ 
Pahang: Cameron Highlands, down to Selangor: 
Mt Moyang, also in the Taiping Hills and on 
Mt Tahan). 

Ecol. In dwarf mossy forest, damp ridges, open 
patch of swamps, 855-1830 m, rather common 
in the Cameron Highlands. Fl. predominantly 
Oct. -Dec, fr. March. 

3. var. heusseri (J. J. S.) Sleum. Reinwardtia 5 
(1960) 129. — R. retusum [non (Bl.) Benn.] Steen. 
Arch. Hydrobiol. Suppl. 11 (1932) 318, f. 21, 
22 {phot. veg.). — R. lon^iflorum Lindl. var. 
heusseri J. J. S. Bull. Jard. Bot. Btzg III, 13 (1935) 
447. 

Much branched shrublet, 50 cm to I m, with 
both lepidote and puberulous branchlets. Leaves 
in 5-13-merous pseudowhorls, rather narrow- 
oblong to ovate-oblong, base cuneate to sub- 
cordate, puberulous above along the midrib. 
Corolla sparsely lepidote and subdensely ± 
patently short-pubescent outside, with an immense 
variety in colours of a pure white, ivory, a pale 
sulphur and an intense red, and many grades of 
salmon between pale yellow and red, furthermore 
all grades between red and white, as known from 
the cultivated Azaleas. 

Distr. Malesia: Sumatra (Atjeh: Gajolands; 
East Coast: Sibolangit and near Pematang Siantar; 
around Lake Toba, especially Samosir Peninsula). 

Ecol. In open rocky places on old, dry volcanic 
soil, apparently always near solfataras, capable 
to stand the gasses of SO2, 750-1525 m. Fl. 
predominantly in June-July. 

Vern. Bunga batu, M. 

Note. Possibly a hybrid between R. jasmini- 
florum and R. longiftorum, but certainly nearer to 
the first, with which it agrees considerably in the 
leaves and the flowers, the influence of the latter 



Jan. 1966] 



Ericaceae (Sleumer) 



567 



mainly visible in the red and yellow colour which 
breaks through to various degree. It is stated 
that the specimens with pure colours are 50 cm 
high in the Samosir Peninsula, whereas the 
specimens with intermediate colours are more 
luxuriant and reach 1 m in height. 

4. var. copelandii (,Merr.) Sleum. Reinwardtia 
5 (1960) 130. — R. copelandii Merr. Publ. Gov. 
Lab. Philip, n. 29 (1905) 42; Philip. J. Sc. 3 (1908) 
Bot. 381; Elm. Leafl. Philip. Bot. 3 (1911) 1105; 
MiLLAis, Rhodod. (1917) 148; ibid. ser. 2 (1924) 
117; Merr. En. Philip. 3 (1923) 243; Copel. /. 
Philip. J. Sc. 40 (1929) 158, t. 7, f. 1-5. 

Similar to var. jasminifiorum, but corolla hairy 
inside in the lower half only. 

D i s t r . Malesia : Philippines (Mindanao : summit 
of Mt Apo), in thickets, 2350-3100 m. Fl. March- 
Nov. 



Vern. Malagas, malambaho. Bag. 

5. \ar. oblongifolium Sleum. Reinwardtia 5 (1960) 
130. — R. verticillatum {non Low ex Lindl.) 
Becc. Malesia 1 (1878) 204. — R. jasminifiorum 
{non Hook. s. str.) Merr. En. Born. (1921) 461. 

Leaves oblong, apex obtuse, base obtuse or 
truncate-rounded, 4-7.5 by 1-2.8 cm; petiole 4-6, 
rarely up to 7 mm. Corolla white, laxly lepidote 
and epilose outside, laxly hairy inside the entire 
length of the tube. Perulae glabrous or laxly 
lepidote, ^ shining. 

Distr. Malesia: Malay Peninsula (P. Tioman, 
Pahang; Mt Blumut, in Johore) and Borneo 
(Sarawak). 

Ecol. Normally in mountain forest, once found 
almost at sea-level in the delta of the Sarawak 
River, rather common on the summit of Mt 
Santubong. 



7. Subsection Euvireya 

Copel./. Philip. J. Sc. 40 (1929) 137, 159; Sleum. Bot. Jahrb. 74 (1949) 541; Reinwardtia 5 
(1960) 134, s. lat. — Fig. 27-^5. 

Scales 'star-shaped' to various degree, mostly (rather) small, already distant from each other 
on the young leaves, the thin marginal zone i irregularly dented or lobed, often early dissolute 
(the remaining centre then giving the impression of a disk-shaped scale). 




Fig. 26. Distribution of Rhododendron subsect. Euvireya. 
Distr. S. Indochina, Australia (N. Queensland), and Malesia. Fig. 26. 



KEY TO THE SERIES 

I. Leaves very small in general. 0.3-1 (rarely in the same specimen partly up to 2) by 0.2-0.6 (rarely 

up to 0.8) cm. Spp. 137-147 1. ser. Linnacoidca 

I. Leaves all or at least for the greater part (in the same specimen) more than 1.6 cm long. 
2. Compact dwarf shrub, forming tussocks or mats. Leaves lincar-lanccolatc to oblanccolatc. Ilowers 
solitary (rarely in twos); pedicel stout, erect, distinctly emerging beyond the foliage; (lower 

nodding. .S>. 148 2. ir/-. Saxifragoldca 

2. Lrcct shrubs or trcclcts. 



568 Flora Malesiana [ser, I, vol. 6* 

3. Leaves linear to narrow-lanceolate, in general (at least partly) not exceeding 7 mm in width. 
4. Leaves 20 or more in each pseudowhorl, 1-1.5 mm wide. Sp. 149 3. ser. Taxifolia 

4. Leaves opposite or 3-8(-15) in each pseudowhorl, 3-7(-ll) mm wide. Spp. 150-153. 

4. ser. Stenophylla 

3. Leaves generally much wider, not linear or narrow-lanceolate. 

5. Stamens 5. Sp. 154 5. ser. Citrina 

5. Stamens (7-)10(-14). 

6. Leaves medium-sized, i.e. 1-4 (sometimes partly in the same specimens up to 6.5) cm long. 

Spp. 155-196 6. ser. Buxifoiia 

6. Leaves generally larger, i.e. all or at least for the greater part in the same specimen longer than 

4 cm. Spp. 197-276 7. ser. Javanica 

1. Series Linnaeoidea 

Sleum. Reinwardtia 5 (1960) 134. — Rhododendron sect. Linnaeopsis Schltr, Bot. Jahrb. 55 (1917) 
144, f. 2. — Rhododendron subsect. Linnaeopsis (Schltr) Sleum. ibid. lA (1949) 541. — Fig. 27 — 29. 

Leaves very small, 3-10 (rarely in part up to 20) by 2-6(-7, rarely up to 8) mm. Slender, often dwarf, 
terrestrial or epiphytic shrubs. 

Distr. Malesia: limited to New Guinea. 

KEY TO THE SPECIES 

1. Ovary all over densely (predominantly) hairy and less densely lepidote. 

2. Style at anthesis relatively short, i.e. ± as long as the ovary, glabrous or hairy at the base only. 
Corolla campanulate to broadly funnelform-campanulate, deeply 5-lobed, exclusively lepidote out- 
side. Very slender creeping or decumbent shrublet 137. R. anagalliflorum 

2. Style at anthesis at least two times as long as the ovary, hairy in the lower yi to % , glabrous for the 
rest. Corolla ± tubular, laxly lepidote and laxly to subdensely hairy outside. Erect (sometimes 
scandent) shrub up to 2 m 138. R. womersleyi 

1. Ovary exclusively and densely lepidote. 

3. Style shortly ± patently hairy nearly to the top 139. R. gracUentinn 

3. Style with scales or some subpatent hairs at the base only, otherwise glabrous, or glabrous all over. 

4. Style at anthesis as long or shorter than the ovary, ± stoutish. 

5. Compact dwarf cushion-like shrublet with short, densely foliate branchlets. Corolla 1-1.2 cm 

long in all 140. R. caespitosum 

5. Slender erect or creeping shrublets, divaricate, with slender elongate branchlets. Corolla (1.5-) 
1.8-2.3 cm long in all. 
6. Leaves opposite or in 3-5(-7)-merous pseudowhorls at the upper 1-3 nodes, entire or nearly so, 

elliptic or subovate- or obovate-elliptic, ± apiculate 141. R. microphyllum 

6. Leaves alternate, scattered along the upper part of the branchlets, conspicuously crenate or 

crenulate, ovate to ovate-lanceolate or -elliptic, subacuminate 142. R. pusillum 

4. Style at anthesis about double as long as or longer than the ovary, ± slender. 
7. Leaves opposite or in 3-5(-7)-merous pseudowhorls at the upper 1-3 nodes. Corolla up to 1.8 cm 
in all. 
8. Corolla 1.5-1.8 cm long. Leaves narrow-elliptic or elliptic, 5-8 by 4-6 mm, apex subacute in 
general 143. R. muscicola 

8. Corolla 1-1.2 cm long. Leaves broad-elliptic or subobovate-elliptic, 6-10(-12) by (3-)4-7(-8) 
mm, apex obtuse-rounded 144. R. parvulum 

7. Leaves scattered along the upper part of the branchlets. Corolla 2.2-3 cm long in all. 

9. Leaves 3-6(-7) by 2.5-4 mm, very slightly or not recurved at the margin . . 145. R. oxycoccoides 
9. Leaves 6-10 by 4—7 mm, ± distinctly recurved at the somewhat thickened or cartilagineous 

margin. 

10. Leaves ovate, apex acuminate, subacute 146. R. disterigmoides 

10. Leaves ovate-elliptic to ovate-oblong, apex ± obtuse in general, but not rarely shortly apiculate. 

147. R. coelorum 

137. Rhododendron anagalliflorum Wernh. Trans. or subsessile stellate-incised scales in the youngest, 
Linn. Soc. II, Bot. 9 (1916) 94; Sleum. Reinward- ± verruculose in the older glabrescent parts; 
tia 5 (1960) 136; Blumea 11 (1961) 121; ibid. 12 internodes 0.5-2.5 cm. Leaves opposite or in 3-5- 
(1963) 106; ibid. 12 (1964) 340. — R. linnaeoides merous pseudowhorls at the upper 3-4 nodes, 
Schltr, Bot. Jahrb. 55 (1917) 144, f. 2. elliptic or oblong- or subobovate-elliptic, apex sub- 
Dwarf epiphytic shrublet, 5-10 (rarely up to 30) acute or rarely more obtuse, base ± broadly 
cm, much divaricately branched, prostrate or cuneate, subcoriaceous, initially lepidote on both 
creeping. Branchlets very slender, 0.5-1 nrni 0, sides, glabrous above at maturity, laxly persistently 
densely clothed with shortly stipitate ('dendroid') lepidote beneath (scales light brown, flat or 



Jan. 1966] 



Ericaceae (Sleumer) 



569 





195S 




Fig. 27. Rhododendron womersleyi Sleum. a. Branchlet with flowers and 3 single leaves, x 2/3. — R. 
^/i/er/fmojWe^ Sleum. 6. Branchlet with flowers, x 2/3,0. leaf, x 2. — R.oxycoccoides Sleum. d. Branchlet 
with flower bud, x 2/3. — /?. parvM/wm Sleum. e. Branchlet with flower, x 2/3, and a single leaf, x 1 1/3- — 
R. caespitosum Sleum./. Branchlet with immature fruit, x 2/3 {a NGF 891 8 Womersley, b-c Brass 9022, 
d Brass 12184, e Eyma 5203, / Brass 9039). 



slightly concave, marginal zone obtusely stellate- 
incised, centre small, not impressed), not or only 
slightly revolute, 3-7 (occasionally up to 14) by 
2.5-3.5 (occasionally up to 6) mm, midrib and 
nerves inconspicuous; petiole slender, 1-1.5 mm. 
Flowers solitary. Outer perulae narrow-ovate- 
acuminate, inner ones oblong-acuminate, apex 
± subulate-attenuate, very finely puberulous or 
glabrous dorsally, fimbriate by stalked scales, 
3-7 by 1-3 mm, often persistent for a rather long 
time. Bracteoles linear, 4-5 mm. Pedicels very 
slender, rather densely clothed with dendroid 
stellate scales, otherwise whether or not finely 
pubescent, (4-)7-12(-15) mm. Calyx obliquely 
cup-shaped, shortly obtusely 5-lobed, c. 2 mm 0. 
Corolla campanulate or broadly funnelform- 
campanulate, thin, 5-lobed halfway or almost so, 
red or pinkish, 12-18(-22) mm long and across 
the limb (20-30 mm when fresh), base 3-6 mm 
0, laxly to subdensely stellate-lepidote outside, 
the upper part of the lobes excepted, glabrous 
inside, lobes obovate-oblong, erecto-patent or 
± expanded, obtuse. Stamens 10, unequal, the 
longest equalling : half the corolla, or slightly 
longer; filaments linear, glabrous; anthers broad- 
oblong, f. 0.7 mm. Disk glabrous. Ovary broadly 
subovoid-cylindric, densely clad withshort, spread- 
ing hairs, less densely with scales between these 
hairs, 2-2.5 mm by 1.5 mm, slightly 5-ribbcd; 
style rather abrupt with the ovary, thickish 
initially, glabrous or sparsely hairy at the very 
base, 2.5-3 mm; stigma thick-capitate, 5-gibbous. 
Capsule fusiform, (1. 2-) 1. 5 -2 cm by 3 4 mm, 
short-hairy and Icpidotc. Seeds 3-4 mm including 
the tails. 

Distr. Malesla: New Guinea (Main Range 



from Mt Carstensz to Mt Victoria). 

Ecol. In montane forest, open grassland or 
alpine thickets, epiphytic on moss clumps on 
trees, or trailing over fallen logs, 1190-3000 
(-3540) m. Fl. Jan.-Dec. 

138. Rhododendron womersleyi Sleum. Reinwardtia 
5 (1960) 136; Blumea 11 (1961) 121; ibid. 12 
(1963) 106; ibid. 12 (1964) 340. — R. aff'. in- 
conspicuum Stonor, Rhodod. Yearbook 6 (1951) 
51, f. 52. — Fig. 27a. 

Stiff", terrestrial, sometimes scrambling shrub, 
very rarely epiphytic on mossy trunk, 0.5-2 m; 
branches loose, erect, rather stout, divaricately 
branched chiefly distally. Branchlets 1-2 rrun 0, 
te-rete, tips densely clothed with brownish, stellate, 
shortly stalked scales, glabrescent below and ver- 
ruculose, early covered with greyish cork. Leaves 
rather densely set along the upper part of the bran- 
chlets, ovate to ovate-elliptic, sometimes subova- 
te-elliptic-oblong, apex short-acuminate, subacute, 
base broad-attenuate to rounded, coriaceous, 
dark green above, paler beneath, subdensely lepi- 
dote on both faces initially, glabrescent and often 
a little shining above at maturity, persistently 
subdensely or more laxly lepidote beneath 
(scales rather large, flat, marginal zone irregularly 
obtusely stellate-incised, centre hardly impressed), 
margin distinctly crcnulatc by the impressed 
scales, not or very slightly revolute, 5-8(-IO, rarely 
up to 15) by 4-6(-7, rarely up to 10) mm, n\idrib 
t distinctly sunk, faintly prominent beneath, 
nerves subobscurc; petiole stoutish, l-1.5(-2) mm. 
Flowers nodding, solitary or in twos, rarely in 
threes. Outer perulae ovate-acuminate, shortly 
subulate or apiculate, inner ones narrower. 



570 



Flora Malesiana 



[ser. I, vol. 6* 



otherwise very similar, glabrous dorsally, margin 
densely shortly subglandular-fimbriate and 
caducously lepidote, 6-14 by 3-6(-8) mm. 
Bracteoles filiform, c. 5 mm. Pedicels stoutish, 
rather densely clothed with very shortly stalked 
stellate scales and more laxly with very short 
papillose hairs, 8-12(-15) mm at anthesis, slightly 
longer in fruit. Calyx disk-shaped, spreading or 
reflexed, c. 2.5 mm 0, lobes broadly triangular, 
obtuse, lepidote and finely hairy, c. 1 mm. Corolla 
nodding, tubular, slightly dilated at the limb, 
bright or deep red to scarlet, 2-2.3(-2.6, rarely -3) 
cm long in all, 3-3.5 cm across the limb, laxly 
stellate-lepidote outside at the tube and the lower 
part of the lobes, moreover sparsely to subdensely 
hairy there (or certainly so in the lower half of the 
corolla tube), glabrous inside, tube subcylindric, 
1.3-1.7(-2) cm long, 3-4(-6) mm at base, 
widened to 5-10 mm at throat, lobes erecto- 
patent, obovate subrotundate, 4-5 (-6) mm 0. 
Stamens 10, slightly longer than the corolla tube, 
slightly unequal; filaments filiform, glabrous; 
anthers oblong, c. 1.5 mm. Disk prominent, 
hairy. Ovary subcylindric, densely patently and 
shortly whitish hairy and subdensely lepidote, c. 
4 by 1.5 mm, gradually tapering to the slender 
style, which nearly equals the corolla in length 
and is set with spreading fine hairs in the lower 
2/3 to %, glabrous above; stigma club-shaped 
or subcapitate, shortly 5-gibbous. Capsule fusi- 
form, hairy, greyish purplish when almost ripe, 
1.5-2 cm by 3-4 mm. Seeds 4 mm including the 
tails. 

Distr. Malesia: E. New Guinea (Eastern, 
Western and Southern Highlands; Morobe Distr.: 
Rawlinson Range). 

Ecol. Mostly in open mossy forest or forest 
edge with grassland, more rarely in grassland 
above the timber-line, fairly common locally, 
3200-3960 m. Fl. Feb.-Sept. Hybridizes with 191. 
R. atropurpureum on Mt Wilhelm. 

Vern. Balimbar, Enga: Poio, kiWhorli, kobu, 
Mendi. 

139. Rhododendron gracilentum F. v. M. Trans. 
R. Soc. Vict. n.s. 1, 2 (1889) 22; Sleum. Reinward- 
tia 5 (1960) 137; Blumea 11 (1961) 121; Jahrb. 
Rhodod. Ges. Bremen (1963) f. 12 — Fig. 28. 
Erect, ascending or subprostrate small shrub 
with divaricate branchlets, (10-) 20^0 (-60) cm 
high, generally terrestrial, rarely epiphytic. 
Branchlets 1-1.5 mm 0, tips subdensely clad 
with rusty, stellate, shortly stipitate scales, else- 
where glabrescent and verruculose; internodes 
0.5-1.5 (rarely up to 4) cm. Leaves opposite or in 
3-7-merous pseudowhorls at the upper 3-4 nodes, 
often somewhat recurved, subovate- or elliptic- 
lanceolate or lanceolate, apex short-acuminate, 
mostly subacute, base acutely attenuate, coriaceous, 
olivaceous-green above, dilutely so beneath, glossy 
and rugulose above, laxly lepidote on both faces 
initially, glabrescent above with age, persistently 
lepidote beneath (scales irregularly and rather 
obtusely stellate-incised at the margin, centre not 
impressed), a little wavy and subcrenulate by the 



± impressed scales on the very margin in dry 
specimens, not or only slightly recurved, (6-)8-12 
(sometimes up to 20) by 2-5(-8) mm, midrib 
subinconspicuous above, slightly obtusely raised 
beneath, nerves obsolete; petiole rather slender, 
1-1.5 mm. Flowers solitary, or rarely in pairs, 
pendent. Flower buds narrow-ovoid, c. 6 by 2 mm. 
Ferulae ovate to oblong-ovate, acute or with a 
short subulate acumen, glabrous dorsally, margin 
shortly gland-fringed and set with early caducous 
scales, up to 6 by 3 mm. Bracteoles filiform, 
glabrous, c. 5 mm. Pedicels slender, densely 
clothed with shortly stalked substeliate scales and 
less densely with very short hairs, 6-10 mm. 
Calyx membranous, deeply 5-lobed, stellate- 
lepidote outside, lobes deltoid, subacute, c. 1 mm. 
Corolla cylindric, red, rose red or dark pink, laxly 




\ 



"Ih 



Fig. 28. Rhododendron gracilentum F. v. M. 

a. Habit, x 2/3, b. stamen, x 2, c. ovary and style, 

X 2, d. submature fruit, x 2/3 (a Womersley.s.«., 

b-c Lam 7747, d NGF 9529 White). 

covered with deeply stellate pale brown scales 
outside, glabrous inside, 1.8-2.4 cm long in all, 
tube 1.2-1.5 cm long, c. 3 mm at base, c. 5 mm 
at throat, lobes erecto-patent or finally much 
spreading, obovate-suborbicular, 4-6 mm 0. 
Stamens 10, unequal, the longest nearly as long 
as the corolla; filaments narrowly linear, glabrous; 
anthers subquadrate, truncate, 1.5 by 1.2 mm, 
connective swollen and darker in colour. Disk 
prominent, glabrous. Ovary oblong-cylindric, 3-4 
by 1-1.5 rrun, densely and exclusively lepidote, 
gradually attenuate to the slender style, which 
nearly attains the corolla tube in length and is 
set with fine, subpatent, short, whitish hairs 
nearly up to the clavate-capitate stigma. Capsule 
erect, oblique, oblong, a little curved, glabrescent, 
1.5-2.2 by 0.3 cm, on slender pedicel c. 1.5 cm. 



Jan. 1966] 



Ericaceae (Sleumer) 



571 



Distr. Malesia: E. New Guinea (Central 
Distr.: Mt Musgrave; Morobe Distr.: Edie Creek 
above Wau). 

Ecol. Terrestrial on open low vegetation on 
rather dry slopes or clayey ground of old gold 
mine workings, also epiphytic in Nothofagus 
forest or in summit growth. 2000-2745 m. Fl. May- 
Oct. 

140. Rhododendron caespitosum Sleum. Rein- 
wardtia 5 (1960) 137. — Fig. 27f, 29. 

Dwarf, epiphytic shrublet. up to 10 cm, much 
branched; compact, forming cushions; branches 
short, robust, rooting. Branchlets very short, slen- 
der, substipitate-stellate-Iepidote in the younger 
parts, giabrescent and minutely verrucose below. 
Leaves dense, scattered or subopposite, elliptic, 
ape.x short-acuminate, subacute or subobtuse, 
base attenuate, coriaceous, laxly to subdensely 
lepidote initially on both faces (scales rufescent, 
marginal zone irregularly stellate-incised, centre 
faintly impressed), giabrescent with age, smooth 
and somewhat lustrous above, dull and persistently 




Fig. 29. Rhododendron caexpifosum Sleum. forming 
a cushion on stem of a grassland trccfcrn (Cyathea 
lomenlosissima Copn..). Lake Mabbcma, .1225 m, 
New Guinea (Brass 9039) (Brass. 1938). 



lepidote beneath, entire or subcrenulate at the 
slightly cartilaginous margin by impressed 
scales, 3-6 by 2-3 mm, midrib and nerves obscure; 
petiole rather slender, c. 1 mm. Flowers solitary. 
Ferulae ovate-acuminate, apiculate, glabrous 
dorsally, subglandular-fimbriate and set with 
early caducous scales along the margin, up to 
6 by 3 mm. Bracteoles filiform, c. 4 mm. Pedicels 
rather stoutish, densely clothed with rusty very 
shortly stalked stellate scales, but no hairs, 5-6 
mm. Calyx short-cupular, obscurely 5-lobed, 
very densely lepidote, c. 2 mm 0. Corolla tubular, 
of a delicate pink, 1-1.2 cm long in all, subdensely 
stellate-lepidote outside, the upper part of the 
lobes excepted, glabrous inside, tube 8-9 mm 
long, c. 2.5 mm at base, slightly widened apically, 
lobes erecto-patent, obovate-orbicular, 3-4 mm 
0. Stamens 10, nearly as long as the corolla tube; 
filaments filiform, glabrous; anthers broad- 
oblong, 0.5 mm. Disk glabrous. Ovary thickly 
obovoid-cylindric, very densely stellate-lepidote 
and set with minute papillae between the scales, 
c. 3 by 1.5 mm; style abrupt with the ovary, 
slenderly columnar, glabrous, c. 2 mm; stigma 
thick-capitate. Capsule obovoid-cylindric, densely 
stellate-lepidote, 8-10 by c. 3.5 mm, the 5 valves 
rather thin, not twisted. Seeds with rather short 
tails on both ends, c. 2 by 0.3 mm. 

Distr. Malesia: W. New Guinea (Mt Wilhei- 
mina-Lake Habbema vicinity). 

Ecol. Forming cushions on trunk of tree-ferns 
in alpine grassland, 2400-3225 m. Fl. Aug.-Sept. 

141. Rhododendron microphyllum J. J. S. Bull. Jard. 
Bot. Btzg 11, 8 (1912) 51; Nova Guinea 12 (1914) 
131, t. 29 c; Sleum. Reinwardtia 5 (1960) 138. — 
R. minimifolium Wernh. Trans. Linn. Soc. II, 
Bot. 9 (1916) 98, p.p. 

Epiphytic slender shrublet, 20-30 cm, rooting, 
divaricately branched. Branchlets f. 1 mm 0, terete, 
tips -t caducously lepidote, scales rufous, shortly 
stipitate, stellate, older parts giabrescent, verru- 
culose for a while, early becoming smooth and 
greyish corticate; internodes (0.6-)l-5 cm. Leaves 
in 3-6-merous pseudowhorls, broadly elliptic to 
obovate-clliptic, sometimes suborbicular, apex 
short-acuminate, subacute, or rather obtuse, not 
rarely apiculate. base broad-attenuate, coriaceous, 
laxly lepidote on both sides initially (scales 
irregularly stcllatc-laccrate, flat, thoscof Ihcunder- 
surface somewhat impressed), t giabrescent above 
with age, persistently lepidote underneath, margin 
narrowly cartilaginous or thickened, subcrenulate 
by impressed scales, (3-)5-9 by (2-)3-6 mm, 
midrib slightly impressed above, somewhat raised 
beneath, or obsolete as are the nerves; petiole 
stoutish. 1-2 (rarely up to 3) mm. f/ourrv solitary, 
nodding. Flower buds narrow-ovoid, acuminate. 
Outer perulae subulate or ovate-lanccolale, 
densely lepidote outside, inner ones broader, 
ovate-acuminate, glabrous dorsally or nearly so, 
shortly glandular-fimbriate by a row of early 
caducous scales, up to 6 by 3 mm. Bracteoles 
filiform, c. 5 mm. Pedicels slender, densely clothed 
with rufous, dendroid, 1. deeply sl.ir-shaped 



572 



Flora Malesiana 



[ser. I, vol. 6* 



scales, no hairs, (6-) 10-1 7 mm. Calyx obliquely 
disk-shaped, slightly obtusely 5-lobed, densely 
stellate-lepidote outside, c. 3 mm 0. Corolla broad- 
ly tubular-subcampanulate, purplish red or pale 
red, laxly to subdensely rufous-stellate-lepidote 
outside, the upper part of the lobes excepted, 
glabrous inside, 1.8-2.1 cm long in all at full 
anthesis, tube 1.2-1.5 cm long, c. 4 mm at base, 
gradually widened to 6-8 mm at throat, lobes 
± orbicular and spreading, 5-7 mm 0. Stamens 10, 
unequal, the longest nearly as long as the corolla 
tube; filaments linear, glabrous; anthers sub- 
quadrate, c. 1.5 rrun. Disk glabrous. Ovary 
cylindric-conical, densely stellate-lepidote, c. 4 by 
2 mm; style rather abrupt with the ovary, short- 
columnar, thickish, glabrous or with some sparse 
subpatent hairs at the very base, c. 3.5 mm; 
stigma subcapitate, distinctly 5-lobed. Immature 
capsule fusiform, 1.5 cm by 2.5 mm, densely 
lepidote. 

Distr. Malesia: W. New Guinea (Main Range 
from Mt Carstenz to Mt Goliath and Mt Wilhel- 
mina-Lake Habbema). 

Ecol. On moss-covered trees in subalpine 
forest, locally common. Fl. March-April, Aug. 

142. Rhododendron pusillum J. J. S. Med. Rijks- 
herb. n. 25 (1915) 1; Nova Guinea 12 (1917) 498; 
ibid. 12 (1918) t. 193; Sleum. Reinwardtia 5 
(1960) 138. — R. candidapiculatum Wernh. Trans. 
Linn. Soc. II, Bot. 9 (1916) 96. 

Dwarfed shrublet of weak, ascendent habit, 
6-25(-50) cm; main branches few, stout, 3-4 
mm 0. Branchlets coming irregularly from the 
branches, much more slender than these (1-1.5 
mm 0), often curved-ascending, densely sub- 
imbricately leaved at least in their upper 
part, tips ± densely clothed with sessile and 
substipitate, rusty-stellate scales, older parts ver- 
ruculose, early covered with cork. Leaves ovate- 
lanceolate or ovate-elliptic, apex short-acuminate, 
subacute, mostly with a paler apiculus, base 
attenuate to the petiole or sometimes nearly 
rounded, often complicate in the upper part, 
stiff-coriaceous, laxly lepidote on both faces 
initially (scales irregularly stellate-incised, flattish, 
not or slightly impressed), glabrescent above with 
age, ± persistently lepidote beneath, margin 
mostly distinctly crenate or crenulate at least in the 
upper half, flat, somewhat cartilaginous and 
paler than the proper lamina, 4-8(-10) by 2-3(-4, 
occasionally up to 6) mm, midrib slightly sunk 
above, respectively raised beneath, often obscure 
as are the nerves; petiole rather stoutish, 1-2 mm. 
Flowers l-2(-3), nodding from an erect pedicel. 
Flower buds ovoid, 10-12 by c. 5 mm. Ferulae 
membranous, ovate-acuminate, shortly subulate or 
apiculate, outer ones lepidote dorsally, inner ones 
completely glabrous, up to 12 by 6 mm. Bracteoles 
linear, up to 8 mm. Pedicels rather stoutish, 
densely set with stipitate stellate scales, no hairs, 
(6-) 10-1 5 (-20) mm. Calyx disk-shaped, very 
shortly subacutely 5-dentate, densely lepidote, 
2-3 mm 0. Corolla tubular-subcampanulate, 
suboblique, initially red, changing to orange at 



later stages, (1.5-)1. 8-2.3 cm long in all, very 
laxly lepidote in the lower part or mostly glabrous 
outside, glabrous inside, tube 1-1.6 cm, base 
rounded, 3-4 mm 0, dilated gradually to 5-6 mm 
at throat, lobes erecto-patent or somewhat 
spreading, obliquely obovate-suborbicular, slight- 
ly retuse, 5-7 (-9) mm 0. Stamens 10, unequal, ± 
half as long as the corolla; filaments linear, gla- 
brous; anthers suborbicular, said to be blackish, 
c. 1.2 mm. Disk glabrous. Ovary ovoid-conical, 
densely stellate-lepidote, 3-4 by 2 mm; style sub- 
abrupt with the ovary, columnar, glabrous, 2-3 
(sometimes up to 4) mm; stigma capitate, 5- 
lobed. 

Distr. Malesia: New Guinea (Main Range 
from Mt Carstensz to the Oranje Mts, frequent 
on Mt Wilhelmina and vicinity). 

Ecol. Alpine grassland, on exposed sandstone, 
in shallow sandy soil or on peat soil at base of 
rocks, also in shrubby borders of subalpine forest, 
3200-3900 m, locally common. Fl. Sept.-Feb. 

143. Rhododendron muscicola J. J. S. Nova Guinea 
18 (1936) 93, t. 19, 1 ; Sleum. Reinwardtia 5 (1960) 
138. 

Weak, divaricately branched shrub. Branchlets 
slender, 1-1.5 mm 0, terete, tips densely set with 
stellate, shortly stalked scales, older parts glabres- 
cent, verruculose; internodes 0.9-3 cm. Leaves in 
5-7-merous pseudowhorls at the upper 2-3 nodes, 
normally narrow-elliptic or elliptic, occasionally 
subobovate-elliptic, apex short-acuminate, sub- 
acute, or rarely more obtuse, base attenuate into 
the petiole, very coriaceous, laxly lepidote on both 
faces when young (scales irregularly and rather 
shallowly obtusely stellate-incised), glabrescent 
above at maturity, persistently and ± impressedly 
lepidote beneath, margin cartilaginous and some- 
what paler than the lamina, ± entire, flat, 5-8 by 
4-6 mm, midrib and nerves obscure; petiole 
stoutish, 1-1.5 mm. Flowers solitary. Flower buds 
ovoid, c. 7 by 3 mm. Ferulae ovate to ovate- 
oblong or obovate, subulate-acuminate, minutely 
sericeous and lepidote dorsally, ciliate, up to 8 by 
3.5 mm. Bracteoles filiform, glabrous, c. 5 mm. 
Fedicels very slender, densely substipitate-stellate- 
lepidote and laxly papillose-hairy, 8-13 mm. 
Calyx obliquely cup-shaped, 5-lobulate, c. 2.5 mm 
0. Corolla narrow funnelform-campanulate, red, 
1.5-1.8 cm long in all, laxly lepidote outside, the 
upper part of the lobes excepted, glabrous inside, 
tube 1-1.2 cm long, 2.5-3 mm at base, gradually 
dilated to c. 5 mm at throat, lobes irregularly 
suborbicular, 4-5 mm 0. Stamens 10, unequal, 
somewhat exserted from the corolla tube; filaments 
linear, a little dilated at the base, glabrous; 
anthers subobovate, 1-1.5 mm. Disk set with some 
hairs at upper margin, not much prominent. 
Ovary oblong-conical, longitudinally 5-ribbed, c. 
3.5 by 1.5 mm, densely stellate-lepidote, gradually 
tapering to the style, the latter rather slender, 
nearly as long as the corolla, with some sparse, 
spreading, yellowish hairs at base, elsewhere 
glabrous, or glabrous all over; stigma subclavate- 
capitate. 



Jan. 1966] 



Ericaceae (Sleumer) 



573 



Distr. Malesia: W. New Guinea (Nassau Mts, 
W. side), once found at 2600 m in moss-grown 
forest. Fl. Oct. 

144. Rhododendron parvulum Sleum. Reinwardtia 
5 (1960) 139. — Fig. 27e. 

Small weak shrublet, only some branchlets 
known, these slender, 0.6-1 mm a, tips densely 
clad with dark, rusty, substipitate scales, glabres- 
scent elsewhere, minutely tuberculate and rough 
to the touch; internodes 1.2-4 cm. Leaves m 3-4- 
merous pseudowhorls or opposite at the upper 
2-3 nodes, broad-elliptic to subobovate-elliptic, 
ape.x rounded-obtuse, base very broadly attenuate 
or rounded, coriaceous, very young ones laxly 
to subdensely lepidote on both faces, glabrescent 
above with age, lepidote for a longer time beneath 
(scales irregularly stellate-lacerate, flat, slightly 
impressed), dull, margin paler than the blade, 
subcartilaginous, entire or subcrenulate by the 
impressed scales, 6-10(-12) by (3-)4-7(-8) mm, 
midrib and nerves obsolete; petiole rather slender, 
lepidote, c. 1 mm. Flowers solitary or in twos. 
Ferulae narrow ovate-acuminate, manifestly and 
subdensely appressed-hairy, glandular-fimbriate 
and initially set with a row of (early caducous) 
scales along the margin, up to 6 by 2.5 mm. 
Pedicels very slender, densely clothed with sub- 
stipitate, deeply stellate-incised scales and very 
short slender subpatent hairs, 7-10 mm at anthesis. 
Calyx disk-shaped, 2 mm 0, deeply obtusely 5- 
lobed, lepidote dorsally. Corolla tubular, 1-1.2 
cm long in all, subdensely stellate-lepidote outside, 
upper half of lobes excepted, glabrous inside, 
tube c. 8 mm, c. 2.5 mm at base, widened to 3.5 
mm at throat, lobes m spreading, suborbicular, 
c. 4 mm 0. Stamens 10, unequal, the longest 
reaching r: halfway the corolla lobes; filaments 
filiform, glabrous; anthers obovate, hardly 1 mm. 
Disk glabrous. Ovary obconical-cylindric, very 
densely stellate-lepidote, c. 2 by 1.2 mm, gradually 
attenuate to the rather slender style (c. 3.5 mm); 
stigma clavate-subcapitate. 

Distr. Malesia: W. New Guinea (Wissel Lakes 
region: summit of Mt Mutaro). One collection. 
Fl. Sept. 

145. Rhododendron oxycoccoides Sleum. Rein- 
wardtia 5 (1960) 139. — Fig. 27d. 

Epiphytic shrublet. Branchlets much divaricate, 
0.5-1 mm 0, tips densely rusty-stellate-lepidote, 
older parts glabrescent and smooth. Leaves laxly 
scattered along the distal part of the branchlets, 
ovate, short-acuminate, subacute or more obtuse, 
base broadly rounded, coriaceous or subcoria- 
ceous, initially lepidote on both sides (scales 
irregularly stellate-incised, not or faintly immer- 
sed), early glabrous above, ! persistently lepidote 
beneath, margin a little thickened, slightly or not 
recurved, 3-6(-7) by 2.5-4 mm, midrib and 
nerves invisible; petiole rather slender, 0.5-1 mm. 
Flowers solitary. Pcrulae membranous, ovate- 
lanccolatc, inner ones long subuiutc-acuminatc, 
quite glabrous and shining, up to 1.5 by 0.5 cm. 
Pedicels rather slender, 4-6 mm, densely stellate- 



lepidote. Calyx disk-shaped, limb very short, 
wavy, lepidote outside, c. 2.5 mm 0. Corolla 
tubular, slightly zygomorphous, pink, thin, 2.5- 
2.7 cm long in all, glabrous in- and outside, tube 
1.8-2 cm, c. 3 mm at base, widened to c. 5 mm 
at throat, lobes erecto-patent, suborbicular, 
5-7 mm 0. Stamens 10, unequal, the longest 
reaching halfway the corolla lobes; filaments 
linear, glabrous; anthers subquadrate, 1-1.2 mm. 
Disk prominent, glabrous. Ovary subcylindric, 
densely lepidote, 3.5 by 2 mm, gradually tapering 
to the rather slender style (c. 1.4 cm); stigma 
capitate. 

Distr. Malesia: New Guinea (Bernhard Camp, 
Idenburg R., c. 139°-3°30'), once found in 
stunted, mossy forest on an exposed summit, 
2100 m. f/. Jan. 

146. Rhododendron disterigmoides Sleum. Rein- 
wardtia 5 (I960) 140. — Fig. 27b-c. 

Terrestrial shrub, 30-80 cm, with robust erect 
branches. Branchlets rigid, rather stout, 1.5-2 mm 
0, tips clad with rufescent substipitate stellate 
scales, older parts glabrous, not or not much 
rugose, early covered with greyish cork. Leaves 
subdensely scattered on the youngest part of the 
branchlets, ovate, apex acuminate, apiculate but 
rather blunt, base rounded, thick-coriaceous, 
persistently laxly lepidote beneath (scales rather 
small, irregularly stellate-incised, faintly or not 
impressed), margin thickened and a little paler 
than the lamina, distinctly revolute, crenulate by 
impressed scales, 6-10 by 4-6 mm, midrib obtusely 
prominent beneath, obsolete above as are the 
nerves; petiole thickish, c. 1 mm. Umbels 2-4- 
flowered. Pedicels rather slender, densely sub- 
stipitate-stellate-lepidote, not hairy, 8-12 mm. 
Calyx very shortly cupular, shortly obtusely 5- 
lobed, densely lepidote dorsally, c. 2.5 mm 0. 
Corolla tubular, deep red, 2.2-2.3 cm long in all, 
laxly to subdensely lepidote at the tube outside, 
glabrous inside, tube c. 1.7 cm long, 4 mm 
at base, widened to c. 7 mm at throat. Stamens 10, 
unequal, :}; as long as the corolla or a little 
exserted; filaments linear, glabrous; anthers 
broadly obovate-obiong, c. 1.8 by 1 mm. Disk 
glabrous. Ovary subconical-cylindric, very densely 
lepidote, 4-5 by 2 mm, gradually tapering to the 
glabrous, columnar style (8-9 mm); stigma sub- 
clavate-capitate. 

Distr. Malesia: W. New Guinea (Lake Habbe- 
ma), common in shrubberies on peaty ridges, 
3225 m. One collection. Fl. Aug. 

147. Rhododendron coelorum Wernh. Trans. Linn. 
Soc. II, Bot. 9 (1916) 96; Sleum. Reinwardtia 5 
(1960) 140; Blumca II (1961) 121. 

Apparently decumbent terrestrial shrublet, c. 
15 cm. Branchlets slender, firm, tips covered with 
stipitate stellate scales, older parts glabrescent, 
laxly verruculo.sc, early covered with greyish cork. 
Leaves rather densely scattered along the upper 
branchlets, ovatc-clliptic to ovate-oblong, apex 
shortly obtusely acuminate or apiculate, base 
i rounded, thick-coriaceous, mature ones glub- 



574 



Flora Malesiana 



[ser. I, vol. 6* 



rous, rugulose and a little lustrous above, laxly 
lepidote beneath (scales irregularly obtusely 
stellate-lobed, slightly or not impressed), margin 
thickened, :l: entire, manifestly revolute in dry 
specimens, 6-10 by 4-7 mm, midrib slightly 
impressed above and raised beneath, nerves 
obscure; petiole thickish, 1-1.5 mm. Flowers 1-2 
(-3-4). Pedicels rather slender, subdensely stellate- 
lepidote, epilose, 15-17 mm. Calyx disk-shaped, 
obscurely 5-lobed, lepidote outside, c. 2.5 mm o. 
Corolla tubular, slightly oblique, 2.5-3 cm long 
in all, laxly stellate-lepidote outside at the tube. 



glabrous inside, 5-6 mm at base, widened to c. 
1 cm at throat, lobes erecto-patent, suborbicular, 
5-6 mm 0. Stamens 10, unequal, ± as long as 
the corolla; filaments linear, glabrous; anthers 
suborbicular, 1.5 mm. Disk prominent, glabrous. 
Ovary subcylindric, densely lepidote, c. 6 by 2 mm, 
gradually attenuate to the slender, quite glabrous 
style, which is nearly as long as the corolla; 
stigma obconical-capitate. 

Distr. Malesia: New Guinea (Mt Carstensz), 
at 3200-3810 m. One collection. Fl. Jan. 



2. Series Saxifragoidea 

Sleum. Reinwardtia 5 (1960) 141 . — Fig. 30, 34 d— e. 

Compact shrub, forming cushions or mats, very densely foliate. Leaves linear-lanceolate or oblanceo- 
late. Flowers solitary, or more rarely in twos, nodding from the apex of a robust, upright pedicel which is 
much outstanding above the foliage. 

Distr. Malesia: monotypical in New Guinea. 





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Fig. 30. Rhododendron saxifragoides J. J. S. Mt Sugarloaf summit area, 3650 m, E. New Guinea (HooG- 

LAND & PULLEN 7131) (HOOGLAND, 1960). 



Jan. 1966] 



Ericaceae (Sleumer) 



575 



148. Rhododendron saxifragoides J. J. S. Med. 
Rijksherb. /;. 25 (1915) 3; Nova Guinea 12 (1917) 
503; ibid. 12 (1918) t. 198; Sleum. Reinwardtia 5 
(1960) 141; Blumea 11 (1961) 121. — R. aff. 
fuchsioides Stonor, Rhodod. Yearbook 6 (1951) 
51, f. 51. — Fig. 30, 34 d-e. 

Completely dwarf, rooting shrub, 5-15 (rarely 
up to 30) cm high, forming tussocks, large 
cushions or dense mats. 0.3-1 m o, much divari- 
cately branched, woody parts ~ in the ground. 
Branchlets stout, mostly abbreviate and very 
short, stellate-lepidote at the ± free apex which 
is densely packed with somewhat spreading leaves, 
sometimes more elongate, somewhat decumbent 
and showing 2 internodes. the leaves then in 6-10- 
merous pseudowhorls at the apex and the ultimate 
node. Leaves linear-lanceolate or oblanceolate, 
apex short-acuminate, not rarely apiculate or 
almost mucronate, gradually attenuate to the ± 
flattened base, very coriaceous and stiflF, a little 
shining especially above, practically glabrous 
above at maturity, laxly lepidote beneath (scales 
rather small, marginal zone irregularly and often 
~ obtusely dentate or angular, centre a little 
sunk), margin — revolute in dry specimens, (1.6-) 
2.3-4 (very rarely up to 5.5) by 0.3-0.5(-0.7, very 
rarely up to 0.8) cm, midrib impressed above, 
obtusely prominent beneath, nerves 2-3 steeply 
ascending pairs, very slightly sunk above, or 
often inconspicuous as they are beneath; petiole 
flattened, darker brown than the lamina in dry 
specimens, 3-6(-8) by 1-2 mm. Flowers solitary, 
or rarely in twos, nodding at first, erect in later 
stages. Ferulae ovate, subulate-acuminate, mem- 
branous, lepidote initially, early glabrescent 
dorsally, laxly fimbriate, up to 1.5 by 0.5 cm. 
Bracteoles linear, glabrous, fimbriate, up to 1 cm. 
Pedicels erect, reddish, 1-1.5 mm 0, laxly to sub- 
densely lepidote, sometimes also laxly set with 



short hairs, which go in later stages, (2.5-)4-6(-7) 
cm at anthesis. Calyx oblique, disk-shaped, wavy 
and obtusely 5-lobed (sometimes with longer, 
up to 2 mm, subacute lobes), often fringed with 
subglandular hairs, c. 4 mm 0, lepidote and 
whether or not short-hairy. Corolla obliquely 
tubular below, expanded at the lobes, blood-red, 
pale scarlet or pinkish red, rather densely stellate- 
lepidote outside, the margin of the lobes excepted, 
puberulous at the tube inside, 2.5-3 cm long in 
all, tube broad-cylindric, slightly curved, (1.7-) 
2-2.5 cm long, base c. 5 mm 0, widened to 8-10 
mm at apex, lobes broadly spathulate or suborbicu- 
lar, c. 1 by 0.7-0.9 cm. Stamens 10, subequal, 
attaining about half the length of the corolla 
lobes, purple; filaments linear, laxly patent-hairy 
to the lower third, glabrous for the rest; anthers 
broad-oblong-obovate, c. 2.5 mm. Disk prominent, 
glabrous. Ovary conical, a little constricted at 
base, rather abruptly tapering to the style, 5-6 
by 2.5-3 mm, velutinous and lepidote, hairs 
covering the scales; style as long as or slightly 
shorter than the stamens, stoutish, hairy at the 
base, glabrous elsewhere; stigma rather large, 
oblique. Capsule erect, thick-fusiform, short-hairy 
and lepidote, 1.5-2 by (0.5-)0.7-0.8 cm, splitting 
from apex into 5 valves, axis and style persistent 
for a long time, valves reflexed, somewhat twisted, 
rather thick; fruit-pedicel 6-11 cm. Seeds tailed 
on both ends, c. 2.5 mm. 

Distr. Malesia: New Guinea (Main Range from 
Mt Carstensz to the Oranje Mts; Western and 
Southern Highlands: Hagen Range, Mt Giluwe, 
Mt Sugarloaf). 

Ecol. Alpine grassland, occasionally on 
shrubby ridges, 3225-4000 m, locally abundant 
and a conspicuous feature in bogs or forest 
glades, on peaty soil. Fl.fr. Aug. -Dec. 

Vern. Kamungunjari, Enga: Poio, kinwi, Mendi. 



3. Series Taxifolia 

Sleum. Reinwardtia 5 (1960) 141. 
Leaves linear, 1-1.5 mm wide, in dense ± 20-merous pseudowhorls. 
Distr. Malesia: Philippines (Luzon). 



149. Rhododendron taxirolium Mkrr. Philip. J. 
Sc. 30 (1926) 419; Copil. /. ihid. 40 (1929) 156, 
t. 6; Sleum. Reinwardtia 5 (I960) 141. 

Epiphytic shrub, c. 1 m. Branches terete, gla- 
brous. Branchlets slender, papillosc-pubcrulcnt and 
sparingly lepidote; internodes 1-3 (rarely up to 5) 
cm. Leaves crowded to dense pseudowhorls at 
the upper 3-4 nodes, 20 or more together, linear, 
sessile or nearly so, rather rigid and coriaceous, 
brown when dry, paler and shining beneath, 
apex oblusc, base attenuate into the petiole, 
glabrous above at full maturity, laxly lepidote 
beneath (scales flat, small, seemingly disk-shaped, 
but in fact irregularly lobed to various degree 
in the marginal zone, centre small, somewhat 
impressed), margin shallowly crcnulate by im- 
pressed scales, flat, (\.H )2 3.5 cm by I 1.5 mm, 
midrib very slightly impressed abf)vc and obtusely 



raised beneath, nerves obsolete; petiole 0-2 mm, 
not clearly distinct from the lamina. Flowers 
solitary or in twos. Perulae membranous, broad- 
ovate, abruptly subulate at the apex, glabrous 
dorsally, margin set with caducous scales, up to 
1.2 by 0.6 cm. Bracteoles linear, very laxly hairy 
and lepidote, c. 6 mm. Pedicels slender, pubescent 
and laxly lepidote, c. I cm. Calyx disk-shaped, 
lepidote and pubescent outside, obtusely and 
shortly 5-Iobed, 3-4 mm 0. Corolla tubular-sub- 
campanulate, white, thin, 2-2.2 cm long in all, 
sparingly substcllatc-lepidote outside, a broad 
marge of the lobes excepted, pubescent within, 
tube c. I cm long, at base r. 4 nmi 0, widened to 
7-8 mm at apex, lobes broad-obovatc. apex 
rounded, c. 1 by 0.8 cm. Sianicns 10, subequal, ) 
as long as the corolla tube; filaments linear, 
subdcnscly to laxly patently hairy to nearly the 



576 



Flora Malesiana 



[ser. I, vol. 6^ 




Fig. 31. Rhododendron stenophyllum Hook./, a. Habit, x 2/3. — R. hooglandii Sleum. b. Habit, x 2/3, 
c. detail of upper resp. undersurface of leaf with epidermal tubercles, x 6 (a Griswold 121, b~c HooG- 

LAND & PULLEN 5576). 



top; anthers oblong, c. 2 mm. Disk glabrous. 
Ovary subcylindrical, densely patently white-hairy, 
with apparently but a few scales between the 
hairs, c. 3 by 1.5 mm; style columnar, hairy below, 
glabrous distally, c. 5 mm; stigma subcapitate. 



Capsule c. 1 cm. 

Distr. Malesia: Philippines (Luzon: Mt Pulog), 
twice found on trees in the mossy forest at c. 
2700 m. Fl. Feb. -March. 



4. Series Stenopbylla 

Sleum. Reinwardtia 5 (1960) 141. — Rhododendron sect. Zygomorphanthe Schltr, Bot. Jahrb. 55 
(1918) 145, p.p. — Fig. 31—32. 

Leaves linear or narrow-lanceolate, opposite or in 3-8 (sometimes up to 15)-merous pseudowhorls, 
3-7 (rarely up to 11) mm wide (at least 1.3 cm long, mostly much longer). 

Distr. Malesia: Borneo and New Guinea. 



KEY TO THE SPECIES 

1 . Ovary densely whitish tomentellous, the hairs covering the small scales . . . 150. R. stenophylluin 
1. Ovary densely and exclusively lepidote. 

2. Leaves (1.3-)1.6-2 cm long 151. R. myrsinites 

2. Leaves longer than 2.7 cm. 

3. Leaves 1.5-3 mm wide 152. R. subulosum 

3. Leaves 4-7 (-11) mm wide 153. R. purpureiflorum 



Jan. 1966] 



Ericaceae (Sleumer) 



577 




Fig. 32. Rhododendron stenophyllum Hook./. Mt Kinabalu, 2130 m (Sleumer 4685) (Sleumer, 1963). 



150. Rhododendron stenophyllum Hook. /. ex 
Becc. Nfalcsia 1 (1878) 203 in ohs., nom. nud.\ 
BuRBiDGE, The Garden of the Sun (1880) 274, 
fig., descr. ahhrev., fif(. repr.: The Garden 18 
(1880) 546; .Stapf, Trans. Linn. See. II, Hot. 4 
(1894) 196; GrBBS, J. Linn. Soc. Bot. 42 (1914) 
104; NffLLAis, Rhodod. (1917) 248; Mfrr. En. 
Born. (1921 ) 463; J. J. S. Bull. Jard. Bot. Blzg III, 
13 (1935) 452, incl. var. anffustifolium J. J. S. 
I.e.; Sleum. Rcinwardtia 5 (1960) 142; Biumea II 
(1961) 122; ihid. 12 (1963) 106; Jahrb. Dcutsch. 
Rhodod. Gcs. Bremen (1965) f. 9, non Makino, 
1910. — Fig. 31a-32. 

Lanky shrub, (O.3-)0.6-l (-1 .2) m, with slender 
stem, epiphytic or on the ground, sparsely and 
divaricately branched. Branchlcts terete, 1-1.5 mm 
0, dilutcly brown, upper inlernodc set with some 
sparse scales initially, early practically glabrous; 



internodes (1 .5-)2-IO cm. New flush starting with 
several scattered, scarious, subulate, early cadu- 
cous thin leaves, ending with normal persistent 
leaves, these F condensed into (6-)8-15-merous 
pseudowhorls at the upper (l-)2-5 nodes, narrow- 
linear, apex ) acute, often subsctaccous or 
nearly pungent, base attenuate into the very short 
and broad, hardly distinguishable petiole, sub- 
sessile, coriaceous, dark glossy green, glabrous 
and rugulose above, medium green and very 
laxly lepidole beneath (scales small, flat, sub- 
orbicular, thin marginal /.one variously angled or 
obtu.sely dented, centre dark, not impressed), 
finally glabrcsccnt on both faces, edge thinned out, 
flat, not rcvolute, (4)5-8 (rarely up to 11) cm 
by (l-)2-3 (rarely up to 4) mm, midrib impressed 
above, prominent beneath, nerves obscure; petiole 
broad, flattened, 0-2 (rarely up to 4) mm. llowers 



578 



Flora Malesiana 



[ser. I, vol. 6* 



nodding, mostly in twos, rarely solitary. Flower 
buds narrow-ovoid, 2-2.5 by 0.6-1 cm, outer few 
perulae subulate, more innerones ovate-acuminate, 
ending in a subulate acumen, inner ones ovate- 
oblong and -± shortly subulate to obtuse, all 
firmly membranous, brownish reddish when fresh, 
glabrous and somewhat lustrous, up to 2.8 by 0.8 
cm. Bracteoles filiform, glabrous, c. 1 cm. Pedicels 
stoutish, laxly substellate-lepidote and ± densely 
puberulous, 0.6-1.2 cm by c. 1 mm. Calyx patel- 
lar, rather fleshy, wavy, very shortly and obtusely 
5-lobed, c. 4 mm 0, laxly lepidote, furthermore 
puberulous at the base. Corolla narrow-campanu- 
late to broadly funnel-shaped, of a shining waxy 
texture, flame coloured, i.e. bright to dark red 
or of a clear orange-scarlet, sometimes light red 
or orange, often with orange tinge especially in 
the interior, or red towards the base, 2-2.5 (rarely 
up to 3) cm long in all, glabrous in- and outside, 
tube 1-1.5(-1.7) cm long, base 4-6 mm 0, gradu- 
ally widened to 8-10 mm, lobes broadly obovate- 
spathulate, slightly spreading, subretuse, 0.8-1.2 
(-1.4) by 0.6-0.8 cm. Stamens 10, ± equalling the 
corolla tube in length; filaments linear, broadened 
and laxly pubescent at the base or the lower 14; 
anthers narrow-oblong, shortly bilobed at the 
base, 3-4 by 1-1.3 mm. Dw/c prominent, glabrous. 
Ovary cylindric or subovoid, abrupt with the 
style, densely shortly whitish pubescent and mi- 
nutely lepidote, 4-5 by 2-2.5 mm; style glabrous, 
6-9 mm; stigma obconical or subcapitate. Capsule 
subcylindric, slightly curved, reddish-purplish, 
laxly covered with very small scales, more densely 
with spreading hairs, c. 3 by 0.4 cm, on fruit- 
pedicel c. 1.5 cm. 

Distr. Malesia: Borneo (Mts Kinabalu, Batu 
Tiban, Trus Madi, and Kemul). 

Ecol. Primary forest, locally plentiful but 
mostly sterile, on Mt Kinabalu a very common 
shrub in the upper montane oak forest, also in 
mossy forest, on mountain ridges, or on low rock 
cliff" at river's edge, in light shade, 1525-2500 m, 
sparsely up to 3350 m. Fl. Jan. -Dec. 

151. Rhododendron myrsinites Sleum. Reinwardtia 
5 (1960) 142. 

Epiphytic shrub, 0.5 m; branches 2-3 mm 0. 
Branchlets c. 1 mm 0, subdensely substellate- 
lepidote; internodes 2-4(-5) cm. Leaves in 3-5- 
merous pseudowhorls at the upper 2 or 3 nodes, or 
close together in the upper third of the internodes, 
lanceolate, apex short-acuminate and plicate, 
minutely apiculate, base broadly attenuate into 
the petiole, coriaceous, laxly lepidote on both 
faces initially, glabrous above in later stages, 
persistently lepidote beneath (scales variously 
substellately incised or lobed in the marginal zone, 
centre minute and slightly impressed), edge sub- 
crenulate by somewhat impressed scales, sub- 
revolute, (1. 3-) 1.6-2 by 0.3-0.4 cm, midrib 
slightly sunk above, prominent beneath, nerves 
obsolete; petiole slender, lepidote, c. 2 mm. 
Flowers solitary or in pairs. Perulae membranous, 
red-brown, shining, ovate, in the upper 5-6 mm 
subulate-acuminate, glabrous, up to 2 by 0.6 cm. 



Pedicels slender, densely stellate-lepidote, c. 1.8 
cm. Calyx much oblique, disk-shaped, obscurely 
lobed, c. 2 mm 0. Corolla tubular, rather fleshy, 
red, quite glabrous in- and outside, c. 3.5 cm 
long in all, tube slightly curved, 2.5 cm, base 
saccate, c. 4 mm 0, widened to c. 8 mm at apex, 
lobes erecto-patent, broadly obovate, 8-9 by 5-6 
mm. Stamens 10, unequal, the longest slightly 
longer then the corolla tube; filaments linear, 
glabrous; anthers obovate, 1-1.3 mm. Disk gla- 
brous. Ovary subcylindric-fusiform, very densely 
lepidote, epilose, c. 6 by 2 mm, gradually tapering 
to the style, the latter rather slender, lepidote at 
the base only, glabrous elsewhere, c. 1.4 cm; 
stigma capitate. 

Distr. Malesia: New Guinea (Bernhard Camp, 
Idenburg R.), on tall trees in mossy forest, at 
2150 m. Once found. Fl. Feb. 

152. Rhododendron subulosum Sleum. Reinwardtia 

5 (1960) 143. 

Weak epiphytic shrublet, 20-30 cm; stem c. 3 
mm 0, few-branched, roots much thickened, c. 5 
mm 0. Branchlets 1-1.5 mm 0, glabrous below, 
subdensely clothed with star-shaped and shortly 
stalked, caducous scales, respectively tardily 
glabrescent and minutely warty at the tips or the 
ultimate internode; internodes 1-4 cm. Leaves in 
3-5-merous pseudowhorls at the upper 1 or 2 
nodes, linear, apex acute or nearly so, base 
narrowed to the petiole, coriaceous, dark green 
and somewhat glossy above, paler beneath, 
glabrous above with age, laxly lepidote beneath 
(scales variously angular or obtusely dented in the 
marginal zone, centre much darker and hardly 
impressed), edge slightly revolute, (3.5-)4-8 cm 
by 1.5-3 mm, midrib ± distinctly sunk above, 
very prominent beneath, red, nerves obscure; 
petiole subterete, red, much darker than the lamina 
in dry specimens, 2-4 by c. 0.5 mm. Umbels 1-3- 
flowered. Outer perulae subulate, more inner ones 
narrow-ovate-subulate, innermost ones ovate, 
abruptly subulate-acuminate in the upper half and 
acute, glabrous dorsally, reddish, up to 2 by 0.5 
cm. Pedicels slender, very densely stellate-lepidote, 
5-7 mm. Calyx obliquely disk-shaped, margin 
membranaceous and very shortly 5-dented, 
lepidote dorsally, c. 2 mm 0. Corolla tubular, 
probably red, c. 1.5 cm long in all, glabrous in- 
and outside, tube c. 1 cm long, 2-3 mm 0, but 
faintly dilated upwards, lobes broadly obovate- 
suborbicular, c. 4 mm 0. Stamens 10, 1.4-1.6 cm; 
filaments filiform, glabrous; anthers subglobose, 
c. 1.5 mm 0. Disk prominent, glabrous. Ovary 
subcylindric, very densely stellate-lepidote, c. 3 by 
1.3 mm, abrupt with the style, the latter rather 
slender, glabrous, 5-6 mm; stigma club-shaped. 
Capsule c. 1.5 by 0.4 cm, valves reflexed and 
twisted. Seeds very narrow, tailed at both ends, 
2-3 mm; fruit-pedicel c. 1.7 mm. 

Distr. Malesia: New Guinea (Doormantop), 
ridge at 1450 m. Once collected. Fl. Nov. 

153. Rhododendron purpureiflorum J. J. S. Med. 
Rijksherb. n. 25 (1915) 3; Nova Guinea 12 (1917) 



Jan. 1966] 



Ericaceae (Sleumer) 



579 



502; ibid. 12 (1918) t. 197; Sleum. Reinwardtia 5 
(1960) 143. — R. podocarpoides Schltr, Bot. 
Jahrb. 55 (1918) 149. 

Epiphytic shrub. 20-50 cm; branches numerous, 
terete, rather slender. Branchlets 1-1.5 mm o, 
subdensely clothed with rufous stellate scales; 
internodes (1.8-)2.5-5.5 cm. Leaves in 3-6-merous 
pseudowhorls at or near the upper 3-4 nodes 
(several much reduced ones as subulate bracts 
along the upper internodes), linear-lanceolate, 
ape.x obtuse or subacute, base attenuate into the 
petiole, rather coriaceous, lepidote on both faces 
initially, glabrescent above with age, la.xly persist- 
ently lepidote beneath (scales irregularly, often 
obtusely lobed or dented in the marginal zone, 
flat, rather pale, centre faintly or not sunk), 
margin flat, 2.7-4.5(-7) cm by 4-6(-7, sometimes 
up to 11) mm, midrib impressed above, rather 
strongly obtusely prominent beneath, nerves 
obscure; petiole lepidote, stoutish, 2-4(-5) mm. 
Umbels 2-4-flowered. Pedicels slender, subdensely 



rufous-stellate-lepidote, 1.2-1.5 cm. Calyx disk- 
shaped, oblique, lepidote, very shortly obtusely 
5-lobed, c. 2.5 mm o. Corolla tubular, somewhat 
zygomorphous, red-lilac or pale pink, very laxly 
substellate-lepidote or glabrous outside, glabrous 
inside, 2.7-3.2 cm long in all, tube subcylindric, 
1.7-2.2 cm, at the 5-saccate base c. 4 mm 0, 
widened to 6-7 rrun at apex, lobes erecto-patent, 
broad-obovate, 7-8(-9) by 5-6 mm. Stamens 10, 
unequal, the longest equalling or somewhat exceed- 
ing the corolla tube; filaments linear, glabrous; 
anthers obovate, c. 1.5 mm. Disk much prominent, 
glabrous. Ovary fusiform, attenuate at the base, 
gradually tapering to the style, densely lepidote, 
c. 4 by 1 .6 mm; style slender, glabrous, ±_ equalling 
the stamens; stigma suboblique, capitate, purple. 

Distr. Malesia: New Guinea (Perameles and 
Schrader Mts, and near Bernhard Camp, Iden- 
burg R.). 

Ecol. Mossy forest on tall trees, 1100-2150 m. 
Fl. Dec, Feb., May-June. 



5, Series Citrina 

Sleum. Reinwardtia 5 (1960) 144. 

Diff"ers from all other series of subsect. Euvireya by having only 5 stamens. 
Distr. Malesia: Sumatra, Java, and Bali. 



154. Rhododendron citrinum (Hassk.) Hassk. Cat. 
Hort. Bog. (1844) 161; Moritzi, Syst. Verz. (1846) 
42; Walp. Rep. 6 (1847) 420; Hook. Bot. Mag. 
(1854) t. 4797; Zoll. Syst. Verz. 2 (1854) 137; 
Planch. Fl. Serres 10 (1854-55) 5, t. 970; Walp. 
Ann. 5 (1858) 452; MiQ. Fl. Ind. Bat. 2 (1859) 
1058; Ann. Mus. Bot. Lugd.-Bat. 1 (1863) 43, 
incl. ft albijiorum MiQ. I.e.; Koord. Teysmannia 10 
(1899) 450; Exk. Fl. Java 3 (1912) 4; Koord.- 
ScHUM. Syst. Verz. 1 (1912) fam. 233, p. 106; 
J. J. S. in K. & V. Bijdr. 13 (1914) 94; Koord. Fl. 
Tjib. 3 (1918) 3; Steen. Bull. Jard. Bot. Btzg 
III, 17 (1948) 387, 388; Amsh. in Back. Bekn. Fl. 
Java (em. ed.) 7 (1948) fam. 162, p. 4; Sleum. 
Reinwardtia 5 (1960) 144; Steen. Endeavour 
21 (1962) 189, t. 2. — Azalea citrina Hassk. 
Flora 25 (1842) Beibl. p. 30; O. Ktze, Rev. Gen. 
PI. 2 (1891) 387. — R. zippelii Bl. Fl. Jav. PI. 
in6d. (1863-83) t. 4. — R. jasminijiontm {nan 
Hook.) Koord. Junghuhn Gcdenkb. (1910) 184, 
cf. Hallifr/. Med. Rijkshcrb. n. I (1910) 35. 

var. citrinum. 

Rather small, epiphytic, divaricately branched 
shrub, 0.7-2 m. Branchlets l-l.5(-2) mm o, terete, 
tips : densely clothed with flattish rufcsccnt 
scales, glabrescent below, greyish corticated in the 
older parts; internodes 2-5 cm. Leaves in 4-5- 
mcrous pseudowhorls at the upper l-2(-3) nodes, 
unequal in the same pscudowhorl, elliptic, 
oblong-elliptic or nearly lanceolate, rarely obovatc- 
clliptic, apex < broadly acuminate, subacute 
or mostly obtuse (terminal gland very small), 
base attenuate to the petiole, coriaceous, dark 
green and glossy above, dilutcly green with brown 
scales beneath, densely lepidote initially in the 



youngest leaves, early glabrescent on the upper 
surface, which is a little wrinkled in dry specimens, 
more persistently and subdensely lepidote beneath 
(scales distant, flat, rather small, variously obtusely 
angled or more deeply dented in the marginal, 
early dissolute zone, dark centre small, somewhat 
impressed), finally ± glabrescent beneath too, 
margin slightly revolute in dry specimens, (1.5-) 
2.5-5(-6.5) by (l-)l.5-2.5(-2.7, rarely up to 4) 
cm, midrib slightly impressed above, prominent 
beneath, nerves 5-8 pairs, widely spreading, 
anastomosing, faintly raised beneath, often 
inconspicuous; petiole lepidote, rather slender, 
scmiteretc, (3-)4-9(-12) mm. Umbels 2-4(-5)- 
flowered, occasionally reduced to a solitary flower. 
Flowers a little fragrant, nodding. Flower buds 
ovoid, outer perulae ovate-acuminate, finely 
sericeous dorsally, the middle line set with early 
caducous scales, inner ones oblong to spathulate, 
glabrous dorsally, all fimbriate along the margin 
or at the top only, up to 10 by 3 mm. Bractcoles 
filiform, practically glabrous, up to 8 mm. 
F*cdicels red, slender, pubcrulent and laxly lepidote 
below, more densely so upwards, c. 1.5 cm at 
anthesis. Caly.x oblique, obtusely 5-lobed (lobes 
1-2 mm, or sonKtimcs one or the other lobe a 
little longer), lepidote dorsally, 3-4 mm o. Corolla 
broadly funnelforni-campanulate, pale or creamy 
yellow, yellowish white, or lemon, lohed nearly 
halfway, 1.5-1.7 cm long in all, outside very laxly 
lepidote or almost glabrous in the Javanese, sub- 
densely lepidote in the Balinese specimens, 
glabrous inside, tube v. 4 mm at base, gradually 
dilated to 6 mm at apex, lobes : spreading, 
somewhat unequal, obovate-spalhulalc, apex 
rounded, very slightly or not rctuse, 7-8 mm. 



580 



Flora Malesiana 



[ser. I, vol. 6* 



Stamens 5, unequal, a little exserted from the 
corolla tube, c. 1.2 cm; filaments linear, glabrous, 
red or orange; anthers oblong, deeply emarginate 
on both ends, 2.2-2.5(-2.8) by c. 1 mm, yellow or 
orange, dorsifix, the connective white. Disk very 
prominent, glabrous. Ovary obliquely oblong- 
ellipsoid, somewhat 5-angled respectively -ribbed, 
lepidote at the base, glabrous for the rest, c. 
4 by 1.5 mm, green, initially gradually tapering 
to the slender glabrous style (5-6 mm), but rather 
abrupt in laterstages;stigmaturbinate-subcapitate. 
Capsule obliquely cylindric, shortly attenuate at 
both ends, slightly curved, red, deeply 5-grooved, 
glabrous, 2-2.5 cm by 4 mm. Seeds very narrow, 
c. 4 mm including the filiform tails at both ends. 

Distr. Malesia: W. Java and Bali. 

Ecol. In very humid primary forest, subalpine 
forest or forest border, (1000-) 1450-2400 (-2900) 
m. Fl. Sept.- March. 



Vern. Tjantigi, S. 

Note. Reported for Central Java and possibly 
growing on Mt Dieng; no specimens seen. 

var. discoloratum Sleum. Reinwardtia 5 (1960) 
145. — R. citrinum inon Hassk. s. sir.) MiQ. Fl. 
Ind. Bat. Suppl. 1 (1860) 251, 586; Ridl. J. Fed. 
Mai. St. Mus. 8 (1917) 58; Sp. Moore, J. Bot. 63 
(1925) Suppl. 57. — /?. multicolor {non MiQ.) 
Sp. Moore, I.e. 

Corolla laxly to subdensely lepidote outside, 
mostly orange red to bright scarlet, rarely light 
lemon yellow. Ovary not rarely completely or for 
a long time covered with scales. 

Distr. Malesia: Sumatra (Atjeh: West Coast; 
Palembang; Lampong). 

Ecol. In mossy iVaccinium-)f orest, on ridges, 
1220-2500 m. 

Vern. Mata kunai, M. 



6. Series Buxifolia 

Sleum. Reinwardtia 5 (1960) 145. — Rhododendron sect. Zygomorphanthe ScHLTR, Bot. Jahrb. 55 (1918) 
145, p.p. — Rhododendron subsect. Malesia Copel./. Philip. J. Sc. 40 (1929) 136, 152. — Rhododendron 
subsect. Linearanthera Copel./. I.e. — Rhododendron subsect. Astrovireya Sleum. Bot. Jahrb. 74 (1949) 
539. — Fig. 33—37. 

Leaves medium-sized, (0.6-)l-4 (rarely in part in the same specimen up to 7) by (0.4— )l-2 (rarely in 
part in the same specimen up to 2.5, very rarely up to 3) cm, of various form other than linear or narrow- 
lanceolate. 

Distr. Malesia. 



KEY TO THE SPECIES 

1 . Ovary hairy and lepidote to various degree (scales sometimes sparse or ~ covered (hidden) by the 

hairs). 
2. Filaments manifestly — patently hairy at least in the lower third. 
3. Corolla broadly tubular-campanulate (tube straight, — equally wide), lobes erect and short, 
equalling but V4~V5 of the total length of the corolla. Leaves obovate-lanceolate, (1.6-)2-3.5 by 

(0.4-)0.5-0.7(-0.8) cm 155. R. frey-wysslingii 

3. Corolla open-campanulate, funnel-shaped or tubular (tube widened towards the lobes and often ± 
curved), lobes erect or expanded, equalling Vi-Vi of the total length of the corolla. 

4. Corolla short, open-campanulate 156. R. pubigermen 

4. Corolla ^ elongate, funnelform, tubular-funnelform or tubular(-zygomorphous). 
5. Umbels 2^(-6)-flowered. 
6. Style hairy at least in the lower third, mostly up to almost the top. 
7. Corolla white. 
8. Style hairy and lepidote 245. R. luraluense 

8. Style exclusively hairy 157. R. vidalil 

7. Corolla pink, or red (scarlet). 

9. Corolla elepidote, glabrous or laxly hairy outside 158. R. batamense 

9. Corolla lepidote, otherwise whether or not hairy outside. 

10. Corolla hairy and lepidote outside 159. R. vandeursenii 

10. Corolla exclusively lepidote outside to various degree. 

1 1 . Style hairy in the lower third 268. R. culminicoluiii 

11. Style hairy in the lower Vz-J^, or almost to the top. 
12. Corolla 2.5 cm. Nervation of the leaves subinconspicuous on both faces. 

160. R. cornu-bovis 
1 2. Corolla at least 3.6 cm. Nervation of the leaves much impressed above, prominent beneath. 

268. R. culminicolum var. nubicola 
6. Style hairy at the base only (up to Ve of its total length), or mostly entirely glabrous. 
13. Style as long as the ovary. 
14. Umbels 2-3-flowered. Leaves 3-4.5 by 1.2 cm 161. R. planecostatum 

14. Flowers solitary. Leaves 1.5-2.3 (-2.7) by 0.5-0.8(-l) cm 162. R. bagobonum 

13. Style distinctly longer than the ovary. 

15. Anthers dorsally glabrous. 



Jan. 1966] Ericaceae (Sleumer) 581 

16. Corolla exclusively lepidote, or glabrous outside, no hairs present at all. 
17. Corolla yellow. Leaves gradually long-acuminate at apex .... 163. R. nieuwenhuisii 
17. Corolla pink to red. Leaves broadly and/or shortly acuminate, or obtuse at apex. 
18. Leaves quite entire. 
19. Corolla laxly lepidote outside 268. R. culminicolum 

19, Corolla completely glabrous outside, (2.5-)3(-3.2) cm 261. R. arfakianum 

18. Leaves minutely though distinctly crenulate. 

20. Corolla completely glabrous outside 164. R. pseudobuxifolium 

20. Corolla lepidote to various degree outside 165. R. commonae 

16. Corolla exclusively hairy, or hairy and lepidote outside. 

21. Leaves linear to linear-oblong, revolute to or almost to the midrib, 2-2.5(-4) cm by 

l.5-2(-3) mm in the revolute state. Umbels 2-3-flowered .... 166. R. abietifolium 

21. Leaves elliptic, or elliptic- or obovate-oblong, revolute to various degree, at least 4 mm 

wide. Umbels 5 (or more)-flowered. 

22. Leaves elliptic- or obovate-oblong, base ± cuneate to the petiole, (4-)6-10(-13) mm 

yf,[^Q 167. R. sheUae 

22. Leaves elliptic, base broadly attenuate to rounded, 1 5-20(-26) mm wide. 

198. R. rugosum var. coriifolium 

15. Anthers dorsally ± densely hairy 168. R. pauciflorum 

5. Umbels 7-12-flowered 

23. Leaves quite entire. Corolla densely lepidote outside 169. R. psammogenes 

23. Leaves manifestly crenulate. Corolla laxly hairy and lepidote outside 

198. R. rugosum var. coriifolium 

2. Filaments glabrous or practically so. 
24. Style hairy at least in the lower '/s, mostly so to or almost to the top. 
25. Corolla (2-)2.5(-2.8) cm long in all. Branchlets verruculose. 
26. Leaves thick-coriaceous, rigid, apex ± rounded, a terminal gland not visible, nerves ± distinctly 
prominent in fully mature leaves. Corolla red 170. R. vitis-idaea 

26. Leaves thin-coriaceous, apex mostly acuminate, rarely rounded, the terminal gland mostly 
forming an outstanding apiculus, nerves subinconspicuous. Corolla white, pink at the throat. 

171. R. rhodostomum 
25. Corolla 3.5-5(-6) cm long in all, red. Branchlets smooth or almost so. 

27. Corolla exclusively stellate-lepidote at the tube outside . . . 268. R. culminicolum var. nubicola 
27. Corolla stellate-lepidote and hairy outside 159. R. vandeursemi 

24. Style hairy at the base only (up to '/• of its total length), or quite glabrous. 
28. Corolla practically glabrous outside. 
29. Umbels 3-4-flowered. Uaves 2.5-3.5(-4) by 1 .2-1 .8(-2) cm. Petiole flattened, 1 .5-3 by l-l .5 mm. 

172. R. alternans 

29. Flowers solitary. Leaves (2-)2.2-3 by (0.7-)0.8-l cm. Petiole subterete, 3-4(-5) by c. 0.8 mm. 

173. R. leptomorpbum 

28. Corolla laxly to subdensely lepidote outside. 

30. Leaves cuneately decurrent into the ± flattened petiole. 

31 . Leaves entire or faintly crenulate, 1 .5-2.5(-3) by 0.8-1 .5 cm. Corolla 1.5-1 .8(-2) cm long m all. 

174. R, pseudonitens 

31. Leaves more manifestly crenulate, (2-)2.5-3.5M) by (0.8-)l-l,5 cm. Corolla 3-3.5 cm, 

175. R. helodes 
30, Leaves (manifestly crenulate) ± truncate at the very base, hardly or not decurrent, the petiole 

well distinguishable, a little or not flattened 165. R. commonae 

1. Ovary exclusively lepidote (but hairs may be present on the style). 
32. Style hairy to various degree (not lepidote). 
33. Style patent-hairy per c. 2 mm above the base. Leaves obovate, 1,2-1,8 by (0.5-)0,7— 1,1 (-1.3) cm. 
Filaments glabrous 176. R. papuanura 

33, Style with a ring of short patent hairs at its very base. Leaves obiong-obovate, 2.3-3,5 by 0.7-1.2 
cm. Filaments pubescent at the base 179. R. ripleyi var. basltrlchum 

32. Style exclusively lepidote up to the middle, or mostly quite glabrous. 

34. Filaments laxly to subdensely and h patently hairy or at least papillosc-pubcrulous below. 
35, Corolla (tubular-funnclform) yellowish white, rather densely hairy at the tube inside. 

177. K. acrophilum 

35. Corolla pink-red to scarlet, glabrous inside. 
36. Corolla up to 3 cm long in all. . i i i,- 

37. Corolla >- open campanulatc, c. 2 cm. Leaves oblanccolatc, apex slightly notched, lackmg a 

terminal gland. Disk glabrous 178- R- banghamlorum 

37. Corolla funnclform-campanulatc, 2.5-3 cm, Uavcs oblong-obovatc or oblong-clliptic rarely 
elliptic or subobovatc-clliptic. apex blunlish, ending in a whether or not prolruduig glaiui, 

179. R. riplcyl 



582 Flora Malesiana [ser. I, vol. 6* 

36. Corolla 5-5.5 cm long in all (subfunnelform-tubular) 180. R. alticolum 

34. Filaments completely glabrous. 

38. Corolla widely campanulate. Leaves oblanceolate 181. R. pyrrhophorum 

38. Corolla tubular or funnel-shaped. Leaves of other shape. 
39. Style ± as longasor shorter than the ovary. Corolla (strictly tubular) 1-1. 3(-l. 6) cm long in all. 
40. Leaves in general elliptic oblong, 2.3-3. 5(-4) by (0.8-)l-1.5 cm; petiole 3-5 mm. 

182. R. luteosquamatum 
40. Leaves in general ovate-elliptic, or elliptic, (l-)1.2-2.4(-3.5) by 0.6-1. 5(-2) cm; petiole 1-2 

(rarely up to 3) mm 183. R. inconspicuum 

39. Style much longer than the ovary. Corolla longer than 2 cm. 
4L Corolla glabrous outside or practically so. 
42. Leaves subsessile, ± imbricately arranged, i.e. the pseudowhorls very close together, petiole 

1-2 by 3^ mm 184. R. lamii 

42. Leaves manifestly petioled, scattered or in manifestly spaced pseudowhorls; petiole (2-)3-5 
by 1-2.5 mm. 
43. Corolla 2-2.5 cm long in all. Calyx and pedicels exclusively lepidote. 
44. Branchlets smooth. Leaves ± obovate, 3-4.5(-5) by 1.5-2(-2.3) cm . 185. R. porpbyranthes 

44. Branchlets verruculose. Leaves lanceolate to subobovate-oblong, 1.5-2 by 0.6-1 cm. 

186. R. simulans 
43. Corolla 3.5-5.5 cm long in all. 

45. Calyx and pedicels exclusively lepidote. 

46. Leaves (1.8-)2-2.7 by (1-)1.4-1.9 cm; nerves subinconspicuous . . . 187. R. ultimum 

46. Leaves 3-4(-5) by L8-2.7 cm; nerves conspicuous above or mostly so on both faces. 

188. R. brassii 
45. Calyx and pedicels lepidote and shortly pubescent. 

47. Umbels 2-flowered, or rarely a solitary flower 189. R. subullferum 

47. Umbels 3-6-flowered 190. R. scarlatiniun 

4L Corolla laxly or mostly subdensely lepidote outside. 
48. Corolla (3-)4^5(-5.5) cm long in all. 

49. Leaves broadly elliptic or ovate-elliptic, sometimes subobovate-elliptic, base rounded or 
mostly slightly cordate 191. R. atropurpureum 

49. Leaves obovate or rarely oblong-obovate, base cuneate 189. R. subuliferum 

48. Corolla (2-)2.5-3 (rarely up to 3.5) cm long in all. 

50. Leaves densely arranged in much approximate 5-8-merous pseudowhorls, broad-elliptic, 
di rounded or subcordate at the base, manifestly crenulate 192. R. buxifolium 

50. Leaves laxly arranged, in distinctly spaced pseudowhorls, ± attenuate at the base, sub- 
crenulate to entire. 

51. Ferulae small, up to 0.8 by 0.4 cm 193. R. wrightianum 

5L Ferulae rather large, 1-2 by 0.4-0.9 cm, laxly lepidote, or mostly glabrous dorsally. 
52. Branchlets distinctly verruculose. Leaves elliptic, sometimes subovate- or subobovate- 
elliptic, 2-2.8(-3) by (0.8-)l-1.4 cm; petiole 2-3 by 1.5-2 mm. Corolla 2.5-3 cm. 

194. R. subcrenulatum 
52. Branchlets smooth. 
53. Leaves oblong to elliptic- or subobovate-oblong, (2-)2.3-3.8 (rarely up to 5) by 1-1.6 

cm; petiole 3-5(-7) by 1-1.5 mm. Corolla 2-2.6 cm 195. R. rubrobracteatum 

53. Leaves narrow-oblong, or elliptic-oblong, 2-2.5 by 0.5-0.9 cm; petiole 2-3 by 0.7-1 mm. 
Corolla 3(-3.5) cm 196. R. calosanthes 

155. Rhododendron frey-wysslingii J. J. S. in Merr. impressed), finally ± glabrescent beneath too, 
Contr. Arn. Arb. 8 (1934) 123;Sleum. Reinwardtia margin very slightly revolute, (l.6-)2-3.5 by 
5 (1960) 149. (0.4-)0.5-0.7(-0.8) cm, midrib impressed above, 
Small terrestrial, much branched shrub, up to obtusely raised beneath in the lower half, becoming 
75 cm. Branchlets terete, tips sparsely to subdensely ± obsolete distally, nerves obsolete; petiole 
covered with substellate ± sessile scales, older lepidote, flat, 1-2 mm. Flowers solitary or more 
glabrescent parts first minutely verruculose, early rarely in twos, nodding. Ferulae narrow ovate- 
becoming smooth; internodes 0.3-3.5 cm. Leaves acuminate, long-subulate, membranous, laxly 
in dense 5-8-merous pseudowhorls at the upper lepidote and papillose at apex, glabrous below, 
1-3 nodes, obovate-lanceolate, apex obtuse or margin set with ± caducous scales, up to 6 by 2 
rounded, base cuneate into the petiole, ± coriace- mm, early caducous. Fedicels rather stoutish, 
ous, young ones cinnamon-lepidote on both 1-1.3 cm in anthesis, densely set with stellate 
faces, mature ones glabrescent above, lepidote for scales and sparingly very shortly hairy. Calyx 
a long time beneath (scales small, flattish, narrow small, disk-shaped, thin, shortly 5-dented, c. 3 
marginal zone subentire or mostly irregularly mm 0, lepidote and often short-hairy dorsally. 
shallowly lobed, centre relatively large, somewhat Corolla broadly tubular-campanulate, thin, red. 



Jan. 1966] 



Ericaceae (Sleumer) 



583 



1.4-1.6 cm long in all, :r densely covered with 
subentire, thickish scales outside, laxly long-hairy 
in the lower half of the tube inside, tube broad- 
cylindric. 1-1.2 cm, rounded and c. 5 mm at 
base, slightly widened to apex, lobes ovate, sub- 
acute, somewhat expanded, c. 3.5 by 3.5 mm. 
Stamens 10, subequal, somewhat exceeding the 
corolla tube; filaments linear, dilated at the base, 
± densely patently hairy to nearly the top; 
anthers subquadrangular, truncate, c. 1.4 mm. 
Disk glabrous. Ovary ovoid-conical, densely 
lepidote and villous, the hairs predominant, 
covering the scales at first, c. 4 by 2 mm, rather 
abrupt with the style, the latter 7-8 mm at anthesis, 
covered to the lower ^/s with whitish hairs; stigma 
capitate. Capsule subcylindric, laxly hairy and 
lepidote, c. 1.3 by 0.4 cm. Seeds 4 mm, the long 
tails included. 

Distr. Malesia: N. Sumatra (Gajo Lands: 
Bur ni Tdlong and RddSlong Volcano), on slopes, 
1800-2000 m. Fl. Jan., June, Sept. 

156. Rhododendron pubigermen J. J. S. in Merr. 
Contr. Am. Arb. 8 (1934) 122; SLEUM.Reinwardtia 
5 (1960) 149. 

Small, generally epiphytic shrub, up to 1.2 m; 
branches slender, divaricate. Branchlets terete, 
c. 1 mm at the upper internodes, which are ± 
densely substellate-lepidote and finely patent- 
puberulous; internodes (l-)2-6 cm. Leaves in 
(4-)6-10-merous pseudowhorls at the upper 2-3 
nodes, oblong- or lanceolate-obovate, apex obtuse- 
rounded, base gradually cuneate, coriaceous, 
rather stiff, lepidote on both sides initially, 
glabrous above at full maturity, subdensely 
and more persistently lepidote beneath (scales flat, 
pale brown, marginal zone relatively narrow, 
irregularly and shallowly incised, centre small, ± 
conspicuously impressed), margin ± revolute in 
dry specimens, (1.3-)1.8-3(-4.5, rarely up to 5.5) 
by (0.5-)0.7-l .2(-l .5)cm, midrib impressed above, 
strongly prominent in the lower half beneath, 
nerves obsolete; petiole ± flattened, lepidote, 
l(-2) mm. Flowers l-2(-3). Flower buds ovoid, 
c. 0.8 by 0.6 cm; perulae suborbicular-ovate, 
outer ones obtuse, inner ones subacuminate, 
glabrous and lustrous dorsally, margin set with 
caducous scales, up to 7 by 5 mm. Bracteoles 
lincar-subspathulate, glabrous, c. 8 mm. Pedicels 
rather slender, densely lepidote and minutely 
pubescent, 1-1.6 cm. Calyx oblique, shortly cup- 
shaped, shortly 5-dentcd or -lobed, c. 3 mm 0. 
Corolla open-campanulate, gradually attenuate 
from the limb to the base, nearly halfway 5-lobed, 
rather fleshy, red, scarlet or crimson, 1.7-1.9(-2.3) 
cm in all, laxly or subdensely substellate-lepidote 
outside, margin of the lobes excepted, glabrous 
inside, lube straight, 8-10 mm, at base rounded 
and 4-6 mm 0, lobes broadly obovatc-spathulatc, 
subtruncatc, slightly spreading, 10-12 by 7-10 
mm. Stamens 10, alternately unequal, the longer 
ones attaining ± half the length of the corolla 
lobes; filaments linear, patently hairy in the lower 
'/i-'/j. glabrous for the rest; anthers broad-oblong 
or tubquadrate, 2 mm. Disk prominent, glabrous. 



Ovary ovoid, c. 3.5 by 2.5 mm, densely set with 
longish subpatent hairs which cover the scales; 
style very abrupt with the ovary, glabrous, (5-) 
6-8 mm; stigma subcapitate. Capsule subovoid- 
elliptic, subdensely long-hairy, laxly lepidote, 
1.2-1.7 cm, the valves twisted. 

Distr. Malesia: Sumatra (East and West 
Coast). 

Ecol. In rain-forest, mossy forest or light, also 
secondary, jungle, 1200-2560 m. Fl. Jan. -Dec, 
fr. June. 

157. Rhododendron vidalii Rolfe, J. Bot. 24 (1886) 
348; Merr. Publ. Gov. Lab. Philip, n. 29 (1905) 
43; Philip. J. Sc. 1 (1906) Suppl. Ill; ibid. 3 (1908) 
Bot. 381; MiLLAis, Rhodod. (1917) 258; Brown, 
Min. Prod. Philip. For. 3 (1921)218); Merr. En. 
Philip. 3 (1923) 245; Millais. Rhodod. ser. 2 
U924) 257; Copel./. Philip. J. Sc. 40 (1929) 152, 
t. 4, f. 3-4, t. 5; Sleum. Reinwardtia 5 (1960) 151; 
Blumea 12 (1963) 106; Jahrb. Rhodod. Ges. 
Bremen (1963) f. 3. — R. verticillatum {non Low 
ex LiNDL.) ViDAL, Rev. PI. Vase. Filip. (1886) 171 ; 
Ceron, Cat. PI. Herb. Manila (1892) 106. — 
R. lussoniense Rendle, J. Bot. 34 (1896) 356; 
Merr. Publ. Gov. Lab. Philip, n. 29 (1905) 43; 
Millais, Rhodod. (1917) 204; ibid. ser. 2 (1924) 
179. _ R, whiteheadii Rendle, J. Bot. 34 (1896) 
356; Merr. Publ. Gov. Lab. Philip. //. 29 (1905) 
43; Philip. J. Sc. 3 (1908) Bot. 381; Millais, 
Rhodod. (1917) 262; ibid. ser. 2 (1924) 260; Merr. 
En. Philip. 3 (1923) 245; Copel./. Philip. J. Sc. 40 
(1929) 154, t. 4, f. 5-11. — R. curranii Merr. 
Philip. J. Sc. 3 (1908) Bot. 255, 381; Millais, 
Rhodod. (1917) 151; ibid. ser. 2 (1924) 121; 
Merr. En. Philip. 3 (1923) 243. — Fig. 33. 

Terrestrial or epiphytic shrub, stiff-woody, 
0.8-2(-3) m, with grey bark; branches divaricate. 
Branchlets 1-2 mm 0, greyish corticate in the 
older parts, tips laxly lepidote and ± patent- 
puberulent; internodes 1.5-5 cm. Leaves crowded 
in 3-5(-7)-merous pseudowhorls at the upper 
l-2(-3) nodes, oblanceolate to obovate-oblong, 
more rarely (broadly) obovate, apex rounded or 
subemarginate, base cuneate to the petiole, 
coriaceous, medium-green and rather shining 
above, paler beneath, laxly to subdensely lepidote 
on both faces when young (scales flat, rufescent, 
the relatively narrow marginal zone irregularly ± 
obtusely stellate-lobed, seemingly roundish, when 
the marginal zone has gone and the centre has 
become blackish with age, the latter slightly or not 
impressed, early ± disappearing above, remaining 
for a longer time beneath), ± glabresccnt w'th 
age on both sides, edge slightly recurved, (1.5-) 
2-4 (in part sometimes up to 5.5) by (0.8-)l-1.5 
(-2, very rarely up to 3) cm, midrib impressed 
above, prominent beneath, nerves 4-6 pairs, 
faintly visible above, slightly raised beneath, veins 
subinconspicuous; petiole 1 flattened, lepidote, 
2-3 (-5) mm. Umbels 2-3(-4)-nowcrcd, sometimes 
reduced to a single flower. Flower buds narrow- 
ovoid, c. 1.5 cm. Perulae membranous, glabrous 
or mostly subdensely to sparingly sericeous and 
laxly lepidote dorsally, short glandular-fimbriatc. 



584 



Flora Malesiana 



[ser. I, vol 6*. 



outer ones broad-ovate, short-acuminate, sub- 
acute, inner ones larger, ovate-oblong to spathulate, 
up to 1.5 by 1 cm. Bractcoies filiform, laxly hairy, 
up to 12 mm. Pedicels rather slender, laxly 
caducously substellate-lepidote, furthermore most- 
ly laxly to subdensely shortly patently hairy (but 
sometimes entirely epiiose!), (1.2-)1.5-2.5 cm 
at anthesis. Calyx small, disk-shaped, obscurely 
dented, c. 3 mm 0, lepidote and ± short-hairy 
dorsally. Corolla funnel-shaped, white (or ap- 
parently sometimes crimson, possibly due to 
hybridizing with a red-flowered other species), 
fragrant, (2.2-)2.5-3 cm long in all, limb c. 3 cm 0, 
laxly lepidote at the tube and the lower half of the 
lobes outside, laxly to subdensely hairy at the 
tube out- and inside, tube straight, (1-)1.5 cm, 
at base c. 4 mm 0, gradually widened to 8-10 mm 
at apex, lobes spreading, broadly cuneate-spathu- 
late, obtuse, 0.8-1.3 by c. 1 cm. Stamens 10, 
subequal, attaining half the corolla lobes; fila- 
ments linear and ± densely subpatently hairy 
in the lower half, filiform and glabrous distally; 
anthers narrow-oblong, c. 3 by 1 mm. Disk 
prominent, glabrous below, short-hairy at the 
upper margin. Ovar>'Subfusiform-cylindric, densely 
patent-hairy, the hairs covering small scales, c. 



5 by 2 mm, rather abruptly tapering to the style, 
which is patently hairy in its lower V^-Vi, glabrous 
towards the top, and ::- as long as the stamens; 
stigma turbinate-capitate. Capsule subcylindric, 
somewhat oblique and curved, subattenuate at 
the base, 1.5-2 by 0.4-0.5 cm, densely pubescent, 
the valves becoming recurved and twisted. Seeds 
c. 3 mm including the long-tailed appendices on 
both ends. 

Distr. Malesia: Philippines (Luzon). 

Ecol. Mostly in moss forest, but also terrestrial 
on banks in rather barren places or grassland, 
epiphytic on large, moss covered trees or terrestrial 
on shrubby mountain side or exposed ridges, 
locally not rare, 1000-2100 m. Fl. Jan. -Dec. 

Vern. Lofong, lopong, tingei, Bon., ballaybay, 
kaykayyin, ayalea. If. 

Uses. The plant is used as a cure for itches. 

158. Rhododendron hatamense Becc. Malesia 1 

(1878) 202; J. J. S. Nova Guinea 12 (1914) 135; 
Sleum. Reinwardtia 5 (1960) 153; Blumea 12 
(1963) 106. 

Low shrub, c. 0.5 m, branches divaricate. 
Branchlets irregular, slender, tips laxly lepidote; 
internodes 0.8-6 cm. Leaves in 3-5(-6)-merous 




Fig. 33. Rhododendron vidalii Rolfe. Mt Polis, 2000 m, Luzon, Philippines (Steiner2175) (Sleumer, 1961). 



Jan. 1966] 



Ericaceae (Sleumer) 



585 



pseudowhorls at the upper 1-2 nodes, ovate- 
elliptic to elliptic, apex short-acuminate or broad- 
attenuate, obtuse or sometimes rounded, or 
apiculate, base broad-cuneate or attenuate, thin- 
coriaceous, glabrescent above with age, sub- 
densely lepidote for a long time beneath (scales 
small, flat, substellate, faintly sunk), finally — 
glabrescent beneath with only the blackish small 
pits remaining, margin slightly cartilaginous, 
mostly rather distinctly revolute, 2-4(-7) by (0.9-) 
1.5-2 (rarely up to 3.5) cm, midrib impressed 
above, prominent beneath, ner\es 4(-6) pairs, 
rather straight below, — looping near the edge, 
finely sunk above. — distinctly raised beneath, or 
subobscure as are the veins; petiole flattish, 
2-3(-4-) mm. Umbels 2-4(-6)-flowered. Ferulae 
ovate-acuminate, short-subulate, membranous, 
glabrous, margin fimbriate by caducous scales, up 
to 10 by 5 mm. Bracteoles filiform, up to 1 cm. 
Pedicels rather slender, densely lepidote and 
patently hairy to subvillous, l-1.8(-2.4) cm. 
Calyx disk-shaped, often reflexed, obscurely 5- 
lobed, c. 3 mm 0, lepidote and hairy dorsally. 
Corolla tubular-funnelform, suboblique, c. 2.5 cm 
long in all, bright to dark red or crimson, waxy 
and glossy, laxly pubescent, chiefly in the lower 
half, or sometimes glabrous, and certainly bearing 
no scales outside, laxly patently hairy in the lower 
half of the tube inside, tube subcylindric, c. 1.5 cm 
long, 4-5 mm at base, a little widened upwards, 
lobes much spreading, broadly obovate-spathulate, 
c. 0.8-1.2 cm. Stamens 10, equalling halfway the 
corolla lobes; filaments linear, laxly patently hairy 
above the base to the lower '/i, glabrous above; 
anthers dark purple to almost blackish, oblong- 
obovate, 2 mm. Disk glabrous. Ovary subovoid, 
densely shortly set with spreading hairs or toment- 
ulose, the hairs covering the scales, c. 5 by 3 mm, 
tapering to the style, the latter 3: equalling the 
stamens, hairy to near the top; stigma subcapi- 
tate. 

Distr. Malesia: New Guinea (Vogelkop 
Peninsula: Arfak, Tamrau and Nettoti Mts). 

Ecol. Epiphytic in mossy Nothofagus forest, 
1550-1900 m, apparently very scattered or rare. 
Fl. Oct.- March. 

Vern. Bingnui, Hattam. 

159. Rhododendron vandeurscnii Sleum. Blumea 1 1 
(1961) 123; ihid. 12 (1963) 108. 

Erect epiphytic or terrestrial shrub, 0.5-1 m; 
upper intcrnodcs rather slender, lepidote, 0.4—4 
(-6) cm, lower ones early glabrescent. Leaves in 
3-4-mcrous pseudowhorls at the upper 2(-3) 
nodes, obovatc or elliptic-obovatc, apex rounded, 
sometimes faintly retusc, base cuneate to the 
petiole, r convex above, coriaceous, laxly lepidote 
on both faces initially, glabrous and a little glossy 
above with age, rather dull and subdcnsely lepidote 
beneath (scales flattish, hyaline marginal zone 
variously dented or incised, early dissolute, centre 
thick and more persistent, scales thus sometimes 
seemingly rotund and entire), edge much revolute, 
(!.5-)2^r-5) by (0.8-)l-l.8 (rarely up to 2.5) 
cm, midrib flat and rather inconspicuous above. 



thick and obtusely prominent below, decreasing 
upwards beneath, nerves c. 5 irregular pairs, 
slightly prominent beneath only, or hardly visible, 
no reticulation; petiole lepidote initially, 3-4(-5) 
by c. 1 mm. Flowers pendent, solitary, rarely in 
twos. Ferulae broadly ovate, subacuminate- 
apiculate, keeled and lepidote dorsally, manifestly 
fimbriate, known up to 6 mm. Pedicels rather 
stoutish, 1.3-2.3 cm, laxly to subdensely stellate- 
lepidote and laxly subpatently hairy. Calyx ± 
obliquely disk-shaped, shortly subacutely 5-lobed, 
lepidote dorsally, 2.5-3.5 mm 0. Corolla tubular, 
not much subcampanulate-dilated upwards, 4.5-5 
(-5.5) cm in all, red or deep crimson, laxly stellate, 
lepidote and short-hairy at least in the lower part 
outside, practically glabrous inside, tube rather 
straight or slightly curved, 3.5-4.3 cm, 4-5 mm 
at base, widened to 1.2-1. 5 (-2) cm at apex, lobes 
erect, broadly subspathulate-obovate or almost 
rounded, 8-10 by 6-8 mm. Stamens 10, ~z as 
long as the corolla; filaments linear (1 mm), 
laxly hairy or almost glabrous; anthers broad- or 
subobovate-oblong, 2.7-3 by c. 1 mm, base 
obtuse. Disk glabrous below, hairy at the upper 
margin. Ovary subcylindric, densely set with sub- 
patent hairs and with scales, 6-7 by 2 nun, 
tapering to the style, the latter ± as long as the 
corolla, laxly subpatently hairy in the lower Vi 
to V2, elepidote; stigma thick-turbinate. 

Distr. Malesia: New Guinea (Central High- 
lands). 

Ecol. Mossy, primary and secondary forest, 
2285-2770 m. Fl. July. 

160. Rhododendron cornu-bovis Sleum. Rein- 
wardtia5 (1960) 152. 

Small, straggling shrub, up to 2 m, with rather 
slender, divaricate branches. Branchlets subterete, 
c. 1 mm 0, tips ± flattened and densely clothed 
with rufescent substellate scales; internodes 1.5-4 
(-6) cm. Leaves in 4-6-merous pseudowhorls at 
the upper 2-3 nodes, obovate-oblong to obovate, 
apex broadly attenuate, obtuse or rounded, base 
cuneate into the petiole, coriaceous, firm, youngest 
ones lepidote on both faces, early glabrescent 
above, persistently subdensely lepidote beneath 
(scales rather large, flat, irregularly substellate- 
lacerate in the marginal zone, centre slightly 
impressed), edge slightly recurved, 2.5-3.5(-4) 
by 0.8-1.4 cm, midrib grooved above, prominent 
beneath, nerves 3-4 pairs, subinconspicuous, often 
obsolete beneath; petiole lepidote, 2-3(-4) mm. 
Umbels 2-3-flowered. Ferulae broad-ovate to 
ovate-oblong, rather obtuse, glabrous dorsally, 
margin fimbriate by caducous scales, up to 8 by 5 
mm. Bracteoles filiform, very laxly lepidote, c. 
5 mm. Pedicels rather slender, subdensely stcllate- 
lepidotc and sparsely very shortly patent-pubescent, 
1-1.4 cm. Calyx disk-shaped, oblique, lepidote 
and short-pubescent dorsally, indistinctly 5-dented, 
2.5 mm 0.Coro//a obliquely tubular-funnel-shaped, 
f. 2.5 cm long in all, pinkish red or crim.son, sub- 
densely or more laxly clothed with rufescent sub- 
stellate scales outside, laxly hairy at the tube 
inside, tube slightly curved, 1.2-1.4 cm, at base 



586 



Flora Malesiana 



[ser. I, vol. 6* 



4-5 mm a, widened to 6-8 mm at apex, lobes ± 
spreading, obovate-spathuiate, 9-10 by 6-7 mm. 
Stamens 10, subequal, reaching halfway the corolla 
lobes; filaments filiform, laxly patently hairy to 
the lower 2/3, glabrous elsewhere; anthers broad- 
oblong, c. 1.6 mm. Disk prominent, glabrous. 
Ovary subcylindric, densely shortly whitish 
greyish hairy and lepidote, the scales i covered 
by the hairs, c. 5 by 2 mm; style a little oblique, 
gradually narrowed from the ovary, laxly patently 
hairy in its lower half, elepidote, glabrous for the 
rest, nearly equalling the corolla in length; stigma 
subcapitate. Submature capsule subfusiform- 
cylindric, somewhat curved, densely hairy and 
lepidote, 2.2 by 0.4 cm. 

Distr. Malesia: W. New Guinea (Waigeou I.: 
top of Mt Nok: Buflfelhoorn), rather common 
there in mossy forest shrub vegetation at 860 m. 
Fl. June. 

161. Rhododendron planecostatum Sleum. Rein- 
wardtia 5 (1960) 153. 

Epiphytic, much branched shrub. Branchlets 
terete, tips appressedly lepidote, glabrous below; 
internodes 2-4 cm. Leaves in 3-5-merous pseudo- 
whorls at the upper 1-2 nodes, oblong-elliptic or 
subovate-oblong, apex shortly acuminate, rather 
obtuse, base cuneate, coriaceous, glabrous above 
at full maturity, subdensely lepidote beneath 
(scales irregularly substellate-dented or -incised 
in the marginal zone, centre minute, slightly 
impressed), somewhat recurved at the edge in 
dry specimens, 3-4(-4.5) by 1-2 cm, midrib 
slightly grooved above the base, but i flattish 
distally on the upper, dilated below and gradually 
narrowed distally on the undersurface, paler than 
the lamina and flat or faintly sunk there, nerves 
c. 6 pairs, irregular, slightly raised above, rather 
inconspicuous beneath, reticulation 0; petiole 
much flattened, 3^ by c. 1.5 mm. Umbels 2-3- 
flowered. Flower buds narrow-ovoid, 1.7-2 by 
0.8 cm. Ferulae ovate-acuminatt, subacute, mem- 
branous, glabrous dorsally, very shortly fimbriate, 
up to 1.7 by 0.7 cm. Bracteoles linear, glabrous, 
c. 1 cm. Pedicels rather robust, (7-)10-15 mm, 
laxly caducously lepidote and more densely 
clothed with short subpatent hairs. Calyx disk- 
shaped, very shortly 5-dented, subglabrous, 2.5 
mm 0. Corolla tubular, c. 2 cm long in all, thin, 
said to be of a beautiful red, glabrous outside, 
laxly hairy in the lower half of the tube inside, 
tube 1.3-1.4 cm long, at base 4-5 mm 0, widened 
to 6-7 mm at apex, lobes erecto-patent, broadly 
obovate, 6-7 by c. 5 mm. Stamens 10, very un- 
equal, 1.2-1.5 cm; filaments linear and densely 
patent-pubescent in the lower third, glabrous and 
filiform for the rest; anthers obovate-oblong, 2 
mm. Disk glabrous. Ovary thick-cylindric, densely 
appressedly hairy, practically elepidote, c. 4 by 2 
mm; style rather abrupt with the ovary, glabrous, 
slender, 4 mm; stigma subclavate-capitate. 

Distr. Malesia: Borneo (Mt Kinabalu), 
epiphyte on big tree over the Liwago R., 1830- 
2135 m. Once found. Fl. Dec. 



162. Rhododendron bagobonum Copel. /. Philip. J. 

Sc. 40 (1929) 151, t. 4, f. 1-2; Sleum. Reinwardtia 
5 (1960) 154; Blumea 12 (1963) 106. — R. cunei- 
folium Staff var. subspathulatum Ridl. J. Str. Br. 
R. As. Soc. n. 33 (1900) 23; Merr. En. Born. 
(1921) 461. — /?. galioides J. J. S. Bull. Jard. Bot. 
Btzg III, 13 (1935) 445. 

Epiphytic shrublet up to 0.6 m, widely branched. 
Branchlets slender, terete, tips only laxly lepidote 
and whether or not patent-puberulous; internodes 
0.9-5 cm. Young shoots with linear, rather long 
persistent ramenta. Leaves in (6-)8-12-merous 
pseudowhorls at the upper 2^(-6) nodes, spathu- 
late-lanceolate, apex short acuminate, obtuse or 
subacute, base long-cuneate into the petiole, sub- 
coriaceous, but stiff", glabrescent above, laxly to 
subdensely and + persistently lepidote beneath 
(scales irregularly dented or lobed at the thin 
marginal zone, centre rather small, slightly im- 
pressed), entire to faintly wavy or crenulate, but 
slightly revolute at the very edge in dry specimens, 
1.5-2.3(-2.7) by 0.5-0.8(-l) cm, midrib ± flat 
above, slightly raised in the lower half beneath, 
nerves obscure; petiole 0.5-1.5 mm. Flowers 
solitary. Perulae narrow ovate-acuminate or 
lanceolate, shortly subulate, membranous, gla- 
brous, up to 1.4 by 0.3 mm. Bracteoles linear- 
spathulate, c. 5 mm. Pedicels stoutish, densely 
shortly patently hairy and lepidote, 4—6 mm. Calyx 
disk-shaped, obscurely obtusely 5-lobed, sparsely 
lepidote and hairy, 2.5 mm 0. Corolla tubular, red 
or scarlet, sometimes red tinged orange, laxly 
lepidote outside, laxly hairy in the lower half of 
the tube inside, c. 1.2 cm long in all, tube straight, 
c. 9 mm long, at base 4—5 mm 0, somewhat narrow- 
ed below the lobes, the latter suberect, ovate to 
suborbicular, c. 2.5 mm 0. Stamens 10, as long as 
the corolla tube; filaments flat linear, laxly patently 
hairy in the lower half; anthers obovate, 1 mm. 
Disk very low, glabrous. Ovary subovoid-cylindric, 
densely pubescent and lepidote (scales covered 
by the hairs), 4-5 by 2.5 mm; style rather abrupt 
with the ovary, slenderly columnar, glabrous, c. 
4 mm. Submature capsule cylindric-subfusiform, 
2-2.5 cm by 3.5-4.5 mm, with a very short (4 mm) 
subpersistent style, laxly hairy and lepidote, on a 
stoutish fruit-pedicel 9-12 mm. Seeds narrow- 
fusiform, c. 3 mm including the long tails. 

Distr. Malesia: Borneo, Philippines (Min- 
danao). 

Ecol. In primary (moss) forest on mountain 
ridges, also in heath forest, locally common on 
Mt Kinabalu mostly in open sunny places, 900- 
2135 m. Fl. Jan.-Dec. 

Vern. Mogang-bagobo, Philippines. 

163. Rhododendron nieuwenhuisii J. J. S. Ic. Bog. 

4 (1910) t. 323; Merr. En. Born. (1921) 462; 
Sleum. Reinwardtia 5 (1960) 154; Blumea 11 
(1961) 122; ibid. 12 (1963) 107. 

Slender, generally epiphytic shrub. Branchlets 
elongate, terete, 1-1.5 mm 0, tips densely rusty- 
stellate-lepidote; internodes (2-)3-7 cm. Leaves in 
3-5(-6)-merous pseudowhorls at the upper 3-4 
nodes, together with subulate ± persistent 



Jan. 1966] 



Ericaceae (Sleumer) 



587 



ramenta (c. 0.7 cm), lanceolate, rarely ovate- 
lanceolate, apex gradually long-acuminate and 
often somewhat curved, subacute, base attenuate, 
acute or subobtuse, subcoriaceous, ± densely 
lepidote on both faces initially, finally glabrescent 
above, more persistently lepidote beneath (scales 
rusty, sessile, deeply and rather regularly stellate- 
incised at the marginal zone, centre small, not 
impressed), edge slightly or not recurved, 2.5-5.5 
(rarely up to 6.5) by 0.7-1.8(-2) cm, midrib sunk 
above, very prominent beneath, nerves 7-1 1 pairs, 
subpatent, a little raised above and impressed 
beneath, lamina thus becoming ± buliate between 
the nerves, veins — obscure; petiole subterete, 
grooved above, densely lepidote, 3-5 by c. 1 mm. 
Flowers solitary or in twos. Flower buds ovoid, 
outer perulae subulate, inner ones larger, ovate- 
acuminate, acute, steliate-lepidote dorsally, fimbri- 
ate, up to 2 by 0.7 cm. Pedicels rather robust, 
densely steliate-lepidote, 3-5 mm. Calyx oblique, 
shortly 5-dented, lepidote dorsally, c. 2.5 mm 0. 
Corolla tubular- funnel-shaped, deep bright yellow, 
5-parted to '/o-'/s, c. 2.8 cm long in all, tube sub- 
cylindric, straight, 1-1.2 cm long, at base 4 mm 
0, slightly dilated upwards, laxly steliate-lepidote 
outside, glabrous inside, lobes obovate, glabrous, 
1.2-1.6(-1.8) by (0.8-)l-1.6 cm. Stamens 10, c. 
1.8 cm; filaments filiform, glabrous to c. 3 mm 
above the base, nearly villous for the following 
4-6 mm, again glabrous distally; anthers linear- 
oblong, 3.5-4.5 by 0.8 mm. Disk prominent, 
hairy at the upper margin. Ovary subcylindric, 
villous, the hairs covering the scales, c. 6 by 2 mm, 
seemingly abrupt with the style, the latter thickish, 
c. 1 cm; stigma club-shaped, oblique, deeply 5- 
lobed. Capsule fusiform-cylindric, densely villous, 
c. 1.8 by 0.35 cm. 

Distr. Malesia: Borneo. 

Ecol. In dense forest, along river, also in 
'heath-forest', (100-)350-800 m, as far as indicated 
on the collector's labels, possibly at higher 
altitudes in Central Borneo. Fl. Sept. -Nov. 

164. Rhododendron pseudobuxifolium Sleum. Rein- 
wardtia 5 (1960) 154. 

Low, apparently not much branched shrub. 
Branchlcls short, stiff, 1.5-2 mm 0, laxly puberu- 
lous and : caducously lepidote; internodes 2-4 
cm. Leaves crowded in the distal part of the 
upper 1-2 internodes, or in oligomerous pseudo- 
whorls, elliptic, apex broad-attenuate, obtuse, not 
rarely minutely apiculate, base broadly attenuate 
or nearly rounded, coriaceous, rigid, glabrous, 
biackish-punctulate and somewhat shining above, 
dull and laxly lepidote for a rather long time 
beneath (scales flat, variously angular or sub- 
stcllatcly dented in the marginal zone, centre 
small and impressed), finally glabrescent on 
both faces, edge distinctly cartilaginous and 
crcnulalc, 1-1.8 by 0.7-J cm, midrib narrow- 
impressed above, robust and obtusely prominent 
beneath, nerves 3 4 pairs, curved and looping 
near the edge, faintly impressed above, slightly 
raised beneath, often hardly visible; petiole slightly 
flattened, 2 3 by r. I mm. Umbels 3 4-flowcrcd. 



Perulae ovate-oblong to broadly oblong-sub- 
spathulate, apiculate or mucronulate, rigidly 
membranous, purplish and lustrous when dry, 
glabrous dorsally, margin caducously lepidote, 
1.3-2 by 0.6-0.9 cm. Pedicels densely substellate- 
lepidote and very shortly pubescent, 1.5-2 cm 
by c. 1 mm. Calyx membranous, disk-shaped, 
margin wavy and reflexed, very shortly obtusely 
5-lobed, c. 3 mm 0. Corolla tubular below, ex- 
panded at the limb, c. 3 cm long in all, wine-red, 
quite glabrous outside, laxly hairy in the lower 2/3 
of the tube inside, tube straight, c. 2 cm, at base 
5 mm 0, slightly dilated upwards, lobes obovate, 
± spreading, c. 1 by 0.8 cm. Stamens 10, very 
unequal, the longest hardly equalling the corolla 
tube; filaments linear, laxly hairy in the lower 
third, glabrous for the rest; anthers broadly 
oblong-elliptic, c. 2 by 1 mm. Disk glabrous. 
Ovary ovoid-conical, densely clothed with whitish 
hairs, which are directed upwards and cover the 
scales, rather abrupt with the style, the latter ± 
slender, glabrous, some hairs at the very base 
excepted, c. 8 mm; stigma capitate, deeply 5-lobed. 
Distr. Malesia: SW. Central Celebes (Lati- 
modjong Range: Mt Rantemario), once found at 
3300 m. Fl. June. 

165. Rhododendron commonae Foerster in Fedde, 
Rep. 13 (1914) 223; Schltr, Bot. Jahrb. 55 
(1918) 154; Lane-Poole, For. Res. Papua (1925) 
130; White & Francis, Proc. R. Soc. Queensl. 39 
(1928) 68; Diels, Bot. Jahrb. 62 (1929) 487; 
Sleum. Reinwardtia 5 (1960) 163; Blumea 11 
(1961) 124; ibid. 12 (1963) 108; Jahrb. Rhodod. 
Ges. Bremen (1963) f. 26. — R. stonori Sleum. 
Reinwardtia 5 (1960) 155. — Fig. 34a-c-35. 

Compact, stiff, not rarely flat-topped terrestrial 
shrub, (0.2-)0.5-1.5 (rarely up to 6) m; branches 
covered with a grey or blackish, peeling cork. 
Branchlets stoutish, upper parts angular, reddish 
purplish and steliate-lepidote, older ones (those 
of the past season) with numerous blackish minute 
pits besides the numerous thick cushions of the 
fallen leaves; internodes (1.5-)2-4.5 cm. Leaves 
spirally arranged, most of them rather dense 
together in the upper half, but in part scattered 
in the lower part of the ultimate shoot, not in 
proper pscudowhorls, elliptic to obovate-clliptic, 
rarely more obovate or obovate-oblong, apex 
shortly broad-acuminate or ± rounded, apical 
gland thickish and a little or hardly prominent, 
base _i- broadly attenuate (very base subtruncatc), 
not or hardly dccurrcnt into the petiole, coriaceous, 
stiff, olivaceous-green or -brownish and L shining 
when dry above, paler (yellowish to brownish) 
and !: dull beneath, provided with numcroui 
blackish pits (scars of former scales) above with 
age, rather den.sely and subpcrsistcntly lepidote 
beneath (scales distant, hyaline marginal zone 
irregularly stcllate-lobed or -incised to various 
degree, centre small, rather dark and deeply sunk), 
glabrescent (and pilled) finally beneath loo. edge 
somewhat cartilaginous and subrevolute. distinct- 
ly subserrulate-crcnulalc. (l.2-)2-3.5(-4.3) by 
(0.8 )1.2 2 cm. midrib impressed above, broad 



588 



Flora Malesiana 



[ser. I, vol. 6^ 




Fig. 35. Rhododendron commonae Foerster. Swampy place at Lake Aunde, Mt Wilhelm, 3550 m, 
E. New Guinea (Sleumer 4166) (Sleumer, 1961). 



Jan. 1966] 



Ericaceae (Sleumer) 



589 




Fig. 34. Rhododendron commonae Foerster. a. Habit, x 2/3, b. ovary and style, X 2/3, c. leaf from under- 
neath, showing the crenulate margin, x 1. — R. saxifragoides J. J. S. d. Habit, x 2/3, e. submature 
fruit with style, x 2/3 (a-c Hoogland & Pullen 6044, d Brass 21 106, e Brass 8l Meijer Drees 10184). 



and obtusely prominent in the lower half, upwards 
less so and ~ obscure distally beneath, nerves 
4-6 pairs, irregular, curved-ascending, looping 
near the edge, slightly impressed in fully mature 
leaves above, faintly raised or hardly visible 
beneath, veins subinconspicuous; petiole thick, 
hardly or not flattened, 2-4(-5) by 1-1.5 (rarely 
up to 2) mm. Umbels (3-)4-6-flowered. Outer 
perulae subulate, intermediate ones ovate-acumi- 
nate, abruptly subulate in the upper V^-Vi^ inner- 
most ones obovate-spathuiate, shining, glabrous 
or lepidote along the dorsal midrib and at the apex 
(never hairy!), irregularly short-fimbriate, up to 
15 by II mm. Bracteoles linear, up to 12 by 
1 mm. Pedicels stoutish, laxly to rather densely 
rusty-stellate-lcpidotc, cpilose or with sparse papil- 
lose hairs between the scales, 12-18 mm at 
anthcsis. Calyx oblique, lepidote at the base, 
deeply 5-lobcd, c. 4 mm 0, lobes narrow-deltoid 
or subulate to lincar-subspathulate, acute or 
obtuse, glabrous, or occasionally long-fimbriatc 
at apex, irregular both in shape and length, 
normally c. 1 mm, occasionally 2-5(-8) mm long 
and 1-1.5 mm wide at the base, apprcsscd to the 
corolla, subpcrsistcnt. Corolla tubular, i oblique 
and curved, (2-)2.5-3(-3.5) cm long in all, 



deep blood-red to purplish or crimson, laxly 
to subdensely stellate-lepidote at the tube and the 
lower part of the lobes outside, glabrous or short- 
hairy inside, tube c. 2 cm, (4-)5-6 mm at the 
5-pouched or swollen base, up to 1 cm at apex, 
lobes obliquely subspathulate-orbicular, 6-8(-10) 
mm 0, markedly veined. Stamens 10, as long as the 
corolla, or slightly exserted; filaments linear, 
glabrous or laxly shortly papillose-hairy; anthers 
obovate-oblong, 2-2.3 by 1 mm. Disk prominent, 
glabrous. Ovary subcylindricdensely covered with 
rather short grey hairs which cover small scales, 
5-6(-7) by 2-3 mm, abrupt with the style, the 
latter stoutish, glabrous or with some hairs at the 
base, nearly equalling the stamens; stigma 
obliquely capitate. Capsule subfusiform-cylindric, 
oblique, 2-2.5 by c. 0.6 cm, subdensely short-hairy 
and lepidote. Seeds c. 3 mm, the tails included. 

Distr. Malesia: New Guinea (Western, 
Eastern and Southern Highlands; Mt Saruwaged 
and vicinity). 

Ecol. Grassland and edge of Papuacedrus 
forest, in open .semi-swampy places, ai.so on 
crests of high ridges, stony ground. 2600-3990 m, 
becoming microphyllous at high altitudes, (lower- 
ing freely in fully exposed position, locally plcnti- 



590 



Flora Malesiana 



[ser. I, vol. 6^ 



ful. Fl. fr. Jan. -Dec. 

Vern. Rohn, Enga: Kepiiam, wamangali, Enga: 
Poio, kuwabe, kwcih, Mendi. 

166. Rhododendron abietifolium Sleum. Blumea 1 1 
(1961) 122; ibid. 12 (1963) 106; ibid. 12 (1964) 
340; Jahrb. Deutsch. Rhodod. Ges. Bremen 
(1965) f. 18. 

Small shrub, c. 1 m. Branchlets slender, tips 
lepidote, early glabrescent and rather shining, 
longitudinally striate lengthwise in dry specimens; 
internodes (l-)2-3(-4) cm. Leaves 8-10(-12) close 
together for 2-3 mm towards the end of the 
upper 2-3 nodes, almost in pseudowhorls, linear 
or linear-oblong, apex very shortly apiculate, or 
obtuse, sometimes subretuse, base attenuate into 
the petiole, coriaceous, rather stiff, lustrous and 
glabrous above, paler and laxly lepidote beneath 
(scales rather small, sessile, marginal zone narrow 
and variously stellate-dented or -divided, darker 
centre rather small, not much impressed), revolute 
± up to the midrib beneath, edge slightly wavy or 
very shallowly crenulate, 2-3 (sometimes up to 4) 
cm long, equally 1.5-3 (-5) mm wide in the revolute 
state, midrib deeply impressed above, thick- 
prominent between the revolute parts of the lamina 
beneath, nerves and veins faintly impressed above, 
obscure beneath; petiole slender, flattened, laxly 
lepidote, 2-3 mm. Umbels 2-3-flowered. Outer 
perulae subulate, inner ones ovate with subulate 
acumen to ovate-oblong and shortly acuminate, 
caducously lepidote fimbriate, puberulent to 
glabrous dorsally, 6-10 by 3-8 mm. Bracteoles 
linear, glabrous, irregularly incised distally, c. 1 
cm by 0.5 mm. Pedicels slender, densely patently 
long-hairy, laxly and caducously lepidote, 1-1.5 
cm. Calyx patellar, hardly 5-lobed, wavy and 
ireflexed, hairy dorsally, c. 1 mm across. Corolla 
broadly tubular below, subcampanulate-dilated 
towards the limb, brownish red in fresh, intensely 
carmine at the lobes, paler at the tube in dry 
specimens, 2.5-3 cm long in all, subdensely set 
with soft, whitish, patent hairs and almost elepi- 
dote at the tube, sparsely lepidote and not hairy 
at the lobes outside, practically glabrous inside, 
tube 1.5-1.8 cm long, c. 4 mm at the slightly 
5-saccate base, c. 1 cm at apex, lobes broadly 
obovate-spathulate, c. 1 by 0.6-0.8 cm. Stamens 
10, ± half as long as the corolla; filaments linear, 
patently hairy for the lower V^-'/s, glabrous above; 
anthers broad-oblong, 2-2.5 by 1 mm, each cell 
provided with a small basal red apiculus or gland. 
Disk prominent, glabrous. Ovary subconical- 
cylindric, densely subappressedly white-hairy, 
caducously lepidote, c. 3 by 1.5 mm, rather 
abrupt with the style, the latter slender, quite 
glabrous, c. 1.3 cm; stigma shortly turbinate. 
Capsule thickly obovoid-cylindric, densely short- 
hairy and laxly lepidote, c. 1.2 by 0.4-0.5 cm, on 
fruit-pedicel 2 cm by 1 mm. 

Distr. Malesia: North Borneo (Mt Kinabalu: 
Mesilau East route). 

Ecol. In summit shrubberies at 3160-3650 m, 
rather frequent locally. Fl. July-Dec.,//-. Aug. 

Note. Forms with longer and wider, narrow- 



oblong, less revolute leaves (3.5-5 by 0.6-1 cm), 
repeatedly collected on the east shoulder of Mt 
Kinabalu between 3050 and 3650 m, might be 
hybrids with either R. sheilae or R. rugosum 
respectively its var. coriifolium. 

167. Rhododendron sheilae Sleum. Blumea 11 
(1961) 122; ibid. 12 (1963) 107; ibid. 12 (1964) 
340. 

Terrestrial shrub, up to 1.8 m, much branched 
distally. Branchlets slender, densely lepidote at 
the tips, glabrescent below; internodes l-3(-4) cm, 
upper ones 1.5-2 mm 0. Leaves in (4-)5-6(-8)- 
merous pseudowhorls at the upper 2-3 nodes, 
narrow-oblong to obovate- or elliptic-oblong, 
broadly attenuate towards the apex, which is 
obtuse or rarely rounded, or even sometimes 
faintly retuse, cuneate towards the petiole, 
coriaceous, erecto-patent, ± caducously lepidote 
initially, glabrous and glossy with age above, 
rather laxly and persistently lepidote beneath 
(scales small, flat, marginal zone rather narrow, 
early dissolute, variously dented or angular, centre 
minute, slightly though distinctly and persistently 
impressed), minutely crenulate along the margin, 
1.5-2.5 (rarely up to 4) by (0.4-)0.6-l (rarely up 
to 1.3) cm, midrib impressed, obtuse and much 
prominent beneath, nerves 4—5 pairs, not much 
conspicuous; petiole densely lepidote, (l-)2-3 by 
c. 1 mm. Umbels (3-)4-6-flowered. Perulae ovate- 
acuminate, apiculate, lepidote dorsally, c. 1.5 cm. 
Bracteoles subfiliform, glabrous, c. 8 mm. Pedicels 
stoutish, rather densely patently short-hairy and 
more laxly and ± caducously lepidote as is the 
calyx, 1.2-2 cm, somewhat accrescent after flower- 
ing time. Calyx obliquely patellar, subobscurely 
5-lobed or -wavy, c. 3 mm 0. Corolla ± funnel- 
shaped, broadly tubular below, dilated towards 
the spreading lobes, deep purple red, subdensely 
to more laxly lepidote and shortly white-hairy 
at the tube and the base of the lobes outside, 
laxly hairy in the lower part of the tube inside, 
glabrous for the rest, 2.5-3.2 cm long in all, tube 
cylindric, c. 1.5 cm, 4-6 mm below, widened 
to 9-10 mm at apex, lobes broadly obovate, 
1-1.3 by c. 1 cm. Stamens 10, subequal, reaching 
about halfway the corolla lobes; filaments linear, 
slightly dilated and laxly hairy below, glabrous 
for the upper 2/,; anthers broadly oblong, c. 2.5 
by 1 mm, each theca ± distinctly protracted into 
a short basal apiculus. Disk prominent, hairy but 
for the upper margin. Ovary conical-cylindric, 
3-4 by 2-2.5 mm, densely white-hairy and more 
laxly set with small subrotund, finally ± caducous 
scales especially distally, abrupt with the style, 
the latter thickish, hardly exserted from the corolla 
tube, quite glabrous; stigma thick-turbinate, very 
shortly 5-lobed. 

Distr. Malesia: North Borneo (eastern 
shoulder of Mt Kinabalu, mainly at junction of 
the Mesilau East and the Royal Society Expedi- 
tion (1961) route, just below pinnacles of main 
massif). 

Ecol. On fairly flat crest of ridges or boulders, 
in dense vegetation of low trees and shrubs of 



Jan. 1966] 



Ericaceae (Sleumer) 



591 



equal height, at 3240-3670 m, very local. FI. fr. 
Aug. 

Note. Presumably a more or less intermediate 
hybrid between R. abietifolium and R. buxifolium 
var. robustum. Almost ripe capsules were found 
te be full of seeds. 

168. Rhododendron pauciflonim K. & G. J. As. 

Soc. Beng. 74. ii (1905) 75; Ridl. F1. Mai. Pen. 2 
(1923) 219; Slelm. Reinwardtia 5 (1960) 156. — 
R. elegans Ridl. J. Linn. Soc. Bot. 38 (1908) 
314, repr. J. Fed. Mai. St. Mus. 2 (1908) 120; 
ibid. 6 (1915) 49, 157; Fl. Mai. Pen. 2 (1923) 220. 

var. pauciflonim. 

Small, epiphytic shrub, with swollen roots and 
divaricate branches. Branchlets 1-2 mm o; upper 
internodes laxly lepidote and patent-puberulous, 
1.5-3 cm. Leaves in 3-6-merous pseudowhorls at 
the upper 2-3 nodes, obovate to obovate-elliptic, 
more rarely oblanceolate-elliptic, apex rounded, 
sometimes retuse, or broadly attenuate, rarely 
apiculaieorsubacuminate, base — broadly cuneate 
to the petiole, coriaceous, dark to medium green 
and a little shining above, paler and dull beneath, 
initially lepidote on both faces, glabrescent above 
with age, more persistently and subdensely lepidote 
beneath (scales irregularly lobed or dented at the 
thin marginal zone, centre — distinctly impressed), 
margin narrowly cartilaginous, a little recurved 
in dry specimens, 2-3 by 1-1.5(-1.8) cm, midrib 
impressed above in the lower 2/3, broad and 
obtusely prominent beneath, nerves 3-4 pairs, 
faintly impressed above and raised beneath, 
mostly rather inconspicuous; petiole flattened, 
laxly pubescent initially, glabrescent, 1-2 by c. 
1 mm, sometimes hardly discernible from the 
lamina. Flowers solitary or in twos. Perulae few, 
ovate-acuminate, short-subulate or mucronate, 
acute, glabrous or very finely sericeous dorsally, 
fimbriate along the margin by caducous scales, 
up to 1.4 by 0.4 cm, persistent for a long time. 
Bracteoles filiform, up to 1 cm. Pedicels rather 
stoutish, densely short-hairy and stellately lepidote, 
8-12 mm. Calyx obliquely cup-shaped, shortly 
obtusely 5-dentcd, c. 2.5 mm 0. Corolla narrow- 
tubular campanuiate, bright red or rose-red, 
1.5-1.8 (-2.2) cm long in all, 1.5 cm at the limb, 
mostly sparingly caducously lepidote, rarely quite 
glabrous outside, glabrous inside, tube straight, 
1-1.3 cm, base rounded 4-6 mm 0, gradually 
widened to 7-8 mm at apex, lobes spreading, 
broad-obovatc to suborbicular, 4-5 mm 0. 
Stamens 10, alternately longer for the length of 
an anther, the longest slightly surpassing the 
corolla tube; filaments linear, laxly to subdensely 
set with longish, spreading, white hairs to the top 
and on the back of the anther, i.e. on the connec- 
tive; anthers broad-oblong, 1.5-2 mm. Disk low, 
glabrous. Ovary ovoid-conical, densely clothed 
with scales and whitish hairs, 3-4 by 2 mm; style 
abrupt with the ovary, glabrous or with some 
hairs at the very base, 5-6 mm; stigma sub- 
capitate. Capsule subcylindric, subdensely hairy 
and lepidote. (I )l.5(-2) by 0.4-0.5 cm, splitting 



into 5 thin, somewhat twisted valves. 

Distr. Malesia: Malay Peninsula (Main Range 
from Mt Kerbau to Mt Batu Puteh; also on Mt 
Tahan). 

Ecol. Mossy forest, on peaty soil, 1370-2135 m, 
apparently rather common but rarely in flower. 
Fl. Feb.-Nov. 

var. calocodon (Ridl.) Sleum. Reinwardtia 5 
(1960) 156. — R. calocodon Ridl. J. Fed. Mai. 
St. Mus. 5 (1914) 38; Fl. Mai. Pen. 2 (1923) 219, 
f. p. 220. 

Corolla laxly covered with rather long wliitish 
hairs outside. 

Distr. Malesia: Malay Peninsula (Main Range 
in Selangor: Mt Mengkuang Lebak, Mt Nuang), 
1465-1770 m. Fl. Feb., May. 

Vern. Lubang pasir, Selangor. 

169. Rhododendron psammogenes Sleum. Rein- 
wardtia 5 (1960) 150. 

Small, erect, terrestrial shrub, up to 1 m. 
Branchlets terete, laxly covered with minute pale 
scales at the tips, glabrous below; internodes 
3-8(-10) cm. Leaves elliptic, broadly attenuate 
towards the apex which is subacuminate or often 
almost rounded and very shortly apiculate, base 
broad-attenuate, more rarely subrotundate, coria- 
ceous, rather stiff", initially lepidote on both faces, 
gradually glabrescent above, subdensely lepidote 
beneath (scales dilutely rufescent, minute, vari- 
ously lobed or dented in the marginal zone, centre 
impressed), slightly revolute at the edge, 2.5-4 
(-5.5) by 1.4-2.8 cm, midrib impressed above, 
stout and prominent beneath in its lower part, 
less prominent distally, nerves 5-6 pairs, irregular, 
rather straight below, — obscurely anastomosing, 
slightly or faintly impressed above, slightly or not 
raised beneath, veins hardly visible; petiole ± 
flattened, thickish, 2-4 mm. Umbels 7-9-flowered. 
Outer perulae ovate, inner ones ovate-oblong to 
subspathulate, membranous, caducously lepidote 
dorsally, or early glabrous, margin fimbriate or 
very shortly irregularly lacerate, up to 1 by 0.5 
cm. Bracteoles linear, laxly lepidote, c. 1 cm. 
Pedicels rather slender, densely clothed with 
thick, variously lobed scales, 2.5-3 cm at anthesis. 
Calyx oblique, patellar, c. 3 mm o, irregularly 
5-dented, teeth normally short (1 mm), but 
sometimes up to 3 mm elongate-triangular and 
acute. Corolla funnel-shaped, subobliquc, white 
or of a pale yellow, thin, 3-3.3 cm long in all, 
densely set with thickish, suborbicular or variously 
dented scales outside margin of the lobes e.xcepted, 
subdensely clad with whitish r spreading hairs 
in the lower half of the corolla tube inside, 
tube cylindric, 1.8-2 cm, at base 3-4 mm 0, 
hardly widened upwards, lobes spreading, 
obovatc-spathulate, shallowly retuse, 1-1.2 by 
0.7-0.8 cm. Stamens 10, unequal, attaining ± 
half the corolla lobes; filaments linear and fc 
densely patently hairy below, much more slender 
and glabrous distally; anthers oblong, c. 2.5 mm. 
Disk glabrous below, shortly yellowish pubescent 
at the upper margin. Ovary subcylindric, densely 



592 



Flora Malesiana 



[ser. I, vol. 6* 



clothed with scales and white, upwards directed 
hairs, c. 5 by 2 mm, gradually attenuate to the style, 
the latter rather slender, laxly patently hairy and 
lepidote in the lower '/a to Y2, glabrous in the 
upper part, as long as or slightly longer than the 
stamens; stigma thick, subturbinate-capitate. 

Distr. Malesia: W. New Guinea (Balim R. 
Valley), once found at 2100 m, amongst bracken 
in unfertile sandy soil. Fl. Dec. 

Note. A similar form, differing in a higher 
lepidote style, which is hairy to Vs only, is found 
in the Lake Habbema region, on sandy soil in 
forest opening. 

170. Rhododendron vitis-idaea SLEim. Reinwardtia 
5 (1960) 156. 

Slender, epiphytic shrub, 0.6-2 m high. Branch- 
lets 1-2 mm 0, tips densely set with stalked 
stellate scales, glabrescent and densely verruculose 
downwards, a little rough to the touch, epilose; 
internodes 2.5-6 cm. Leaves in 3-5-merous 
pseudowhorls at the upper 2-3 nodes, obovate- 
elliptic or obovate, sometimes oblong-obovate, 
apex very broadly attenuate or mostly rounded, 
not apiculate, base cuneate to the petiole, coria- 
ceous, rigid, subdensely or laxly lepidote on both 
faces initially, glabrous and shining above with 
age, lepidote for a long time beneath (scales flat, 
irregularly stellate-incised at the marginal zone, 
centre impressed), finally i glabrescent and set 
with impressed blackish points on the under- 
surface, margin cartilaginous, slightly recurved 
in dry specimens, 1.5-2.5(-3.5) by (0.8-)l-1.5 
(-2) cm, midrib minutely sunk above, obtusely 
prominent beneath, nerves c. 3 pairs, obsolete 
above, somewhat raised and curved-anastomosing 
beneath in full-grown leaves, reticulation 0; 
petiole subterete, lepidote, 2(-3) by 0.8-1 mm. 
Flowers solitary. Ferulae narrow-ovate-acuminate, 
acute, membranous, glabrous dorsally, margin 
short-fimbriate by caducous scales, up to 8 by 4 
mm, p)ersistent for a long time. Pedicels slender, ± 
densely clothed with rufous stellate scales and 
very short spreading hairs, (0.8-) 1-1.5 cm. Calyx 
very obliquely disk-shaped, shortly obtusely 5- 
lobed, c. 2.5 mm 0, lepidote outside. Corolla 
tubular, bright red or scarlet, membranous, tube 
laxly stellate-lepidote and minutely papillose- 
pubescent outside, glabrous inside, somewhat 
curved, 1.5-2 cm, at base 3-4 mm 0, gradually 
dilated to 5-6 mm at apex, lobes ± spreading, 
broad-obovate or suborbicular, 5-7 mm 0. 
Stamens 10, slightly exceeding the corolla tube; 
filaments linear, glabrous; anthers obovate-oblong, 
1.8-2 by 1 mm. Disk prominent, glabrous below, 
pubescent at the upper margin. Ovary subcylindric, 
slightly attenuate at the base, gradually tapering 
to the style, densely shortly patent-hairy and sub- 
densely lepidote, c. 4 by 1.5 mm; style slender, ± 
as long as the stamens, patently hairy in the lower 
half; stigma subcapitate. 

Distr. Malesia: E. New Guinea (Owen Stanley 
Range; Morobe Distr.: Rawlinson Range). 

Ecol. Forest or open hill side, 2195-2590 m. 
Fl. Jan. 



171. Rhododendron rhodostomum Sleum. Rein- 
wardtia 5 (1960) 157. 

Epiphytic shrub, 0.3-0.6 m; branches divaricate. 
Branchlets terete, 1-1.5 mm 0, tips densely set 
with substellate and shortly stalked scales, older 
parts glabrescent and manifestly verruculose, a 
little rough to the touch; internodes 2-4(-6) cm. 
Leaves in 4— 5-merous pseudowhorls at the upper 
l-2(-3) nodes, elliptic or obovate-elliptic, rarely 
broadly obovate-oblong, sometimes obovate, 
apex broad-attenuate, or rarely nearly rounded 
(very tip apiculate or mucronulate by a very 
short thickened gland), base cuneate into the 
petiole, thin-coriaceous, very young ones lepidote 
on both sides, mature ones glabrescent and rather 
dull above, subdensely lepidote for a long time 
beneath (scales variously dented, angular or 
obtusely lobed in the narrow marginal zone, 
thickish and rather large centre impressed, becom- 
ing blackish), edge slightly revolute, 2-3.5 by 
1-1.8(-2.1) cm, midrib slightly impressed above 
in its lower part and raised beneath, nerves sub- 
inconspicuous; petiole flattened, lepidote, 1-3 
(occasionally up to 5) mm. Flowers solitary. 
Ferulae ovate-acuminate, ± mucronulate or 
subulate, membranous, lepidote at the tips 
dorsally, glabrous elsewhere, margin with caducous 
scales and subglandular-fimbriate, up to 12 by 
5 mm. Bracteoles filiform, glabrous, 8 mm. 
Pedicels rather slender, very densely rufous- 
stellate-lepidote, not hairy at all, c. 1 cm. Calyx 
oblique, disk-shaped, lepidote dorsally, very 
shortly obtusely 5-lobed, c. 2.5 mm 0. Corolla 
tubular, fleshy, white, the throat pink, 2.3-2.7 cm 
long in all, subdensely substellate-lepidote at the 
tube outside, glabrous inside, tube cylindrical, 
slightly curved, c. 2 cm, base 6-7 mm 0, ± dilated 
upwards, lobes broad-obovate or suborbicular, 
6-7 mm 0, glabrous, not ciliate. Stamens 10, 
nearly equalling the corolla in length; filaments 
flattened and linear, glabrous, anthers subobovate- 
oblong, 2.5 by 1 mm, cells often contracted at the 
base into a short apiculus. Disk prominent, 
puberulous at the upper margin, glabrous else- 
where. Ovary subcylindric, densely very shortly 
greyish hairy and substellately lepidote, c. 5 by 
2 mm, gradually attenuate to the style, the latter 
rather slender, patently hairy in the lower half, 
glabrous for the rest, ± equalling the corolla in 
length; stigma club-shaped. Immature capsule 
fusiform, 2 by 0.3 cm, dense puberulous and laxly 
lepidote. 

Distr. Malesia: E. New Gumea (Mt Saruwaged 
and Rawlinson Range) . 

Ecol. In mossy bush or mossy bush margin, 
1525-3655 m. Fl. Feb.-March, June. 

172. Rhododendron alternans Sleum. Reinwardtia 
5 (1960) 159. 

Terrestrial or epiphytic shrub, 2-3 m, divari- 
cately branched. Branchlets terete, 2-3 mm 0, tips 
laxly stellate-lepidote, older parts early glabrescent 
and smooth; internodes 4-8 cm. Leaves close 
together in the upper third of the upper intemode, 
i.e. the uppermost ones opposite or in a 3 (-4)- 



Jan. 1966] 



Ericaceae (Sleumer) 



593 



merous pseudowhorl, the other 2 or 3 spiral and 
somewhat distant from each other, elliptic or 
ovate-elliptic, more rarely subobovate- or oblong- 
elliptic, apex shortly and broadly subacuminate, 
or more rarely almost rounded (apical gland 
hardly prominent), base broad-attenuate or 
mostly subrotundate, sometimes subtruncate or 
subcordate, coriaceous, youngest ones subdensely 
lepidote on both faces, fully mature ones glabres- 
cent above, remaining lepidote for a long time 
beneath (scales substellately dented or obtusely 
lobed in the rather wide marginal zone, centre 
minute, slightly impressed), cartilaginous at the 
paler margin, entire or faintly crenulate by im- 
pressed scales, 2.5-3. 5(-4) by 1.2-1.8(-2) cm, 
midrib dilated below, impressed above, prominent 
in the lower half, becoming hardly visible towards 
the apex of the blade beneath, nerves c. 5 pairs, 
spreading, slightly conspicuous above only; 
petiole flattened, 1.5-3 by 1-1.5 mm. Umbels 3-4- 
flowered. Outer perulae ovate, subacuminate or 
obtuse, inner ones ovate-oblong to subspathulate, 
all sericeous dorsally and fimbriate by caducous 
scales, not ciliate, up to 1.8 by 0.7 cm. Bracteoles 
linear, very laxly hairy, up to 1.5 cm. Pedicels 
rather slender, densely clad with rufous substellate 
scales and very short whitish hairs, 1.5-2 cm. 
Calyx oblique, disk-shaped, margin wavy and 
obscurely lobed, lepidote and very shortly hairy, 
c. 2.5 mm 0. Corolla tubular, red, glabrous in- and 
outside, 3-3.5 cm long in all, tube cylindric, 
rather straight, 2-2.5 cm, at the 5-pouched base c. 
4 mm 0, dilated to 6-7 rrun at apex, lobes ± 
spreading, obovate-spathulate or nearly orbicular, 
7-9 mm 0. Stamens 10, unequal, the longest 
nearly as long as the corolla; filaments linear, 
glabrous; anthers oblong, 2.7 by 1 mm, base 
sometimes very shortly biappendicuiate. Disk 
prominent, glabrous. Ovary thick, subovoid- 
conical, densely clothed with rather short and 
subpatent yellowish hairs and with scales between 
these hairs, 4 by 2.5 mm, abrupt with the style, 
the latter slender, quite glabrous, i as long as the 
stamens; stigma subcapitate. Capsule subcylindric, 
attenuate at both ends, 2-2.8 by c. 0.5 cm, sub- 
densely hairy and laxly lepidote, on fruit-pedicel 
c. 2 cm by 1 mm. 

Distr. Malesia: Central Celebes (summit of 
Mt Kambunoj, terrestrial in shrub vegetation, or 
epiphytic in low forest, c. 3000 m, common. Ft. 
July. 

173. Rhododendron leptomorphum Sleum. Rein- 
wardtia 5 (I960) 160. 

Terrestrial shrub, c. 2 m high. Branchlcts 1.5 
mm 0, densely stellatc-lcpidotc at the upper inter- 
nodcs, glabrcsccnt and smooth in the lower parts. 
Leaves in - 4-mcrous pscudowhorls at the upper 
1-2 nodes, narrow-elliptic or oblong-elliptic, apex 
shortly acuminate, subacute, base attenuate to the 
petiole, coriaceous, glabrous and rugulosc above 
at maturity, brown and subdensely lepidote 
beneath (scales flat, irregularly crenatc or sub- 
stellate-lobed in the marginal /one, the rather 
large centre impressed^, cartilaginous margin 



slightly revolute, wavy and subcrenulate, (2-)2.2-3 
by (0.7-)0.8-l cm, midrib very narrowly im- 
pressed below on the upper surface and prominent 
beneath, darker coloured, nerves subinconspicu- 
ous; petiole subterete, lepidote, 3^(-5) by c. 0.8 
mm. Flowers solitary. Pedicels densely stellate- 
lepidote, less densely puberulous, 1.5-2 cm by 
c. 0.6 mm. Calyx disk-shaped, limb membranous, 
reflexed and obscurely dented, c. 4 mm 0. Corolla 
tubular, scarlet, slightly oblique, c. 3 cm long 
in all, glabrous or with several stellate scales at 
its base outside, glabrous inside, tube nearly 
straight, 1.9-2.2 cm, at base slightly saccate, 
c. 6 mm 0, dilated to c. 1 cm at apex, lobes broadly 
obovate-spathulate to suborbicular, 8-10 by 7-9 
mm. Stamens 10, unequal, the longest reaching 
halfway the corolla lobes; filaments linear below, 
filiform distally, glabrous; anthers broad-oblong, 
c. 2.5 by 1.2 mm. Disk prominent, glabrous. 
Ovary subcylindric, densely shortly yellowish 
hairy, no scales visible, 4 by 1.8 mm, ± gradually 
attenuate to the style, the latter stoutish, quite 
glabrous, c. 2.3 cm or slightly exceeding the 
stamens; stigma turbinate-capitate. 

Distr. Malesia: Eastern Central Celebes 
(summit of Mt Lumut), once found at c. 2280 m. 
Fl. Sept. 

174. Rhododendron pseudonitens Sleum. Blumea 1 1 
(1961) 124. — R. nitens Sleum. Reinwardtia 5 
(1960) 162, nan Hutch. 1936. 

Terrestrial shrub, erect-branching or spreading, 
usually less than 1 m, but occasionally attaining 
2 or more m. Branchlets divaricate, stiff, angular 
and subdensely appressedly stellate-lepidote at 
the tips, subterete and glabrescent below; inter- 
nodes 2-6 cm. Leaves spiral and crowded in the 
upper half or third of the upper 1-2 internodes, 
the uppermost leaves forming almost a pseudo- 
whorl, obovate or oblong-obovate, apex very 
broadly and obtusely attenuate or rounded, some- 
times very shortly emarginate, base cuneate to 
the petiole, coriaceous, stiff, glabrous and rugulose 
above, ± dull, smooth and subdensely lepidote 
for a long time beneath (scales small, flat, variously 
stellate-incised at the marginal zone, centre minute, 
impressed, blackish), margin very narrowly 
cartilaginous and revolute, ± entire, 1.5-2.5(-3) 
by 0.8-1.5 cm, midrib impressed above, obtusely, 
thick-prominent in the lower part beneath, nerves 
c. 6 pairs, curved-ascendent and anastomosing, but 
in full mature leaves impressed above, hardly or 
not visible beneath, veins laxly reticulate-impressed 
above; petiole flattened, grooved above, lepidote, 
2-3 by 1-1.5 mm, on a thick leaf-cushion. Umbels 
4-8-flowercd. Perulae ovate-acuminate, apex 
shortly subulate, membranous, lepidote at the 
dorsal mediane, margin set with early caducous 
scales, up to 8 by 4 mm. Bracteoles linear, glabrous, 
c. 8 mm. Pedicels rather slender, subdensely 
clothed with apprcsscd stellate scales, 1.5-2 cm 
by 0.6 mm at anthesis. Calyx oblique, deeply and 
rather irregularly 5-lobcd, lobes narrow-deltoid, 
acute, apprcsscd to the corolla, membranous, 
glabrous, (2-)3-4 mm. Corolla tubular, sub- 



594 



Flora Malesiana 



[ser. I, vol. 6* 



oblique, dark red, 1.5-1.8(-2) cm in all, densely 
stellate-lepidote at the tube outside, glabrous 
inside, tube c. 1.5 cm, base -: dilated, c. 4 mm 0, 
widened to c. 6 mm at apex, lobes oblique, ex- 
panded, suborbicular, c. 5 mm 0. Stamens 10, 
subequal, slightly exceeding the corolla tube; 
filaments linear, glabrous; anthers subquadrate, 
1.5 mm. Disk glabrous. Ovary subcylindric, deeply 
5-grooved longitudinally, densely very shortly 
whitish greyish hairy, with small scales between 
the hairs, c. 4 by 2 mm; style rather abrupt with 
the ovary, nearly as long as the stamens, glabrous 
or with sparse hairs at the very base; stigma 
oblique, capitate, very shortly 5-lobed. Capsule 
thick-fusiform, oblique at the base, puberulous 
and lepidote, rather early glabrescent, 1.5-1.8 by 
0.6 cm. Seeds 2.5 mm including the tailed appen- 
dages. 

Distr. Malesia: New Guinea (Central Distr.: 
Murray Pass, Wharton Range), said to be abun- 
dant in open grasslands or in glades in forest on 
top of range, at 2840 m. Fl. June-Sept. 

175. Rhododendron helodes Sleum. Reinwardtia 5 
(1960) 161. 

Terrestrial shrub, with robust, divaricate 
branches, 0.5 m high. Branchlets terete, c. 3 mm 0, 
densely clad with appressed substellate scales at 
the distal part, glabrescent below; internodes 
3.5-8 cm. Leaves in 4— 6(-8)-merous pseudowhorls 
at the upper 2-3 nodes, oblong or lanceolate- 
oblong, rarely subobovate-oblong in the same 
specimen, apex shortly broad-acuminate, sub- 
acute or bluntish, rarely really blunt, apiculate by 
a gland protruding ± beyond the margin of the 
lamina, base ± broadly and cuneately decurrent 
into the petiole, rigidly coriaceous, laxly impressed- 
lepidote or ± glabrescent above in mature state, 
more distinctly and persistently lepidote beneath 
(scales distant from each other on the intervenium, 
more dense and ± touching each other towards 
the base of the lamina and on the petiole, marginal 
hyaline zone rather large and variously stellate- 
dentate or -lobed, early disappearing, centre 
rather small, much impressed), finally glabrescent 
beneath too leaving lax, impressed, blackish 
points, margin narrow-cartilaginous, irregularly 
crenulate by impressed scales or subentire, (2-) 
2.5-3.5(-4) by (0.8-)l-1.5 cm, midrib impressed 
in the lower 2/3 above, and dilated and obtusely pro- 
minent beneath, gradually disappearing apically, 
2 pairs of nerves suprabasal and high-ascendent, 
2-3 other ones from the midrib, not much con- 
spicuous; petiole much flattened, 3-5 by 2-3 mm. 
Umbels (l-)2-4~f\oweTed. Outer perulae few, short, 
ovate-subulate, inner ones more numerous, 
broadly obovate-spathulate, shortly acuminate, 
subacute, first densely rufescent-lepidote, early 
glabrescent, up to 1.8 by 0.7 cm. Pedicels very 
densely covered with rufous or copper-coloured 
stellate scales, but no hairs, 2-3 cm by c. 1 mm. 
Calyx disk-shaped, c. 5 mm 0, lepidote dorsally, 
the membranous margin reflexed and irregularly 
lobed or lacerate, lobes up to 1 mm and ± 
appressed to the corolla. Corolla tubular, red or 



yellowish red, 3-3.5 cm long in all, subdensely 
stellate-lepidote outside margin of the lobes 
excepted, glabrous inside, tube slightly curved, 
2-2.5 cm, base saccate-dilated, 6-7 mm 0, slightly 
attenuate above, gradually dilated again to 10-12 
mm at apex, lobes erect or somewhat spreading, 
broadly spathulate or suborbicular, very slightly 
emarginate, 0.9-1.1 by 0.7-0.9 cm. Stamens 10, 
unequal, the longest as long as the corolla tube; 
filaments linear, glabrous; anthers obovate, 2 mm. 
Disk prominent, glabrous. Ovary subcylindric, 
rather abrupt with the style, very densely clothed 
with whitish or yellowish, subhirsute hairs, which 
are directed forwards and cover numerous scales, 
c. 8 by 3 mm; style rather slender, c. 2 cm, sur- 
passing the stamens, short-hairy for the lowest 2 
mm, glabrous for the rest; stigma capitate. 

Distr. Malesia: New Guinea (Lake Habbema 
region). 

Ecol. In low shrubberies on long-grass marshes, 
open boggy slopes, locally frequent, at 3225 m. 
Fl. Aug. 

176. Rhododendron papuanum Becc. Malesia 1 

(1878) 201; Sleum. Reinwardtia 5 (1960) 163; 
Blumea 12 (1963) 108; Jahrb. Rdodod. Ges. 
Bremen (1963) f. 31. — ? /?. wrightianum Koord. 
var. piliferum J. J. S. Nova Guinea 12 (1914) 
131.- Fig. 36. 

Much branched, epiphytic or terrestrial shrublet, 
20-50 cm, with fleshy roots. Branchlets 1-2 mm 0, 
the upper 1-2 internodes subdensely set with 
minute rufescent stalked scales, glabrescent and 
minutely verruculose or a little scabrid to the 
touch in the lower parts, finally quite glabrous; 
internodes 2-7 cm. Leaves in 4—6 merous pseudo- 
whorls at the upper 2-3 nodes, obovate, apex 
rounded, rarely slightly retuse, base broad- 
cuneate, coriaceous, glabrous above at full age, 
subdensely lepidote beneath (scales rather small, 
variously angled or substellately lobed at the 
marginal zone, centre dark, small and distinctly 
impressed), margin slightly revolute, (1-)1.2-1.8 
by (0.5-)0.8-l.l(-1.3)cm,midrib impressed above, 
obtusely prominent in the lower Vi-Va beneath, ± 
disappearing distally, nerves obsolete; petiole 
somewhat flattened, lepidote, c. 2 by 0.6-1 mm. 
Umbels (l-)2-3-flowered. Perulae membranous, 
ovate to ovate-oblong, obtuse, glabrous dorsally, 
fimbriate by caducous scales, up to 6 by 3 mm. 
Bracteoles filiform, glabrous, up to 6 mm. Pedicels 
slender, erect, densely clad with shortly stalked 
stellate rusty scales, practically epilose, c. 1.5 cm 
at anthesis, up to 2.5 cm in fruit. Calyx rusty- 
lepidote. c. 2.5 mm 0, irregularly 5-dented, lobes 
1-1.5 (one or two of them up to 2.5) mm, sub- 
acute, spreading or reflexed. Corolla tubular, deep 
pink to purplish red or carmine, glossy, 1 .3-1.5 cm 
long in all, subdensely lepidote outside, glabrous 
inside, tube straight, 0.8-1 cm long, at base c. 3 
mm 0, slightly widened to apex, lobes suborbicular, 
4-5 mm 0, ± spreading at anthesis. Stamens 10, 
c. 1.2 cm; filaments linear, glabrous; anthers 
obovate, 1 mm. Disk low, glabrous. Ovary sub- 
cylindric, densely stellate-lepidote, completely 



Jan. 1966] 



Ericaceae (Sleumer) 



595 




Fig. 36. Rhododendron papuanum Becc. Mt Nettoti, 1 750 m, NW. New Guinea (Van Royen & Sleumer 

7410) (Sleumer, 1961). 



epilose, with 5 deep longitudinal furrows, 4.5 by 
1.5 mm, subabrupt with the style, the lattersiender, 
iepidote at the base, laxly patently hairy for the 
following 2 mm, glabrous for the rest, 6-7 mm 
in all; stigma dilated, obliquely short-obconical. 
Submature capsule subobovoid-fusiform, densely 
Iepidote, 1.8-2.3 by c. 0.4 cm, surmounted by the 
style (8-10 mm). 

Distr. Malesia: New Guinea (Vogelkop 
Peninsula: Arfak, Nettoti and Tamrau Mts, 
possibly also in the Oranje Mts). 

Ecol. Terrestrial in open places on moss- 
cushions, or epiphyte in mossy forest, (1525-) 
1800-2550 m. II. July-Dec. 

177. Rhod(>dcndron acrophilum Merr. & Quisumb. 
Philip. J. Sc. 82 (1953; 333; Sleum. Rcinwardtia 
5 (I960) 163. 

Shrub, I m; branches slender, divaricate. 
Branchlcts 1-2 mm a, terete, covered with flattish, 
stellate scales at the upper 1-2 intcrnodcs, glabrcs- 
ccnt in the lower parts; intcrnodcs 1.5 4 cm. 
Leaves in ± 4-merou8 pseudowhorls at the upper 



3-7 nodes, oblong-elliptic, or sometimes rather 
obovate-elliptic, apex short-acuminate, subacute 
or subobtuse (terminal gland not protruding 
beyond the leaf-margin), base ± broadly cuneate 
to the petiole, coriaceous, young ones sub- 
densely Iepidote on both faces (scales irregularly 
stellate-lobed or -lacerate at the marginal early 
disappearing zone, centre slightly impressed), 
glabrescent above in later stages, more tardily 
so beneath, edge slightly revolute, 2.5-5 by 1-2 
cm, midrib very slightly impressed above, obtusely 
prominent at least in the lower half beneath, 
nerves 6-8 pairs, ascendent, slightly raised on both 
sides; petiole flattened, Iepidote, 1-2.5 by 1-1.5 
mm. Flowers mostly solitary, rarely in twos. 
Pcrulae ovate-oblong to oblong, membranous, 
Iepidote dorsally, especially towards the top, up to 
2 by 0.6 cm. Pedicels stoutish, subdcnscly stellate- 
Icpidote, cpilo.se, r. I cm. CVj/^Aobliquoly thickened, 
disk-shaped, obscurely lobcd, c. 3 mm o. Corolla 
tubular-funncl-shapcd, yellowish white, c. 3 cm 
long in all, 5-lobcd halfway, sparsely stellatc- 
Icpidole all over outside, rather densely whitish 



596 



Flora Malesiana 



[ser. I, vol. 6* 



hairy at the lower 2/3 of the tube inside, tube 
slightly dilated upwards, 4-5 mm at the pouched 
base, lobes erecto-patent, obovate-spathulate, 
c. 1.5 by 1.2 cm. Stamens 10, unequal, 1.7-2 cm; 
filaments linear and ± densely white-hairy in the 
lowest 5-6 mm, glabrescent or glabrous and fili- 
form towards the top; anthers narrow-oblong, 
thecae very shortly appendiculate, 2.7-3 by 0.8-1 
mm. Disk glabrous, prominent. Ovary cylindric, 
rather abrupt with the style, very densely stellate- 
lepidote, c. 5 by 2 mm; style stout, completely 
glabrous or with some scales at the very base, 
1-1.4 cm; stigma turbinate, with 5 rounded lobes. 
Submature capsule subfusiform-cylindric, rather 
densely lepidote, 3.5 by 0.5 cm, on pedicel c. 1 cm 
by 1-1.3 mm. 

Distr. Malesia: Philippines (Palawan), once 
found in mossy forest on top of Mt Mantalinga- 
han, Brooke's Point, at c. 1725 m. Fl. May. 

178. Rhododendron banghamiorum (J. J. S.) Sleum. 
Reinwardtia 5 (1960) 163. — R. pubigermen J. J. S. 
var. banghamiorum J. J. S. in Merr. Contr. Arn. 
Arb. 8 (1934) 123. 

Shrub, c. 1.2 m, slenderly branched. Branchlets 
subterete, laxly lepidote, c. 2 mm 0; internodes 
2-5 cm. Leaves oblanceolate, set in 5-7-merous 
pseudowhorls at the upper 2-5 nodes, apex 
rounded, shortly retuse, base cuneate to the 
petiole, which is winged to nearly the base by the 
decurrent lamina, coriaceous, initially subdensely 
lepidote on both faces, glabrescent above, per- 
sistently lepidote beneath (scales small, marginal 
zone irregularly stellate-lacerate, rather early 
dissolute, centre small, red brown, impressed), 
margin mostly slightly revolute, entire or sub- 
crenulate by impressed scales, 2-3.5 by (0.5-) 
0.6-1 cm, midrib impressed above, obtusely 
prominent in the lower half beneath, becoming 
flat towards the apex of the lamina, nerves and 
reticulation obscure; petiole flattened, 1-2 by c. 
1 mm. Umbels 2-3-flowered, sometimes reduced 
to a solitary flower. Perulae ovate-subacuminate, 
glabrous and shining dorsally, ciliolate, up to 
6 by 4 mm. Bracteoles filiform below, subspathu- 
late distally, glabrous, c. 8 mm. Pedicels rather 
slender, densely stellate-lepidote, epilose, 1.2-1.5 
cm. Calyx shortly cup-shaped, mostly very 
oblique, c. 3 imn 0, densely lepidote, lobes obtuse, 
c. 1 mm. Corolla ± open campanulate, membra- 
nous, rose-red, c. 2 cm long in all, lobed ± half- 
way, laxly lepidote outside, quite glabrous inside, 
tube 3-4 mm at base, gradually widened to c. 
1 cm at apex, lobes obovate-subspathulate, 6-7 
mm wide. Stamens 10, slightly unequal, c. 1.2 cm; 
filaments linear, subdensely db patently hairy 
below; anthers broadly obovate-oblong, c. 2 by 
1.2 mm, obtuse. Disk glabrous. Ovary ovoid- 
conical, ± abrupt with the style, very densely 
lepidote, c. 4 by 2 mm; style slender, glabrous, 
6-8 rtun at anthesis; stigma subturbinate-capi- 
tate. 

Distr. Malesia: N. Sumatra (Gajo Lands: 
Redelong Volcano), c. 1830 m. One collection. Fl. 
Jan. 



179. Rhododendron ripleyi Merr. Not. Nat. Ac. 
Nat. Sc. Philad. 47 (1940) 4; Sleum. Reinwardtia 
5 (1960) 164. 

KEY TO THE VARIETIES 

1. Style quite glabrous. 
2. Disk hairy at the upper margin. Scales light 

coloured 1. var. ripleyi 

2. Disk quite glabrous. Scales dark chestnut 

brown 2. var. cryptogonium 

1. Style with a ring of short patent hairs at its 
very base 3. var. basitrichum 

1. var. ripleyi. 

Shrub, 1.2-2.5 m; branches terete, divaricate. 
Branchlets 1-2 mm 0, the upper 1-2 internodes 
subdensely set with flattish scales, older parts 
quite glabrous, finally covered with a thin grey 
cork; internodes 2-5 cm. Leaves in 4-8-merous 
pseudowhorls at the upper 2-5 (-6) nodes, but 
mostly 1-2 leaves a little lower than the others, 
oblong-obovate to oblong-elliptic, rarely elliptic 
or subobovate-elliptic, apex ± obtuse, or apiculate 
by a protruding obtuse gland, base cuneate to 
broad-attenuate, coriaceous, rather densely lepi- 
dote initially on both sides, glabrescent above with 
age, ± persistently and distally lepidote beneath 
(scales small, of a rather light colour, the thin 
narrow marginal zone irregularly crenate or lobed, 
centre darker, much impressed, a blackish pit in 
old leaves), margin finely crenulate by impressed 
scales and ± distinctly recurved, (2-)2.5-3.5 by 
(0.7-)l-1.5(-1.8) cm, midrib slightly impressed 
above, somewhat prominent but in general rather 
flattish beneath, nerves 4—5 pairs, spreading, im- 
pressed above in old leaves, ± inconspicuous 
beneath, or ± obsolete on both faces; petiole ± 
flattened, lepidote, 2-3 (-5) by 1-2 mm. Umbels 
(l-)2-, sometimes 3(-4)-flowered. Perulae ovate- 
elliptic, short-acuminate, membranous, glabrous 
dorsally, very shortly fimbriate, often persistent 
at the base of the pedicels, up to 1 by 0.5 cm. 
Bracteoles linear, glabrous, up to 1 cm. Pedicels 
stoutish, ± densely appressedly substellate- 
lepidote, epilose, 1-2 cm at anthesis. Calyx 
shortly cup-shaped, margin obtusely shallowly 
5-lobed, c. 3.5 mm 0, lepidote outside. Corolla 
funnelform-campanulate, the base contracted into 
a short and broad tube, 2.5-3 cm long in all, rose- 
red, laxly substellate-lepidote outside, glabrous 
inside, tube c. 1.5 cm long, gradually widened 
from the base (c. 4 mm 0) to 1 cm at apex, lobes 
subpatent, spathulate-obovate to suborbicular, 
0.8-1.3 by 0.6-0.8(-l) cm. Stamens 10 (sometimes 
8), unequal, the longest reaching halfway the 
corolla lobes; filaments linear, ± densely set with 
longish, white, subpatent hairs for c. 2 mm above 
the base, glabrous for the rest; anthers broad- 
oblong, base of each cell somewhat saccate, 2 by 
nearly 1 mm. Disk prominent, short-hairy at the 
upper margin only. Ovary ovoid-conical, abrupt 
with the style, densely stellate-lepidote, epilose, c. 
3.5 by 2 mm; style rather slender, quite glabrous, 
8-1 3 (-17) mm; stigma capitate, ± deeply 5-lobed. 



Jan. 1966] 



Ericaceae (Sleumer) 



597 



Distr. Malesia: N. Sumatra (Gajo Lands: Mt 
Losir). 
Ecol. In mossy jungle, 2500 m. Fl. April. 

2. var. ciTptogonium Sleum. Reinwardtia 5 (1960) 
164. 

Differs by a quite glabrous disk, below sparsely 
to very sparsely hairy filaments and darker scales. 
Leaves as in \ar. ripleyi, but up to 6 by 2.3 cm. 
Flo\\ers cerise red or scarlet. 

Distr. Malesia: N. Sumatra (Gajo Lands), 
1850-3000 m. in ridge scrub, locally common. 
Fl. Jan., Feb. 

Note. The dark chestnut scales suggest a hybrid 
of R. ripleyi with R. malayanum; they are very 
dense in the youngest, not yet full developed 
leaves, touching each other as in R. malayanum. 

3. var. basitrichum Sleum. Reinwardtia 5 (1960) 
164. 

Style with a ring of short patent hairs at its very 
base. Leaves oblong-obovate, 2.3-3.5 by 0.7-1.2 
cm, with a distinct apical gland as in var. ripleyi. 
Corolla scarlet. 

Distr. Malesia: N. Sumatra (Gajo Lands: Mt 
Kemiri), in ridge scrub at 2800 m. Fl. March. 

Note. Possibly a hybrid of R. ripleyi with R. 
pubigermen. 

180. Rhododendron alticolum Sleum. Reinwardtia 

5 (1960) 164. 

Small tree with sparse foliage. Branchlets 
divaricate, nodose, caducously appressed-lepidote 
at the upper internodes, older parts early covered 
with ashy cork; internodes 1.5-2.5 cm long, 
1.5-2.5 mm o. Leavei in 3-4-merous pseudowhorls 
at the upper 1-2 nodes, obovate or elliptic-obovate, 
apex very broadly attenuate and obtuse or not 
rarely rounded, base broadly attenuate to the 
petiole, coriaceous, rather rigid when full-grown, 
dark green above, yellow-green beneath, glabres- 
cent above, rather laxly lepidote beneath for a 
long time (scales small, marginal zone very thin, 
early dissolute, variously dented or lobed, centre 
minute, dark, impressed), edge narrowly cartilagi- 
nous, (2-)2.5-4 by (1.2-)1.5-2.3(-2.7) cm, mid- 
rib narrow and impressed above in the lower, flat 
or slightly raised in the upper part above, as thick 
and prominent as the petiole, gradually less so 
apically, nerves in 6-7(-8) pairs, }, irregularly 
curved-ascendent, anastomosing, faintly impressed 
in old leaves above, a little prominent beneath, 
reticulation lax, visible beneath only; petiole some- 
what flattened, grooved above, 5-7(-9) by 1-1.5 
mm. Umbels (l-)2-3-flowercd. Pcrulac ovate- 
acuminate, laxly lepidote dorsally. Bracteoles 
filiform below, slightly dilated above, c. 1 cm. 
Pedicels rather slender, laxly lepidote, 1.5-2 cm. 
Calyx obliquely patellar, irregularly and shortly 
5-dentcd, c. 4 mm 0. Corolla tubular in general, 
tomewhat funnclform-dilalcd towards the limb, 
5-5.5 cm long m all, apparently red, laxly sub- 
stellate-icpidute at the tube outside, glabrous 
inside, tube slightly curved, c. 4 cm long, at base 
c. 5 mm 0, widened upwards to c. I cm, lobes 



i expanded, obovate-spathulate, c. 1.5 by 1-1.3 
cm. Stamens 10, unequal, the longest attaining 
half the length of the corolla lobes; filaments 
linear and laxly papillose-puberulous below, more 
filiform and quite glabrous distally; anthers 
oblong-obovate, c. 2.5 by 1 mm. Disk very 
prominent, glabrous. Ovary obovoid-cylindric, 
very densely lepidote, c. 6 by 2 mm, rather 
abruptly tapering to the style, the latter rather 
slender, lepidote but at the very base, glabrous for 
the rest, slightly exserted; stigma thick, subcapitate. 
Capsule subcylindric, narrowed towards the base, 
exclusively subdensely lepidote, submature 2.2 
by 0.5-0.6 cm. 

Distr. Malesia: New Guinea (Central Distr.: 
Mt Albert Edward), once found, c. 3600 m. Fl. 
May-July. 

Note. Growths together with R. culminicolum 
var. culminicolum, and differs from it by larger 
corollas and an entirely epilose ovary. Possibly 
only a variety of R. culminicolum or a local hybrid 
of it with another species. 

181. Rhododendron pyrrhophorum Sleum. Rein- 
wardtia 5 (1960) 165. 

Slender, much divaricately branched shrub. 
Branchlets terete, c. 1 mm 0, laxly to subdensely 
set with flat appressed scales but at the ultimate 
internodes, epilose, older parts glabrescent and 
smooth; internodes 2-6 cm. Leaves in 5-7-merous 
pseudowhorls at the upper 1-2 nodes, oblanceolate, 
apex rounded-obtuse, base cuneate and somewhat 
decurrent into the petiole, coriaceous, very young 
ones subdensely but distantly lepidote on both 
faces, mature ones glabrescent, blackish punctulate, 
minutely rugose and ± dull above, subdensely 
persistently lepidote beneath (scales irregularly 
crenate, lobed or shortly incised in the marginal 
chestnut brown zone, centre darker and slightly 
sunk), edge shortly distinctly revolute, 2.5-4 
(-4.5) by 0.5-1 (-1.2) cm, midrib markedly sunk 
above, stoutish and much prominent beneath, 
nerves inconspicuous; petiole flattened, densely 
lepidote, 2-4 by c. 1 mm. Flowers solitary or in 
twos. Perulae ovate-oblong, membranous, glabrous 
dorsally, lustrous, margin with early caducous 
scales, up to 8 by 4 mm. Bracteoles linear-filiform, 
glabrous, c. 6 mm. Pedicels rather slender, 
densely stellate-lepidote, epilose, 1-1.5 mm. 
Calyx patellar, very oblique, obscurely sub- 
acutely 5-lobed, densely lepidote dorsally, 2.5 
mm 0. Corolla campanulate, thin, fiery red, 5-lobed 
halfway, 1.5-1.8 cm long in all, tube at base c. 
4 mm 0, gradually dilated to c. 1.5-2 cm at apex, 
subdensely or more laxly stellate-lepidote outside, 
glabrous inside, lobes broadly obovate-spathulate, 
slightly spreading, 6-8 mm wide. Stamens 10, 
unequal, reaching halfway the corolla lobes; 
filaments linear, slightly dilated at the base, 
becoming filiform distally, quite glabrous; 
anthers elliptic, c 1.8 by 1 mm. Disk prominent, 
glabrous. Ovary ovoid-conical, very densely 
stellate-lepidote. 3 by 1.5 mm, very abrupt with 
the style, the latter rather slender, glabrous, c. 
8 mm; stigma obconical, deeply 5-lobcd. 



598 



Flora Malesiana 



[ser. I, vol. 6* 



Distr. Malesia: N. Sumatra (Gajo Lands: Mt 
Goh Lembuh), once found on forest ridge, 1850 
m. Fl. Feb. 

182. Rhododendron luteosquamatum Sleum. Rein- 
wardtia 5 (1960) 166; Blumea 11 (1961) 125; ibid. 
12 (1963) 108. 

Slender shrub up to 3 m, with stiff, erect, at 
apex divaricate branches. Branchlets 1-2 mm a, 
densely clothed with subsessile, yellowish, sub- 
stellate scales in the young, somewhat flattened 
parts, lower parts terete, glabrescent, brownish 
and a little rough to the touch; internodes (2-)4-7 
cm. Leaves in 3-merous pseudowhorls at the 
upper 4—6 nodes (one or the other leaf occasionally 
on the internode), or opposite, mostly elliptic- 
oblong, or partly subovate- or obovate-elliptic, 
shortly acuminate towards the apex, the very 
apex — obtuse, base attenuate to the petiole, ^ 
coriaceous, medium green above, very pale below, 
in submature state subdensely set with yellow 
scales which are thickened and raised above, 
somewhat impressed beneath, at maturity glabres- 
cent and — smooth above, — persistently lepidote 
beneath (scales rather small, dense but not 
touching each other, markedly impressed, vari- 
ously substellate-dented or-lobed in the thin, early 
caducous marginal zone, centre minute), margin 
very shortly revolute and minutely crenulate by 
impressed scales, (2.3-)2.5-3.5(-4) by (0.8-)l-1.5 
cm, midrib narrowly impressed above, much 
prominent below in the lower, less so in the upper 
part, nerves 4-5 pairs, slightly raised beneath 
only, or subinconspicuous; petiole subterete, 
lepidote, 3-5 by c. 1 mm. Umbels 3-4-flowered. 
Outer perulae ovate, inner ones ovate-oblong, 
dorsally lepidote at the apex, glabrous elsewhere, 
margin set with caducous scales, up to 7 by 4 mm. 
Bracteoles filiform, up to 6 mm. Pedicels stoutish, 
c. 10 mm at anthesis, very densely clothed with 
yellow substellate scales. Calyx patellar, obscurely 
lobed or wavy, densely lepidote externally, c. 3 mm 
0. Corolla tubular, suboblique, c. 10 mm long in 
all, tube at base c. 3 mm 0, hardly dilated distally, 
red or pale pink, very densely yellow-lepidote at 
the tube, laxly so at the lobes outside, glabrous 
inside, lobes 4, erecto-patent, suborbicular, c. 
3.5 mm 0. Stamens 10, c. 8 mm; filaments filiform, 
base dilated, glabrous; anthers obovate-truncate, 
1 mm. Disk glabrous. Ovary subcylindric, very 
densely clad with substellate, yellow scales, 3 by 2 
mm; style abrupt, slender, glabrous, 4 mm; stigma 
subcapitate, shortly 5-lobed. Capsule cylindric, 
very densely lepidote, 1.3-1.5 by 0.5 cm, valves 
finally recurved and twisted. 

Distr. Malesia: E. New Guinea (Morobe 
Distr.: Edie Creek Road, above Wau), 1830-2060 
m, frequent there on land slides respectively old 
gold workings and high, clayey road banks. Fl. fr. 
May-July. 

183. Rhododendron inconspicuum J. J. S. Med. 
Rijksherb. n. 25 (1915) 1; Nova Guinea 12 (1917) 
499; ibid. 12 (1918) t. 194; Kaneh. & Hatus. Hot. 
Mag. Tokyo 56 (1942) 479; Sleum. Reinwardtia 



5 (1960) 167; Blumea 11 (1961) 125; ibid. 12 
(1963) 108. 

Erect, slender, rather compact shrub, 1-3 (-5) 
m, or slender tree, occasionally up to 10 m, much 
divaricately branched; bark pale green. Branchlets 
stoutish, 1.5-2 mm 0, terete, reddish, upper inter- 
nodes densely covered with substellate — stalked 
scales, lower ones glabrescent, minutely warty and 
rough to the touch, early covered with cork; 
internodes 1.5-4 cm. Leaves partly assembled at 
or near the upper 3-4 nodes in 3-merous pseudo- 
whorls, partly scattered in the upper half of the 
internodes, elliptic or ovate-elliptic, more rarely 
ovate, or partly obovate- or oblong-elliptic, apex 
broadly attenuate, obtuse or sometimes rounded, 
base cuneate to rounded, rarely subcordate, 
coriaceous, rather dark green above, much paler 
and often yellowish green or brownish beneath, 
very densely lepidote initially on both faces, early 
glabrescent but rather densely marked with 
blackish minute pits above, more persistently 
and densely lepidote beneath (scales roundish to 
angular, distant, or only partly touching each other 
with the thin, — large marginal zone, which goes 
rather early and is irregularly crenate or shortly 
substellate-lobed, centre dark yellow, i thickish 
and a little impressed), margin subcartilaginous, 
rz distinctly crenulate, slightly or not revolute, 
(l-)1.2-2.4(-3.5) by (0.6-)0.8-1.5(-2) cm, midrib 
narrowly sunk above, much prominent beneath 
in the lower part, less so distally, nerves 4— 6(-8) 
pairs, spreading, faintly impressed above and 
raised beneath in fully mature leaves, often i 
inconspicuous; petiole somewhat compressed 
dorsally, densely lepidote, 1-2 (rarely up to 3) 
mm. Umbels 3-5(-7)-flowered. Perulae ovate, 
outer ones acute or apiculate, inner ones — obtuse, 
thin-membranous, with some scales on the dorsal 
middle line, or quite glabrous and shining, margin 
very shortly irregularly dented or fimbriate (not 
ciliate!), up to 10 by 4 mm. Bracteoles linear 
below, subspathular apically, scantily lepidote, up 
to 1 cm. Pedicels rather slender, very densely set 
with substipitate scales, (8-) 10-1 4 mm. Calyx 
disk-shaped, irregularly, very shortly and obtusely 
5-lobed, densely lepidote dorsally, 2.5-3.5 mm 0. 
Corolla obliquely wide-tubular, 1.1-1.3(-1.6) cm 
long in all, pink, red, or bright crimson, very 
densely covered with thickish, substellate, yellow 
scales outside, glabrous inside, tube rather 
straight, 0.7-1.2 cm long, at base c. 4 mm 0, 
widened to c. 6 mm at apex, lobes — erecto- 
patent, obliquely suborbicular, 3-4.5 mm 0. 
Stamens 10, unequal, ± reaching halfway the 
corolla lobes; filaments filiform, glabrous; anthers 
obovate-subquadrate, 1 mm. Disk low, glabrous. 
Ovary ovoid-conical, very densely set with thicken- 
ed suborbicular scales, 3-4 by 1.5-2 mm, rather 
abruptly tapering to the style, the latter stoutish, 
quite glabrous, ± as long as the ovary; stigma 
subcapitate, deeply 5-lobed. Capsule subfusi- 
form-cylindric, longitudinally 5-ribbed, somewhat 
curved, (1. 2-) 1.5- 1.7 cm by 4-5 mm, densely 
lepidote, crowned by the subpersistent style (4-5 
mm). Seeds c. 3 mm including the fine tail-like 



Jan. 1966] 



Ericaceae (Sleumer) 



599 



appendages. 

Distr. Malesia: New Guinea (Vogelkop 
Peninsula: Arfak Mts; along the Main Range to 
Mt Dayman in Milne Bay Distr.). 

Ecol. In mossy forest or forest openings, or in 
Fflcc//!/H/?i-shrubbery, apparently not rarely in 
woody growth of old landslides, also on edge of 
deep gully or very steep mountain side with grassy 
slopes, shrubs and stunted trees in full sun, often 
on infertile sandy soil, sometimes on limestone 
ridges, locally common, 1800-3400 m. Fl. fr. 
Jan. -Dec. 

Vern. Kanamungar, rohn, Enga: Poio and 
Kepilam. 

Note. Similar in habit to 94. R. yelliotii Warb. 
{subsect. Albovireya) and sharing with that species 
many foliar and floral characters. The main 
difierences between the two species are: in R. 
inconspicuum the scales on the undersurface of 
the mature leaves are — dense, but in general 
distant from, and touching each other but occa- 
sionally or for a small number; the perulae are 
caducously lepidote along the dorsal middle line, 
fimbriate, but not properly pubescent at all, 
mostly quite glabrous and — shining, whilst in 
R. yelliotii the scales are much denser, generally 
all of them touching each other; the perulae are 
lepidote and shortly subsericeous-pubescent dor- 
sally, the margin manifestly ciliate. Much related to 
R. luteosquamaium, which is perhaps only a variety 
of R. inconspicuum. 

184. Rhododendron lamii J. J. S. Nova Guinea 18 
(1936) 96, t. 20, 1; Sleum. Reinwardtia 5 (1960) 
168. 

Lax shrub with long branches, up to 2 m. 
Branchlets terete, 5-6 mm o, upper internodes 
laxly lepidote, lower ones with the thick cushions 
of fallen leaves and an irregularly warty cork; 
internodes 0.8-1.7 cm. Leaves in crowded 6-7- 
merous pseudowhorls, broad-elliptic-obovate, 
apex broadly subacuminate or mostly obtuse to 
rounded, with a thick and somewhat protruding 
gland, base broadly attenuate and decurrent into 
the petiole, subsessile, stiff"-coriaceous, initially 
subdensely lepidote on both faces (scales small, 
marginal zone narrow, irregularly substellate- 
lobed or -crenate, centre slightly or hardly sunk), 
early glabrescent above, less quickly so beneath, 
edge distinctly crenulate, a little or not recurved, 
1.8-3.3 by (0.8-)l-l.8cm, midrib faintly impressed 
above, rather flat beneath, nerves 4-6 pairs, 
erccto-patent, rather obscure on both sides, 
reticulation subinconspicuous; petiole flattened, 
1-2 by 3-4 mm. Umbels 3-4-flowcrcd. Ferulae 
relatively large, membranous, glabrous dorsally, 
outer ones ovate, apiculatc, c. 1 by 0.8 cm, more 
inner ones ovalc-oblong, 2-3.5 by c. 1.5 cm, inner- 
most ones oblong-obovatc or -spathulatc, up to 
5 by 2 cm, not fimbriate. Bractcoles linear, gla- 
brous, up to 2 cm by I mm. Pedicels rather densely 
lepidote, epilosc, c. 1 cm by I mm at anthesis. 
Calyx oblique, disk-shaped or rim-likc, shortly 
jubaculcly 5-lobcd. c. 4 mm a. Corolla funnel- 
shaped, dark red, rather fleshy, glabrous in- and 



outside, 5-lobed nearly halfway, c. 3 cm long in 

all, tube at base c. 4 mm o, gradually dilated to c. 
1 cm at apex, limb spreading to 4.5 cm o, lobes 
obovate-suborbicular, 1.2-1.5 cm o. Stamens 10, 
unequal, slightly exserted; filaments linear, dilated 
above the base, glabrous; anthers obovate, base 
subacutely attenuate into 2 appendages which are 
very close together, c. 3.5 by 1.8 mm. Disk 
prominent, glabrous. Ovary subcylindric, 6 by 
2.5-3 mm, very densely lepidote, tapering to the 
stoutish glabrous style, which somewhat exceeds 
the stamens in length; stigma obconical, dis- 
tinctly lobed. Capsule thick-fusiform, c. 2.5 by 
0.7 cm, on pedicel nearly 3 cm by 2 mm. 

Distr. Malesia: W. New Guinea (Mt Door- 
man), on open slope with rocks, 3280 m. Fl.fr. Oct. 

185. Rhododendron porphyranthes Sleum. Blumea 
12 (1963) 108. 

Weak, erect, 1 -few-stemmed, terrestrial shrublet, 
0.3-0.5 m, few-branched distally. Branchlets 
slender, young ones purplish, laxly lepidote; 
internodes 3-9 cm by 1-2 mm. Leaves 4-6 on 
each of the upper 2-3 internodes, 3 or 4 of them 
laxly arranged in the upper third of the internode, 
1 or 2 near the top, obovate to elongate-obovate, 
apex rounded, sometimes a little emarginate, base 
cuneate to the petiole, immature ones subdensely 
or more laxly lepidote on both faces, mature ones 
thin-coriaceous, shining, early glabrescent above, 
tardily so beneath (scales rather small, flattish, 
marginal zone angular or stellate-dented to various 
degree, centre small and impressed), edge slightly 
revolute and subcrenulate by impressed scales, 
3^.5(-5) by 1.5-2(-2.3) cm, midrib slightly 
grooved in the lower, flat in the upper part above, 
as thick as the petiole at the base, gradually 
decreasing upwards beneath, nerves 3-4 pairs, 
rather obscure, no reticulation; petiole winged in 
the upper part by the decurrent lamina, subterete 
below, (6-)7-10 by c. 1 mm. Umbels 2-flowered. 
Flower bud oblongoid, c. 2 by 0.8 cm; outer 
perulae ovate, distinctly keeled, inner ones obovate 
to spathulate, all shining and laxly lepidote dor- 
sally, fimbriate by early caducous scales, up to 1.2 
by 0.7 cm. Pedicels slender, subdensely substellate- 
lepidote, I cm. Calyx oblique, patellar, wavy, 
c. 2 mm across. Corolla subcampanulate-tubular, 
± zygomorphous, nodding, waxy, magenta in 
fresh specimens, lustrous, quite glabrous, c. 2.3 
cm long in all, tube rather straight, 1.5-1.8 cm, 
at base c. 3 mm o and distinctly 5-pouched, 
dilated to c. 6 mm at apex, lobes erect, broadly 
obovate-spathulatc, 6-8 mm o. Stamens 10, 
reaching halfway the corolla lobes; filaments 
linear, glabrous; anthers subovatc-elliptic, base 
obtuse, c. 2 by 1.5 mm. Di-ik prominent, glabrous. 
Ovary subcylindric, 2.5 by 1.2 mm, tapering to the 
style, the latter rather slender, glabrous. 7-8 mm; 
stigma clavate. 

Distr. Malesia: W. New Guinea (Arfak Mts: 
Mt Sarumot near Iray, Anggi Gigi Lake), one 
collection. 

Ecol. Noihofagus forest at 2320 m, locally rare. 
Fl. Jan. 



600 



Flora Malesiana 



[ser. I, vol. 6^ 



186. Rhododendron simulans Sleum. Reinwardtia 5 
(1960) 168. — R. wrightianum Koord. var. 
ovalifolium J. J. S. Nova Guinea 18 (1936) 94, p.p. 

Shrub, c. 0.7 m; branches rather robust, covered 
with a blackish bark. Branchlets terete, tips 
densely set with stalked scales, lower parts 
glabrescent, rather densely verruculose and some- 
what scabrid, 1.5-2 mm 0; internodes 2-6 cm. 
Leaves crowded at the ultimate 2-3 nodes or in 
3-4 merous pseudowhorls, lanceolate or oblong 
or subobovate-oblong, apex short-acuminate, sub- 
acute, the terminal gland hardly prominent, 
cuneate towards the base in the petiole and ± 
decurrent, the very base often subtruncate, thick- 
coriaceous, stiff, fully mature ones here and there 
with a flat scale above, or mostly quite glabrescent, 
subdensely lepidote beneath (scales variously 
angular or lobed, flat, centre small, impressed in 
minute elevations of the epidermis), margin 
cartilaginous, distinctly revolute especially to- 
wards the base, 1.5-2 by 0.6-1 cm, midrib im- 
pressed above, robust in the lower part and ± as 
thick as the petiole beneath, gradually narrowed 
upwards, nerves inconspicuous; petiole subterete, 
2-3 by c. 1 mm. Umbels 2(-3, very rarely up to 4)- 
flowered. Ferulae membranous, outer ones subu- 
late, inner ones ovate-acuminate, apex subulate- 
attenuate, quite glabrous and glossy, up to 2 by 
0.6 cm. Pedicels stoutish, densely set with rufous 
stellate scales, no hairs, c. 1.5 cm. Calyx obliquely 
disk-shaped, red, very shortly irregularly 5-dented, 
lepidote dorsally, c. 3 mm 0. Corolla tubular 
below, funnel-shaped apically, rather fleshy, bright 
or fiery red, with a scale here and there, or mostly 
glabrous outside, glabrous inside, c. 2.5 cm long 
in all, tube rather straight, 1.6-2 cm long, base 
5-saccate, c. 5 mm 0, widened to 7-10 mm at 
apex, lobes a little spreading, broadly obovate or 
suborbicular, 7-10 mm 0. Stamens 10, subequal, 
2.2-2.4 cm long; filaments linear, c. 1 mm wide 
at the base, glabrous; anthers elliptic, 2-2.5 by 
1 mm. Disk very prominent, glabrous. Ovary 
subovoid-cylindric, very densely rufous-stellate- 
lepidote, ± gradually attenuate to the style, c. 
5 by 2.5 mm; style stoutish, glabrous, c. 1.7 cm; 
stigma thick-obconical, manifestly 5-lobed. Cap- 
sule subcylindric, 2-2.3 by 0.6 cm, the thin valves 
twisted, pedicel c. 2 cm by 1 mm. 

Distr. Malesia: New Guinea (Mt Doorman), 
on steep open slopes with rocks, 3300-3500 m. 
Fl. Oct. 

187. Rhododendron ultimum Wernh. Trans. Linn. 
Soc. H, Bot. 9 (1916) 99; Sleum. Reinwardtia 5 
(1960) 169. 

Small shrub. Branchlets c. 2 mm 0, pruinose 
and subdensely covered with substellate scales at 
the ultimate internodes, glabrescent, with numer- 
ous thick cushions of fallen leaves; internodes 
2.5-6(-8) cm. Leaves rather densely spirally 
arranged along the upper internodes, broad- 
elliptic to elliptic-obovate, apex ± rounded, with 
a small protruding apical gland, base broad- 
attenuate to the petiole, coriaceous, stiff, rather 
laxly to subdensely lepidote initially on both 



sides, early glabrescent above, ± persistently 
lepidote beneath (scales rather small, flat, sub- 
orbicular, marginal zone large, thin, variously 
angled or substeliate-dented or crenate, centre 
small, dark faintly impressed), margin minutely 
crenulate and slightly revolute, (1.8-)2-2.7 by 
(1-)1.4-1.9 cm, midrib slightly impressed above, 
broad and obtusely prominent beneath in the 
lower part, nerves subinconspicuous; petiole sub- 
terete or somewhat flattened, 2-3 (-4) by 1-2 mm. 
Umbels (2-)3-4-flowered. Outer perulae ovate, 
inner ones obovate to spathulate, apex short- 
acuminate, or obtuse, membranous, entirely 
glabrous, outer ones 1-1.5, inner ones 2-2.5 by 
0.8-1.5 cm. Bracteoles linear, glabrous, up to 1.8 
by 1 mm. Pedicels densely substellate-lepidote, 
epilose, 1.2-1.5 cm by 0.8 mm. Calyx obliquely 
disk-shaped, obscurely lobed, lepidote dorsally, 
c. 3 mm 0. Corolla funnel-shaped in outline, 3.5- 
4.5 cm long in all, rose-red, rather fleshy, glabrous 
in- and outside, tube straight, 2-2.5(-3) cm long, 
at base c. 5 mm 0, widened to 9-10 mm at apex, 
lobes broad-obovate to suborbicular, erecto- 
patent, 1.4-1.7 by 1.2-1.3(-1.5) cm. Stamens 10, 
unequal, reaching ± halfway the corolla lobes; 
filaments linear, glabrous; anthers obovate- 
oblong, c. 2.8-3 by 1.3-1.5 mm, each cell ending in 
a short basal apiculus. Disk prominent, glabrous. 
Ovary subcylindric, densely substellate-lepidote, 
c. 6 by 2.5 mm, tapering to the style, the latter 
lepidote at the base, glabrous for the rest, slightly 
longer than the stamens; stigma short-obconical. 
Distr. Malesia: New Guinea (Mt Carstensz), 
3200-4100 m, twice found. 

188. Rhododendron brassii Sleum. Reinwardtia 5 

(1960) 170. 

Lax, suberect shrub, 1-2.5 m. Branches 6-8 mm 
0, greyish corticate. Branchlets subterete, reddish 
and pruinose when dry, smooth, tips laxly clothed 
with small substellate early caducous scales; inter- 
nodes 5-16 cm. Leaves spiral, rather equidistant 
along the upper internodes, elliptic or elliptic- 
or oblong-obovate, apex broadly obtusely attenu- 
ate or rounded, apiculate by a thick ± prominent 
gland, cuneate towards the base and somewhat 
decurrent on the petiole, coriaceous, stiff, appar- 
ently lepidote initially on both faces, very early 
glabrescent and with minute pits above, ± 
persistently and subdensely lepidote beneath 
(scales small, suborbicular, variously substellate- 
angular or -dented in the marginal zone, centre 
minute, slightly impressed), edge cartilaginous 
and distinctly subserrulate-crenulate, 3-4(-5) by 
1.8-2.7 cm, midrib impressed above especially at 
base and obtusely prominent beneath, nerves 6-8 
pairs, ascending at an angle of c. 45°, slightly but 
markedly impressed above, raised or subin- 
conspicuous beneath, veins laxly reticulate, visible 
mostly beneath only; petiole flattened, 3-6 by 
1.5-2 mm. Umbels 3-5(-6)-flowered. Flower buds 
ovoid-oblong, 3.5 by 1.2 cm, perulae membranous, 
quite glabrous, outer ones ovate, obtuse and 
shorter, inner ones ovate-oblong, subacuminate, 
2-3.5 by c. 1.5 cm. Bracteoles linear, c. 2 cm by 



Jan. 1966] 



Ericaceae (Sleumer) 



601 



1-1.5 mm. Pedicels laxly substellate-lepidote 
below, densely to very densely so upwards, epilose, 
c. 2 cm by c. 1 mm. Calyx very obliquely disk- 
shaped, shortly (1 mm) obtusely 5-lobed, lepidote 
dorsally, 4-5 mm 0. Corolla funnelform-tubuiar, 
somewhat oblique and fleshy, orange at the tube, 
yellowish at the lobes, or (greenish) yellow or 
orange suffused with red, glabrous in- and out- 
side, (4-)4.5-5(-5.5) cm long in all, tube c. 3 cm 
long, rather straight, at base a little pouched, 5-6 
mm 0, widened to 1-1.5 cm at apex, lobes erecto- 
patent, broadly obovate-spathulate or suborbicu- 
lar, 1.2-1.5 by 1-1.5 cm. Stamens 10, very une- 
qual, the longest reaching halfway the corolla 
lobes; filaments linear, glabrous; anthers obovate- 
elliptic, 2.5-3 by 1.3-1.5 mm, cells with a basal 
apiculus. Disk prominent, glabrous. Ovary subo- 
void-cylindrical, very densely clothed with rufous 
suborbicular or variously angled and flat scales, 
c. 5 by 2.5 mm, gradually tapering to the style, the 
latter rather slender, laxly lepidote at the base for 
c. 1 cm, glabrous for the rest, shortly exserted 
from the corolla at anthesis; stigma thick-capitate. 
Submature capsule obliquely fusiform-cylindric, 
1.5 by 0.6 cm; pedicel c. 3 cm by 1.5 mm. 

Distr. Malesia: W. New Guinea (Lake 
Habbema vicinity and Wamena R. on N. slope of 
Mt Wilhelmina). 

Ecol. Shrubby forest borders, or grassy edges 
of Podocarpus forest, or open shrubberies on a 
ridge, locally plentiful, 3200-3225 m. Fl. Aug.- 
Sept. 

189. Rhododendron subuliferum Sleum. Reinward- 

tia 5 (1960) 171. 

Epiphytic or terrestrial shrub, 1-2 m. Branches 
terete, c. 5 mm 0, with grey bark. Branchlets c. 2 
mm 0, rather densely clothed with rufous sub- 
stellate and very shortly stalked scales at the upper 
internodes, early glabrescent in the lower parts, 
which are a little rough to the touch; upper inter- 
nodes 1-3, lower ones 3-9 cm. Leaves in 4-6- 
merous pseudowhorls at the upper 1-2 nodes, 
obovate, more rarely oblong-obovate, apex _r 
rounded-obtuse, sometimes minutely emarginate, 
the apical gland subinconspicuous, base cuneate 
to the petiole, coriaceous, rigid, fully developed 
ones glabrous, and densely rugulose above, 
densely lepidote beneath (scales variously angular 
or obtusely substellatc-dented in the thin and 
early caducous marginal zone, small persistent 
centre thickened, reddish brown and slightly 
impressed), margin subcartilaginous, minutely 
crenulate or wavy and subentire, slightly or not 
revolute, 2-3(-3.5, very rarely up to 4) by 1-1.5 
(-2.2) cm, midrib slightly impressed above, 
obtusely thick-prominent in the lower part 
beneath, nerves 3-4 pairs, subinconspicuous; 
petiole lepidote, flattened, 3-4 by 1-1.5 (rarely up 
• o 2) mm. Flowers in twos, rarely solitary. Outer 
pcrulae narrow-subulate, i-1.5 cm long, inner ones 
ovate, tapering into a subulate acumen. 0.8-1.2 by 
0.5-0.6 cm, membranous, glabrous dorsally, 
margm with early caducous scales, often still 
present at anthesis. Bracteules filiform, c. 1 cm. 



Pedicels rather slender, densely clothed with sub- 
stellate scales and very short hairs, 1.2-1.5 cm by 
0.6 mm. Calyx patellar, oblique, very shortly 
irregularly 5-dented and revolute, lepidote and 
pubescent externally, ciliolate, c. 3 mm 0. Corolla 
tubular, (4-)4.5-5(-5.5) cm long in all, very laxly 
and caducously lepidote or quite glabrous out- 
side, glabrous inside, dark red, tube 3-4 cm long, 
straight, at the base with 5 minute pouches, 4-5 
mm 0, gradually dilated to 0.8-1 (-1.3) cm at 
apex, lobes erect, broadly spathulate-obovate or 
suborbicular, 1-1.3(-1.5) by 0.8-1.3 cm. Stamens 
10, unequal, the longest a little exserted from the 
corolla; filaments linear, 1 mm wide at the base, 
glabrous; anthers broad-oblong or obovate-oblong 
c. 3 by 1 mm, cells obtuse or provided with an 
indistinct basal apiculus. Disk prominent, gla- 
brous. Ovary subcylindric, densely substellate- 
lepidote, 6-8 by c. 2 mm, gradually attenuate to the 
style, the latter quite glabrous, as long as or 
slightly longer than the corolla; stigma sub- 
capitate. 

Distr. Malesia: New Guinea (Bele R. Valley, 
NE. of Lake Habbema). 

Ecol. High epiphyte, frequent in midmountain 
forest or, apparently terrestrially, frequent in old 
sandy clearings of mossy forest, at 2300 m. Fi. 
Oct. -Nov. 

190. Rhododendron scarlatinum Sleum. Reinward- 

tia 5 (1960) 168. 

Apparently small and slender shrub; branches 
with grey bark and thick cushions of the fallen 
leaves. Branchlets terete, 1-1.5 mm 0, laxly sub- 
stellate-lepidote and pruinose at the upper inter- 
nodes, lower parts glabrescent and smooth; inter- 
nodes 2.5-5 cm. Leaves partly crowded in pseudo- 
whorls at the upper 1-2 nodes, partly spirally 
arranged along the internodes, elliptic, apex ± 
rounded, not rarely very shortly apiculate, base 
nearly rounded or broadly attenuate, thin- 
coriaceous, firm, laxly lepidote initially, glabrescent 
and a little rugose with age above, laxly and ± 
persistently lepidote beneath (scales flat, sub- 
orbicular, variously angular or irregularly dented 
in the rather wide chestnut coloured marginal 
zone, centre minute, darker, hardly impressed), 
margin subcartilaginous, undulate-crcnulatc or 
subentire, 2-3 by 1-1.8 cm, midrib very narrowly 
impressed above, broader, ± flattened and faintly 
or not prominent beneath, nerves subinconspicu- 
ous; petiole compressed dorsally, 2-3 by 1-2 mm. 
Umbels 3-6-flowered. Bractcolcs filiform, glabrous, 
f. 1 .3 cm. Pedicels subdensely clothed with stellate 
scales and very short patent hairs, 1 .5-2 cm by 0.5 
mm. Calyx disk-like, irregularly and very shortly 
5-dented (but one or the other tooth may be 
elongated), lepidote and hairy dorsally, c. 3 mm 
0. Corolla tubular, scarlet, r. 3.5 cm long in all, 
glabrous in- and outside, tube straight, cylindric, 
f. 2.5 cm long, base 5-pouched, 5 6 mm 0, 
hardly widened upwards, lobes crcclo-patent, 
broadly obovatc-spathulatc, c. I by 0.7-0.8 cm, 
very slightly rctusc, Stamens 10, unequal, 2.7-3.3 
cm; filaments linear, glabrous; anthers oblong- 



602 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 37. Rhododendron atropurpureum Sleum. Mt Wilhelm, E slope at 3600 m, E. New Guinea (Sleumer 

4160) (Sleumer, 1961). 



Jan. 1966] 



Ericaceae (Sleumer) 



603 



obovate or nearly obcordate, 3 by 1.5 mm, cells 
attenuate towards the base, apiculate at the very 
base. Disk prominent, glabrous. Ovary subovoid- 
cylindrical, very densely substellate-lepidote, 
epilose, rather abrupt with the style, the latter 
slender, glabrous, 2.8-3.2 cm; stigma obconical. 
Immature capsule subcylindric, 1.5 by 0.4 cm. 

Distr. Malesia: Central Celebes (Latimodjong 
Range: W. slope of Mt Rante Mario), once found 
in forest, 2950 m. Fl. June. 

191. Rhododendron atropurpureum Sleum. Rein- 
wardtia 5 (1960) 172; Blumea 11 (1961) 125; 
Jahrb. Rhodod. Ges. Bremen (1963) f. 25; Blumea 
12 (1963) 109; ibid. 12 (1964) 340. — Fig. 37. 

Erect shrub up to 3 m. Branchlets c. 2.4 mm 0, 
densely clad with substeliate, shortly stalked 
rufescent scales and very short papillose hairs at 
the upper internodes, older parts glabrescent and 
minutely warty; internodes 3.5-9 cm. Leaves 
spirally arranged in the upper 2/3 of 2 internodes, 
the upper 2-3 leaves — in a pseudowhorl, broad- 
elliptic or ovate-elliptic, sometimes subobovate- 
elliptic, apex broad-attenuate, subacuminate or 
mostly — rounded-obtuse, the thick apical gland 
a little prominent, base rounded or mostly slightly 
cordate, coriaceous, rigid, fully mature ones still 
with the remains of scales, a little rough, finally 
entirely glabrous above, subdensely lepidote 
beneath (scales small, membranous marginal zone 
irregularly crenate or denticulate, early dissolute 
- scales seemingly orbicular then -, centre per- 
sistent, rather large and thick, blackish red and 
markedly impressed), margin indistinctly carti- 
laginous and manifestly crenulate, flat, 1.7-2.6 
(-3) by (l.2-)1.5-2.2 cm, midrib reddish chestnut 
brown and more densely (or for a longer time) 
lepidote than the rest of the upper surface, slightly 
impressed above, robust and obtusely prominent 
beneath, nerves 3-4(-5) pairs, spreading, faintly 
impressed above, slightly raised beneath, often 
subinconspicuous above or rarely so on both 
faces; petiole flattened, densely lepidote, 4—5 by 
1-1.5 mm. Umbels 2-3-flowered. Outer perulae 
ovate, blunt, inner ones obovate, glabrous, 
lepidote-ciliate, 1-1.5 by 0.8-1.2 cm. Pedicels 
very densely clothed with dark rufous stellate- 
incised and shortly stalked scales, no hairs, 1-1.5 
cm by 1 mm. Calyx patellar, shortly obtusely 
5-lobed, 3-4 mm 0, densely lepidote outside. 
Corolla tubular-funnel-shaped, suboblique, red to 
crimson when fresh, becoming magenta in the 
drying process, (3.5-)4— 4.5 cm long in all, densely 
clad with thickish, substeliate scales outside, 
margm of the lobes excepted, glabrous inside, 
tube rather straight, 2.5-3 cm long, base with 5 
small pouches, 5-6 mm 0, widened to 1-1.3 cm 
at apex, lobes • sprcadmg, broad-obovatc or sub- 
orbicular, slightly rctusc, 1-1.5 by 1-1.3 cm. 
Stamens 10, unequal, reaching halfway the 

corolla lobes; filaments linear, 1.5 mm wide at 
base, glabrous; anthers broadly clliptic-oblong, 
3.5-4 by 1.5 mm, each cell bearing a short basal 
apiculus. Disk prominent, glabrous. Ovary ovoid- 
conical, gradually attenuate to the style, exclusively 



very densely substellate-lepidote, 5-6 mm long, 
c. 3 mm at base; style nearly equalling the 
stamens, subdensely to laxly lepidote in its lower 
third, glabrous for the rest; stigma subturbinate- 
capitate. Capsule fusiform-cylindric, shortly atten- 
uate at base, longer so towards apex, a little 
oblique, 2-2.5 by 0.6 cm, densely lepidote. 

Distr. Malesia: E. New Guinea (Eastern High- 
lands: Mt Wilhelm, where it hybridizes with 138. 
R. womersleyi; Western Highlands: Mt Kinkain), 
common here in edge of subalpine forest or alpine 
shrubbery, at 3590-3840 m. Fl. June-Sept. 

Vern. Kage, Chimbu. 

192. Rhododendron buxifolium Low ex Hook./, in 
Hook. Ic. PI. (1852) t. 890; Walp. Ann. 5 (1858) 
146; Staff, Trans. Linn. Soc. II, Bot. 4 (1894) 
197; MiLLAis, Rhodod. (1917) 131; Merr. En. 
Born. (1921) 460; Sleum. Reinwardtia 5 (1960) 
173; Blumea 12 (1963) 109; Jahrb. Deutsch. 
Rhodod. Ges. Bremen (1965) f. 14. 

var. buxifolium. 

Shrub, (1.2-)3-5 m, becoming smaller and 
flat-topped at high altitudes. Branches at the ends 
c. 6 mm 0, terete, with blackish bark. Branchlets 
stoutish, upi>er 1-2 internodes brownish, initially 
laxly clothed with substeliate scales and sometimes 
also with very fine papillose hairs, lower parts 
early glabrescent, lower ones greyish corticate and 
with thick cushions of fallen leaves; internodes 
l-3(-4) cm. Leaves dense, in much approximate 
5-8-merous pseudowhorls at the upper 3-4 nodes, 
broad-elliptic, sometimes subovate- or obovate- 
elliptic, apex broadly attenuate, often rounded- 
obtuse, sometimes slightly emarginate, the apical 
gland rather inconspicuous, base rounded to sub- 
cordate, stifi"-coriaceous, subdensely lepidote on 
both faces initially, rather early glabrescent or 
with minute pits above, more persistently lepidote 
beneath (scales irregularly crenate or lobed in the 
thin, early dissolute marginal zone, persistent 
orbicular centre small, thick, dark brown to 
blackish, rather deeply impressed), edge distinctly 
crenulate, not properly revolute, 1-2 by 0.6-1.2 
cm, midrib narrow and slightly sunk above, 
broad and obtusely prominent beneath, nerves 
5-6 pairs, somewhat irregular, spreading, ± 
joined near the edge, slightly impressed above in 
old leaves, t inconspicuous beneath, veins lax 
and faintly sunk above, obsolete beneath; petiole 
flattened, rugose, 1.5-2.5 by l-l.5(-2) mm. 
Umbels (2-)3-5 (sometimes up to 8)-flowcrcd. 
Outer perulae ovate, the dorsal midrib tapering 
to a short subulate acumen, finely sericeous and 
laxly to subdensely lepidote externally, inner ones 
ovate-obtuse to spathulatc and mostly 1 glabres- 
cent, up to 1.5 by I cm. Bracteoles linear below, 
subspathulatc distally, up to 2 cm by I mm. 
Pedicels ! densely substellate-lepidote, whether or 
not with very sparse short hairs, (I -) 1. 5 -2 (some- 
times up to 2.5) cm by 0.7 mm. Calyx obliquely 
disk-shaped, very shortly irregularly 5-lobed or 
-dented, often ! revolute. 3-4 mm 0. Corolla 
broad-funncl-shapcd or subcampanulate, sub- 



604 



Flora Malesiana 



[ser. I, vol. 6* 



oblique, pink to deep red or rich carmine, waxy, 
scentless, subdensely lepidote outside, glabrous 
inside, 2.5(-3) cm long in all, tube ± straight, c. 
1.5 cm long, at the somewhat pouched base 4—5 
mm 0, widened to ± 1 cm, lobes mostly ± spread- 
ing, obovate-spathulate, 1-1.5 by 1-1.3 cm. 
Stamens 10, subequal, ± equalling the tube; fila- 
ments linear, glabrous; anthers oblong, (2-)2.5-3 
by 1 mm, cells obtuse or mostly narrowed into a 
short, ± distinct basal apiculus. Disk prominent, 
glabrous. Ovary subovoid-conical, apex obtuse 
and much abrupt with the style, very densely 
substellate-lepidote (but occasionally with some 
very sparse hairs on some or other ovary in the 
same specimen), c. 4-5 by 2-3 mm; style glabrous, 
lb as long as the stamens; stigma subclavate- 
capitate. Capsule thickly ovoid-fusiform, oblique, 
1-1 .2 by 0.5-0.6 cm, lepidote, finally ± glabrescent, 
crowned by the rest of the then thick style. Seeds 
c. 2.5 mm including the tails. 

Distr. Malesia: North Borneo (Mt Kinabalu). 

Ecol. In Ericaceous forest above 2745 m, but 
apparently more frequent within shrub vegetation 
in open places on granite amongst rocks, on crests 
of ridges, in fissures up to the very top of Mt 
Kinabalu at 4096 m. Fl. Jan. -Dec. 

var. robustum Sleum. Reinwardtia 5 (1960) 173; 
Blumea 12 (1963) 109. — R. verticillatum {non 
Low ex LiNDL.) Low ex Hook./, in Hook. Ic. PI. 
(1852) t. 884; Walp. Ann. 5 (1858) 447, p.p.; 
Staff, Trans. Linn. Soc. \l. Hot. 4 (1894) 197; 
Merr. En. Born. (1921) 463, p.p. 

A form with larger leaves, 1.8-3 (-3. 5) by 1.2-2 
(-2.5) cm, otherwise not distinct. 

Distr. Malesia: North Borneo (Mt Kinabalu) 
with the normal form. 

Ecol. A constituent of the Ericaceous forest, 
but also in open places (upper limit of scrub 
forest at slanting rock) on granite, sometimes 
together with var. buxifolium, (244O-)3350-3800 m. 

193. Rhododendron wrightianum Koord. Nova 
Guinea 8 (1912) 880; J. J. S. ibid. 12 (1914) 130, 
incl. var. ovalifolium J. J. S. I.e. 131, t. 29B; ibid. 
18 (1936) 94; Sleum. Reinwardtia 5 (1960) 173; 
Blumea 11 (1961) 125. 

KEY TO THE VARIETIES 

1. Ferulae sericeous and i lepidote outside, at 

least in the upper part. 
2. Leaves l-1.5(-2) by 0.5-0.9 (rarely up to 1.5) 
cm. Pedicels lepidote, otherwise whether or not 

papillose 1. var. wrightianum 

2. Leaves (1.3-)2-3(^) by 0.7-1.7 cm. Pedicels 

lepidote and rather densely short-pubescent. 

2. var. cyclopense 

1. Perulae outside laxly lepidote, not hairy at all. 

Leaves 1-2 by 0.6-1. 2(-l. 5) cm. Pedicels 

lepidote, otherwise whether or not very shortly 

pubescent 3. var. insulate 



Branchlets subterete, 1-2 mm 0, densely set with 
shortly stalked, stellate scales at the upper 1-2 
internodes, not hairy, lower parts glabrescent 
and warty by the remaining stalks of the then gone 
scales, finally greyish corticate; internodes (1-) 
2-4(-6) cm. Leaves in 3(-5)-merous pseudowhorls 
at the upper l-3(-4) nodes, obovate-oblong to 
obovate, apex mostly obtuse or rounded, base 
± broadly cuneate, coriaceous, laxly lepidote 
beneath (scales irregularly crenulate or sub- 
stellately dented at the rather narrow marginal, 
early dissolute zone, centre dark, proportionally 
large and somewhat impressed), margin sub- 
crenulate by impressed scales or entire, minutely 
cartilaginous and ± recurved, 1-1. 5 (-2) by 
0.5-0.9 (rarely up to 1.5) cm (in sterile young 
shoots up to 4 by 2 cm), midrib slightly or not 
impressed, rather prominent beneath, nerves 2-3 
spreading pairs, ± obsolete; petiole 1-2 by 0.5-1 
mm. Umbels (l-)2-3-flowered. Outer perulae 
ovate, subacuminate or obtuse, inner ones oblong, 
all manifestly sericeous and laxly lepidote dorsally 
at least in the upper half, and shortly fimbriate, 
often still persistent at flowering time, up to 8 by 
4 mm. Bracteoles filiform, up to 7 mm. Pedicels 
slender, very densely set with ± stellate and shortly 
stalked scales, and mostly with very short papillose 
hairs, 0.5-1.5 cm. Calyx shortly cup-shaped, 
oblique, the 5 lobes or teeth irregular (0.5 mm), 
c. 2.5 mm 0, densely lepidote externally. Corolla 
tubular, dilated upwards, glossy-red to deep 
crimson, thin suboblique, (2.2-)2.5-3 cm long in 
all, laxly to subdensely stellate-Iepidote outside, 
glabrous inside, tube slightly curved, i 2 cm long, 
at base 4-5 mm 0, dilated to 8-10 mm at apex, 
lobes erecto-patent, suborbicular, c. 7 mm 0. 
Stamens 10, unequal, reaching ± halfway the 
corolla lobes; filaments linear, glabrous; anthers 
broad-oblong-elliptic, 2 by 1 mm, the base of each 
theca subapiculate. Disk glabrous. Ovary sub- 
cylindric, densely lepidote, 4-5 by 1.5-2 mm, 
gradually tapering to the style, which is lepidote in 
its lower ^U-^U ^^^ 1.7-2 cm long; stigma sub- 
clavate-capitate. Submature capsule narrow-fusi- 
form, c. 3.5 cm by 3.5 mm. 

Distr. Malesia: New Guinea (Oranje and 
Nassau Mts; Milne Bay Distr.: Mt Garatun). 

Ecol. In mossy forest, occasionally (high) 
epiphyte, in shrubby opening, on sandy or peaty 
soil, 1370-2800 (-3250) m. Fl. Oct.-March. 

2. var. cyclopense J. J. S. Nova Guinea 12 (1914) 
130; Sleum. Reinwardtia 5 (1960) 173; Blumea 12 
(1963) 109. 

Leaves (1.3-)2-3(-4) by 0.7-1.7 cm. Pedicels 
lepidote and rather densely short-pubescent. 
Perulae as in var. wrightianum. 

Distr. Malesia: N. New Guinea (Mt Cycloop). 

Ecol. Epiphytic or on moss-covered soil, in 
± open Nothofagus-Rapanea ridge forest, 1480- 
1800 m. f/.June-Oct. 



1. var. wrightianum. 

Small diffuse, terrestrial or epiphytic shrub, 
0.2-1 m. Branches much divaricate, 2-4 mm 0. 



3. var. insulare Sleum. Reinwardtia 5 (1960) 174. 

Leaves 1-2 by 0.6-1. 2(-l. 5) cm. Perulae laxly 

lepidote dorsally, not hairy at all, up to 8 mm in 



Jan. 1966] 



Ericaceae (Sleumer) 



605 



length. Pedicels lepidote, sometimes furthermore 
laxly and very shortly hairy. 

Distr. Malesia: New Guinea (Milne Bay 
Distr.: Normanby and Goodenough Is.). 

Ecol. Common high epiphyte in tall mossy 
forest, 820-1600 m. Fl. Jan., Oct. 

194. Rhododendron subcrenulatum Sleum. Rein- 
wardtia 5 (1960) 174. 

Erect, terrestrial shrub, 0.3-0.5 m; branches 
terete, 4-5 mm o; bark blackish grey. Branchlets 
divaricate, c. 2 mm o, densely set with substellate 
and distinctly stalked scales at the upper internodes, 
early glabrescent at the lower ones, early greyish 
corticate and minutely warty, a little rough to the 
touch; internodes 2.5-5 cm. Leaves in 3-5-merous 
pseudowhorls at the upper 3-5 nodes, it narrowly 
elliptic or more rarely subovate- or subobovate- 
elliptic, short-acuminate towards the apex, ± 
blunt at the very apex by a thick gland, base 
broad-attenuate or subtruncate-obtuse, very coria- 
ceous, subdensely lepidote on both faces, early 
glabrescent above with age, ± persistently lepidote 
beneath (scales rather small, marginal zone thin, 
variously dented or irregularly crenate, centre 
thickish and much impressed), edge narrowly 
cartilaginous and slightly or not revolute, sub- 
crenulate or entire, 2-2.8(-3) by (0.8-)l-1.4 cm, 
midrib ~ distinctly and narrowly sunk above, 
thick and obtusely prominent in the lower part 
beneath, nerves inconspicuous; petiole densely 
lepidote, a little flattened, 2-3 by 1.5-2 mm. 
Umbels (l-)2-4-flowered. Outer perulae ovate, 
with a 4-5 mm long subulate acumen, more inner 
ones ovate-oblong (acumen c. 1 cm), innermost 
ones subspathulate, obtuse, 1-1.5 by 0.5-0.8 cm, 
lepidote dorsally in the upper partorquite glabrous 
(no sericeous pubescence), margin initially 
fimbriate by scales. Bracteoles linear, c. 1 cm. 
Pedicels very densely clad with rufous stellate scales 
(no hairs), 1.5-2 cm by c. 1 mm. Calyx sub- 
obliquely disk-shaped, shortly obtusely 5-lobed, 
very densely stellate-lepidote, 3-4 mm 0. Corolla 
tubular, suboblique, dark red, 2.5-3 cm long in 
all, subdensely stellate-lepidote outside margin of 
the lobes excepted, glabrous inside, tube rather 
straight, 2-2.3 cm long, base but faintly saccate, 
c. 4 mm 0, dilated to 8-10 mm at apex, lobeserecto- 
patent, suborbicular, 5-7 mm. Stamens 10, un- 
equal, the longest almost as long as the corolla; 
filaments linear, glabrous; anthers broad-oblong, 
2.5 by 1 mm, blunt at the base. Disk glabrous, 
or short pubescent at the upper margin only. 
Ovary subcylindric, very densely stellate-lepidote, 
6-7 by 2.5 mm, gradually attenuate to the style, 
which • equals the stamens and is lepidote in its 
lower Vj-'/j. glabrous for the rest; stigma sub- 
capitate; pedicel 3 cm. 

Distr. Malesia: W. New Guinea (Lake Habbe- 
ma), 3225-3345 m, common on edges of mossy 
forest, or in open mossy thickets, also in thin 
shrubberies on a dry ridge, f'l. Aug. 

195. Rhododendron rubrobractcatum .Si.i ijm. Rein- 
wardlia 5 (l^W)) 175. 



Small, epiphytic or terrestrial shrub, up to 60 
cm. Branchlets 1.5-2 mm 0, laxly substellate- 
lepidote at the upper 1-2 internodes, glabrescent 
and smooth downwards; internodes 2-7 cm. 
Leaves in 3-5-merous pseudowhorls at the upper 
2-3 nodes (with one or two leaves inserted lower 
than the rest), oblong to elliptic oblong, apex 
short-acuminate, subacute or blunt, not rarely 
apiculate by a gland, base broad-attenuate, more 
rarely truncate-subrotundate, coriaceous, minutely 
rugose above in old leaves, with scale remains or 
quite glabrous, rather laxly and more persistently 
lepidote beneath (scales irregularly substellate- 
dented at the thin marginal zone, centre slightly 
impressed), margin minutely cartilaginous and 
paler than the proper lamina, entire, wavy or sub- 
crenulate, hardly revolute, (2-)2.3-3.8 (very 
rarely up to 5) by 1-1.6 cm, midrib ± impressed 
above, obtusely prominent beneath, nerves sub- 
inconspicuous; petiole a little flattened, lepidote, 
3-5(-7) by 1-1.5 mm. Umbels (2-)3(-5)-flowered. 
Perulae ovate-acuminate, attenuate or almost 
subulate into a short acumen, red, membranous, 
rather glossy, glabrous, initially with scales along 
the margin, (l-)1.2-2 by 0.4-0.9 cm. Bracteoles 
filiform, c. 6 mm. Pedicels rather slender, very 
densely rufous-stellate-lepidote, furthermore ± 
densely papillose, 1.3-2 cm. Calyx obliquely 
disk-shaped, obscurely and bluntly 5-dented, 
lepidote dorsally, c. 3 mm 0. Corolla tubular, 
suboblique, red or of a delicate pale red, or pale 
orange-pink, thin, 2-2.6 cm long in all, laxly to 
subdensely stellate-lepidote outside, glabrous in- 
side, tube rather straight or somewhat curved, 
1.6-2 cm long, base 5-pouched c. 5 mm 0, widened 
to 7-8 mm at apex, lobes erecto-patent, broadly 
obovate or suborbicular, slightly retuse, 5-6 mm 
0. Stamens 10, unequal, the longest almost as 
long as or sometimes slightly longer than the 
corolla; filaments linear, glabrous; anthers broad 
oblong, r. 2 mm, base obtuse. Disk glabrous. 
Ovary subcylindric, very densely stellate-lepidote, 
c. 4 by 1.5 mm, -± gradually narrowed to the style, 
which is rather slender, glabrous and nearly 
equals the stamens; stigma subcapitate. Immature 
capsule fusiform, c. 2.2 by 0.35 cm. 

Distr. Malesia: New Guinea (Lake Habbema 
and vicinity). 

Ecol. In open thickets of higher ridges, also 
epiphytic on high trees in mossy forest, 2900-3345 
m. Fl. Aug., Oct. 

Note. A Rhododendron with similar leaves and 
larger corollas (3 cm) was found on Mt Hcllwig. 
1900 m. 

196. Rhododendron calosanthes SttUM. Blumea I I 
(1961) 125. 

.Shrub, c. 40 cm. Branchlets slender, laxly sub- 
stellatc-lcpidote at the youngest, glabrescent at the 
older parts, smooth, I 2 mm o; internodes (1. 5-) 
2-5 cm. Leaves 4--6 on each of the ultimate 2 or 3 
nodes at short intervals, or partly (2 or 3) more 
close together or even in a pscudowhorl. narrowly 
oblong or oblong-elliplic, apex subacutely atten- 
uate or subacuminate, base attenuate to sub- 



606 Flora Malesiana [ser. I, vol. 6* 

cuneate, coriaceous, initially laxly lepidote on both stellate-lepidote at the tube inside, otherwise quite 
faces, tardily glabrescent above, subpersistently glabrous, 3 (-3.5) cm long in all, tube straight, 
lepidote beneath (scales of medium size, variously 2(-2.5) cm long, at base 5-6 mm 0, dilated to 
substellate-dented or -incised at the thin marginal 8-10 mm at apex, lobes erecto-patent, broadly 
zone, centre small, r: impressed), narrowly obovate-oblong, c. 1 by (0.5-)0.6-0.8 cm. Stamens 
marginate, margin subcrenulate by impressed 10, very unequal, the longest hardly exserted from 
scales or ~ entire, 2-2.5 by 0.5-0.9 cm, midrib the corolla tube; filaments linear, glabrous; 
hardly impressed above or even flat, thick and anthers obovate-oblong, base obtuse, 1.5-1.8 by 
obtusely prominent beneath, nerves and veins 1 mm. Disk prominent, glabrous. Ovary sub- 
inconspicuous; petiole 2-3 by 0.7-1 mm. Flowers obliquely cylindric, 4 by 1.5 mm, densely set with 
solitary or mostly in twos, nodding. Ferulae substellate, ferrugineous scales (no hairs), tapering 
membranous, red, glabrous and shining dorsally, to the slender style, which bears some scales at the 
ovate-oblong, outer ones acuminate-apiculate, base and is glabrous for the rest; stigma turbinate, 
inner ones obtuse, up to 2 by 1 cm. Pedicels shortly 5-lobed. 

rather slender, densely stellate-lepidote, 1.5-2 cm. Distr. Malesia: Central New Guinea (Mt 

Calyx obliquely disk-shaped, often included by the Antares, Star Mts). 

basalsaccatepartof the corolla tube, hardly lobed, Ecol. Alpine or semi-alpine shrub vegetation, 

± 2.5 mm 0, lepidote dorsally. Corolla a little 3000-3200 m, rare. Fl. July, 
zygomorphous, red or fiery red, thin, laxly sub- 

7. Series Javanica 

Sleum. Relnwardtia 5 (1960) 176. — Rhododendron sect. Schistanthe Schltr, Bot. Jahrb. 55 (1917) 
\AQ,p.p. — Rhododendron sect. Hapalanthe Schltr, ibid. 55 (1918) 155. — Rhododendron sect. Zygomor- 
phanthe Schltr. I.e. 145, p.p. — Rhododendron subsect. Euvireya Copel./. Philip. J. Sc. 40 (1929) 137, 
159, s. str. — Rhododendron subsect. Leiovireya Copel./. I.e. 137, 167. — Rhododendron subsect. Schizo- 
vireya Sleum. Bot. Jahrb. 74 (1949) 538. — Fig. 38 — 45. 

Leaves rather large, all or at least for their greater number in the same specimen longer than 4 cm, 
mostly much longer, and of various shape. 

Distr. S. Indochina, Malesia. 

KEY TO THE SPECIES 

1 . Corolla outside rather laxly to densely (rarely very densely) short-hairy, otherwise whether or not 

lepidote. 
2. Leaves very shortly soft-hairy beneath at least when young .... 197. R. verticillatum/. velutinum 
2. Leaves without hairs beneath. 
3. Style glabrous or with some spreading hairs at the base or up to the lower third, rather abruptly 

set off against the ovary. 
4. Leaves ± buUate; scales stalked, i.e. subdendroid to dendroid 198. R. rugosum 

4. Leaves not bullate (though the main nerves are markedly impressed above); scales not stalked, flat. 

199. R. keditii 
3. Style pubescent to almost the top, ± gradually tapering from the ovary. 

5. Leaves cordate at the base, subsessile 200. R. christi 

5. Leaves rounded to attenuate at the base, distinctly petiolate. 

6. Corolla 5-6.5 cm long in all. Leaves 10-13 by 5-7 cm 201. R. curviflorum 

6. Corolla up to 4.5 cm long in all. Leaves up to 8.5 by 4.5 cm. 

7. Corolla 3^(-4.5) cm long in all 202. R. villosulum 

7. Corolla c. 2.5 cm long in all 158. R. hatamense 

I. Corolla outside not or practically not hairy (some hairs may be present at the base, cf. 197. R. verti- 
cillatum), otherwise whether or not lepidote. 
8. Ovary glabrous or lepidote, not hairy at all. 
9. Ovary very densely and persistently lepidote. 
10. Filaments quite glabrous. 
11. Anthers obovate or obcordate, i.e. much narrowed towards the base, each cell with a distinct 
basal apiculus. 
1 2. Leaves oblong, entire, apex shortly acuminate, subacute; petiole 0.8-1 .2 cm. 203. R. glabrlflorum 
12. Leaves obovate, subserrulate-crenulate, apex obtuse to rounded, petiole 0.3-0.5 cm. 

204. R. pachycarpon 
11. Anthers broad-oblong, base obtuse. Leaves elliptic, entire, apex subacuminate to rounded; 

petiole 0.4-1 (-1.2 cm) 205. R. celebicum 

10. Filaments distinctly hairy at least in the lower part. 
13. Style with some sparse hairs at the very base, or not hairy at all, otherwise whether or not 
lepidote. 
14. Leaves manifestly reticulate (impressed or prominent) on both faces .... 206. R. sayeri 



Jan. 1966] Ericaceae (Sleumer) 607 

14. Leaves faintly or not reticulate. 

15. Corolla (7-)7.5-8 cm long in all. Anthers 8 nam 207. R. pachystigma 

15. Corolla up to 6 cm long in all. Anthers up to 6.5 mm. 
16. Leaves ovate-lanceolate. Corolla 5.5-6 cm. Anthers narrow-oblong, 5.5-6.5 mm. 

208. R. angulatum 
16. Leaves oblong-elliptic or more rarely elliptic. Corolla 2-3.5 cm. 
17. Calyx shortly cup-shaped, distinctly irregularly dented. Anthers 1-1.5 mm. 

209. R. loerzingii 
17. Calyx disk-shaped, obliquely 5-angular. Anthers 3.5-4 mm. 
18. Leaves in 4-5-merous pseudowhorls at the upper nodes, 6.2-8.3 by 2.4-3.2 cm. 

210. R. impressopunctatum 
18. Leaves scattered along the upper intemodes, 9-12 by 3.5-5 cm . . 211. R. xanthopetalum 
13. Style rather densely set with short spreading hairs (and some scales) in its lower half. 

212. R. hirtolepidotum 
9. Ovary initially laxly to subdensely lepidote, ± early glabrescent during the anthesis, or glabrous 
from the beginning. 
19. Filaments quite glabrous. 

20. Nerves (15-) 17-20 pairs, from the midrib at an almost right angle, straight ± towards the 
margin, curved before the edge. Style (4-)5-6 mm. Leaves spirally arranged. 

213. R. pseudomunidense 

20. Nerves (6-)8-13 pairs, from the midrib at a more acute angle, mostly ascending or curved. 

Style exceeding 6 mm. 

21. Most or all leaves scattered along the intemodes. 

22. Corolla 5-lobed to c. ^/j of its length, orange, salmon, light or brilliant red. Pedicels (1.5-) 

2-3 cm in anthesis, 3-4 cm in fruit 214. R. seranicum 

22. Corolla lobed halfway, yellow. Pedicels 1-1.5 cm at anthesis, 1.5-2 cm in fruit. 

215. R. loboense 
21. Leaves in distinct pseudowhorls, though 1-2 leaves may be inserted up to 1 cm below the rest. 

23. Corolla white 216. R. williamsii 

23. Corolla yellow, salmon or red. 

24. Leaves ± coriaceous. Corolla 3-3.5 cm, yellow to salmon or orange, sometimes light red. 

217. R. robinsonii 
24. Leaves i thick-coriaceous. Corolla 2-2.5 cm, red to dark red or crimson, rarely orange red. 

218. R. rarilepidotum var. rarilepidotum 
19. Filaments ± densely hairy at least in their lower part. 
25. Style longer, often much longer than the ovary, at least 7 mm. 
26. Corolla 1.5-2(-2.5) cm. Leaves ± subcoriaceous, lanceolate or oblong-lanceolate, (3-)4-7 
by (0.7-)l-l.5(-2, very rarely up to 2.5) cm. Capsule 1.8-2.2 by 0.5 cm . . 219. R. multicolor 
26. Corolla longer. Leaves ir stiffly coriaceous and mostly also the capsules larger. 
27. Corolla lobed to 2/3, tube pubescent inside along 5 vertical stripes . . . 220. R. brevitubum 
27. Corolla lobed about halfway, equally short-pubescent, or glabrous inside. 
28. Leaves sessile or nearly so (the base ± truncate-rounded). 
29. Corolla (3-)3.5-4 cm. Capsule 2.5-3 by 0.6-0.7 cm 221. R. sessilifolium 

29. Corolla (5-)5.5-6.5 cm. Capsule (4- )4. 5-5. 5 (-6) by 0.6-0.7 cm .... 222. R. beccarii 
28. Leaves distinctly petioled (the base cuneate to rounded). 

30. Leaves narrow in relation to their length, lanceolate, apex subacutely acuminate, 14-18 
(-23) by (2-)2.5-4(^.5) cm 272. R. rctivenium 

30. Leaves wide in relation to their length, apex subacute to obtuse. 
31. Corolla tube narrow, 3-4(-5) rrun at base, elongate, equalling c. */& of the total length 

of the corolla, which is laxly lepidote outside 223. R. kemulense 

31. Corolla tube wider, 5-8 mm at base, shorter, equalling mostly c. Y2 (rarely up to '/j) of 
the total length of the corolla, which is glabrous outside. 
32. Nerves (1 6-) 1 8-20 pairs, close to each other, ± straight and rectangular from the midrib. 

224. R. moultonii 
32. Nerves 7-10(-13) pairs, more distant from each other and i: curvcd-asccnding. 

33. Corolla (3-)3.5-4 cm. Leaves shortly petioled 221. R. sessilifolium 

33. Corolla generally much longer. Leaves relatively long petioled . . . 225. R. javanicum 
25. Style short, ' as long as the ovary. 

34. Leaves ovate to oblong-ovate; petiole 4-6 by 3 mm 226. R. perplcxum 

34. leaves elliptic to oblong; petiole 12-18 by 3-4 mm 227. R. crasslfolluin 

8. Ovary hairy to various degree, otherwise whether or not lepidote. 
35. Style hairy (or hairy and lepidote) to at least half its length, often so to nearly the top. 
36. Leaves vclutinous all over the undcrsurfacc at least initially, glabrescent to various degrees in 

later stages 197. R. vcrtlclllatum /. vclutinum 

36. Leaves without hairs underneath. 



608 Flora Malesiana [ser. I, vol. 6* 

37. Leaves ± long-attenuate and (sub)acutely (subcaudate-)acuminate at the apex. 
38. Corolla tubular, tube very slightly expanded upwards, 4-5 cm long .... 228. R. rhodopus 
38. Corolla funnel-shaped, i.e. tube short (up to c. 1.5 cm), if longer, manifestly dilated upwards. 
39. Corolla 7.5-9(-10) cm. 
40. Pedicels 2.4-4 cm 229. R. baenitzianum 

40. Pedicels (6-)7-l 1 cm 230. R. englerianum 

39. Corolla up to 6.5 cm. 

41. Corolla lobed to c. 2/3, tube 1-2.1 cm. 

42. Leaves ovate to lanceolate-ovate; petiole rather slender, 1-1.5 mm across. 

231. R. wentianum 

42. Leaves lanceolate to subovate- or oblong-lanceolate; petiole stout, 2-3 mm across. 

232. R. cuspidellum 
41. Corolla lobed up to halfway, tube 3-4 cm. 

43. Leaves acutely attenuate at the base 233. R. commutatum 

43. Leaves rounded or slightly cordate at the base 234. R. subcordatum 

37. Leaves shortly (acutely or mostly rather bluntly) acuminate or attenuate, or obtuse to rounded 

at the apex. 
44. Leaves lanceolate to (sub)oblong-lanceolate. 
45. Anthers (3-)4-5 mm. 
46. Pedicels hairy o«^ lepidote. Petiole (0.4-)0.6-l cm. Corolla (3.7-)4.5-5.5(-6) cm. 

235. R. vanvuurenij 
46. Pedicels lepidote, epilose. Petiole 1.5-2 cm. Corolla 6.5-7 cm .... 228. R. rhodopus 

45. Anthers 2 mm. Pedicels lepidote, epilose 236. R. leptobrachion 

44. Leaves oblong, elliptic or ovate, more rarely subobovate. 
47. Corolla at least 14 cm long in all. 
48. Leaves oblong-elliptic, apex ± obtuse, 13-17 by 6-8 cm. Corolla 5-merous. 

237. R. schlechteri 

48. Leaves ± oblong, apex ± subacuminately attenuate, (18-)20-28 by 6-10.5 cm. Corolla 
7-merous 238. R. leucogigas 

47. Corolla up to 10 cm long in all. 

49. Corolla glabrous (elepidote) or practically so outside. 

50. Corolla c. 2.5 cm long 158. R. hatamense 

50. Corolla longer than 4 cm. 
51. Corolla tubular-funnel-shaped, 4-5 cm. 

52. Leaves 7-10.5 by 2.8-4.4 cm. Anthers exappendiculate 239. R. lompohense 

52. Leaves (7-) 12-1 6 by (4-)5-7.5 cm. Anthers glandular-appendiculate at base. 

240. R. buruense 

51. Corolla tubular, (6.5-)7-7.5 cm 241. R. bloembergenii 

49. Corolla laxly to densely lepidote outside. 
53. Corolla lobed to VS-^ of its total length, tube up to 1 cm long . . . 242. R. macgregoriae 
53. Corolla lobed to V^-V^ of its total length, tube longer than 1.5 cm. 
54. Filaments glabrous, or practically so (very sparse hairs found at the base only). 
55. Leaves ovate to ovate-elliptic, base cordate 243. R. rosendahlii 

55. Leaves elliptic to obovate-oblong, rarely obovate, base broadly attenuate to the petiole. 

268. R. culminicolum 
54. Filaments distinctly hairy at least in the lower third. 

56. Leaves (2.5-)4-8 (rarely in the same specimen in part up to 9) by (1.5-)2-4(-5) cm. 
57. Corolla white or pale pink. 

58. Leaves distinctly petioled. 

59. Inflorescences (8-)9-13-flowered 244. R. comparabile 

59. Inflorescences (3-)4— 5-flowered 245. R. luraluense 

58. Leaves subsessile. Inflorescences (4-)5-7(-9)-flowered .... 131. R. orbiculatum 
57. Corolla coloured otherwise. 
60. Corolla greenish, yellowish greenish, yellow or orange-yellow. 
61 . Corolla greenish or yellowish greenish, 4-5 mm across the base . . 246. R. flavoviride 
61. Corolla yellow or orange-yellow, 7-8(-10) mm across the base. 
62. Corolla widely tubular-subcampanulate. Perulae up to 1.5 by 0.8 cm. 

247. R. christianae 
62. Corolla broadly funnel-shaped. Perulae 3.5(-5) by 1.5(-2) cm . . 248. R. laetum 
60. Corolla pure red. 
63. Reticulation of the leaves dense, subimpressed above, distinctly raised beneath. 

249. R. scabridibracteum 

63. Reticulation of the leaves practically absent 250. R. longiflonim 

56. Leaves (5-)7-15(-18) by (2.5-)3-8 cm. 

64. Pedicels exceeding 6 cm at anthesis at least in part in the same specimen. 



Jan. 1966] Ericaceae (Sleumer) 609 

65. Leaves shortly (rarely up to 1 cm) petioled. Corolla c. 8 cm . . 230. R, englerianum 

65. Leaves petioled for 2-4 cm. Corolla c. 4 cm 251. R. polyanthemum 

64. Pedicels shorter than 6 cm at anthesis at least in the greater part of the same specimen. 

66. Outer and middle perulae up to 2.5 cm long, glabrous, lepidote or finely sericeous and 
lepidote dorsally. 

67. Corolla white or pale pink, tubular to almost salver-shaped. 
68. Leaves petioled for 1.5-2 cm 252. R. kochii 

68. Leaves subsessile 131. R. orbiculatum 

67. Corolla yellow, orange, pale to deep red, ± funnel-shaped. 

69. Leaves sessile or subsessile, petiole l-2(-3) by 3-4 rrun 253. R. brevipes 

69. Leaves r: manifestly petioled. 

70. Leaves thick-coriaceous, stiflF, base ± rounded or often subcordate. 

197. R. verticUlatum /. verticillatum 

70. Leaves subcoriaceous to coriaceous, base usually ± cuneate, sometimes rounded. 

71. Perulae ± densely lepidote in the upper part both in- and outside. Scales on the 

leaves of a scurfy, somewhat detersile type 254. R. aurigeranum 

71. Perulae lepidote and whether or not sericeous outside, mostly early glabrescent, 
glabrous inside. Scales on the leaves not scurfy, though (very) early caducous. 

250. R. longifloruin 

66. Outer and middle perulae distinctly larger, sometimes up to 5 cm long, all over densely 

covered with a rather scabrid, ± appressed tomentum of shorter or longer and ± 

stoutish hairs outside for a long time, glabrescent in later stages. 

72. Corolla a: tubular, densely lepidote outside. Leaves manifestly reticulate at least 

beneath 249. R. scabridibracteum 

72. Corolla ± funnel-shaped, ± laxly lepidote outside. Leaves faintly or not reticulate. 

73. Leaves — coriaceous, broadly to subovate-elliptic, subsessile or shortly (rarely up 

to 7 mm) petioled. Corolla of a pure yellow initially, turning orange or reddish at the 

lobes in later stages; anthers 4-5 mm 248. R. laetum 

73. Leaves thin-coriaceous, elliptic to oblong-elliptic, rarely subovate-elliptic or ovate- 
oblong, generally manifestly (5-15 mm) petioled. Corolla orange, or tube yellow 
and lobes orange-red from the beginning; anthers (5-)6-8 mm . . 255. R. zoelleri 
35. Style hairy at the base, or rarely hairy up to the lower third, or glabrous. 

74. Leaves markedly bullate 256. R. maxwellil 

74. Leaves not bullate. 
75. Leaves narrow-lanceolate, (0.6-)l-2 cm wide. 
76. Leaves (2.7-)5-7.5 cm long, sessile or nearly so, reticulation distinctly finely prominent on 
both faces. Corolla red 257. R. nervulosum var. nervulosum 

76. Leaves 10-15 cm long, petioled for (3-)4-6 mm, reticulation, if any, faintly visible on the 
upper surface only. Corolla yellow or orange 258. R. salicifolium 

75. Leaves lanceolate or of other shape, at least in part in the same specimen wider than 2 cm. 

77. Leaves 4—8 (rarely in the same specimen partly up to 10) cm long. 

78. Style markedly abrupt with the short-conical or obovoid-cylindrical ovary, glabrous or with 
some scattered hairs at the very base. 

79. Leaves sessile or nearly so 259. R. lanceolatum 

79. Leaves manifestly petioled. 
80. Corolla broadly funnel-shaped, lobed at least halfway. 
81. Leaves rounded-obtuse at apex. Perulae up to 8 by 5 mm, dorsally sericeous. 

260. R. sumatranum 

81. Leaves acuminate, the very apex only subobtuse. Perulae up to 2 by 1 cm, quite glabrous 
dorsally 218. R. rarilepidotum var. ootrichum 

80. Corolla tubular or funnel-shaped, lobcd to c. 1/3. 

82. Corolla (2.5)-3-3.5 cm long, quite glabrous outside. Leaves in distinct pseudowhorls. 

261. R. arfakianum 
82. Corolla laxly lepidote outside. 
83. Leaves in distinct pseudowhorls. Corolla (4.5-)5-5.5(-6) cm . . . 262. R. fuchsii 
83. Leaves scattered along the branchlcts. Corolla (3.5-)4-5 cm . . . 263. R. impositum 
78. Style continuous or subabrupt with the ovoid to cylindrical ovary, pubescent up to the 
lower third, or glabrous. 
84. Corolla lobed halfway or more. 
85. Corolla t densely lepidote outside, lobes much often horizontally, spreading. Pedicel 
slender, generally longer than the corolla 242. R. macgrcgorlae 

85. Corolla glabrous outside, lobes crecto-patent. Pedicel rather stout, much shorter than the 
corolla 264. R. Icylcnse 

84. Corolla lobcd less than halfway. 

86. Style reaching hallway the stamens 265. K. brachygynum 



610 Flora Malesiana [ser. I, vol. 6* 

86. Style nearly as long as or longer than the stamens. 

87. Corolla white. Ferulae up to 3.5 by 1.5 cm 266. R. mindanaense 

87. Corolla light to dark red, or orange. Ferulae, as far as known, up to 2 by 1 cm. 
88. Ovary predominantly hairy (tomentose), bearing a few, early caducous scales. Capsule 

hairy, not lepidote 267. R. renschianum 

88. Ovary hairy (even tomentose) awt/ distinctly lepidote (scales, often hidden by the hairs on 
the ovary, persistent and at least well visible on the capsule). 

89. Corolla completely glabrous outside 261. R. arfakianum 

89. Corolla laxly to subdensely lepidote outside 268. R. culminicolum 

77. Leaves all or at least most of them in the same specimen longer than 8 cm. 
90. Corolla laxly to subdensely lepidote outside. 
91. Leaves ± long and acutely (subcaudate-)acuminate. 

92. Leaves lanceolate or oblong-lanceolate, 1.5-3 cm wide 269. R. mollianum 

92. Leaves (2.5-)3.5-6.3 cm wide. 
93. Leaves ovate to lanceolate-ovate; petiole slender, 1-1.5 mm . . . 231. R. wentianum 
93. Leaves lanceolate, sometimes subovate- or oblong-lanceolate; petiole stout, 2-3 mm 0. 

232. R. cuspidellum 
91 . Leaves shortly and acutely or mostly obtusely acuminate or attenuate, or obtuse at all at the 
apex. 
94. Corolla 3.5-6 cm long in all, tube at least 1.6 cm long 
95. Style at least thrice as long as the ovary. 

96. Corolla funnel-shaped 270. R. poremense 

96. Corolla obliquely tubular 268. R. culminicolum 

95. Style + as long as the ovary 265, R. brachygynum 

94. Corolla (1.8-)2-2.8 cm long in all, tube 0.5-1 cm. 

97. Leaves sessile or nearly so 259. R. lanceolatum 

97. Leaves distinctly petioled 242. R. macgregoriae 

90. Corolla quite glabrous outside from the beginning. 
98. Style ± as long as the ovary. 
99. Leaves subsessile. Corolla red 257. R. nervulosum var. exuberans 

99. leaves 1.5 cm petioled. Corolla yellow 265. R. brachygynum 

98. Style much longer than the ovary. 

100. Leaves manifestly cordate 271. R. stresemanni 

100. Leaves rounded (rarely subcordate) or attenuate at the base. 

101. Ovary sparsely and shortly hairy (lens!). Leaves lanceolate to lanceolate-oblong, finely 

and remarkably reticulate on both faces 272. R. retivenium 

101. Ovary subdensely to densely and rather longish white-hairy. 
102. Leaves large, (12-)14-23 by (5-)6-13 cm (leaf index 2-2.5), stiflfly coriaceous; petiole 
stout and flattened, 3-10(-15) by 3-5(-6) mm. 
103. Umbels (6-)7-12-flowered. Pedicels laxly lepidote, epilose. Anthers 5 mm. 273. R. lowii 

103. Umbels 2-flowered. Fedicels densely short-hairy, elepidote. Anther c. 8 mm. 

274. R. intranervatum 
102. Leaves mostly smaller, only (2-)2.5-5 (partly in the same specimen occasionally up to 
9) cm wide (leaf index 3-5, but only 1 if wider than 5 cm), less stiff; petiole mostly more 
slender. 

104. Leaves oblong-ovate, nervation manifestly impressed above . . . 275. R. toxopei 
104. Leaves lanceolate to oblong or elliptic-oblong, sometimes subovate or subobovate- 

oblong, nervation but faintly or not impressed above. 
105. Leaves broadly oblong to subovate- or subelliptic-oblong. Corolla (3-)4— 5(-5.5, 

rarely up to 6) cm 225. R. javanicum 

105. leaves oblong to lanceolate. Corolla mostly 6-7(-8) cm, rarely shorter. 

276. R. brookeanum 

197. Rhododendron verticillatum Low ex Linfdl. J. long and fleshy. Branchlets downy and ± laxly, 

Hort. Soc. Lond. 3 (1848) 86, c. icon. p. 89; early caducously lepidote at the upper internodes; 

Planch. Fl. Serres 5 (1849) 477^80, fig.; Walp. internodes (2-)5-10(-15) cm by 3-^(-5) mm. 

Ann. 5 (1858) 477, p. p.; Millais, Rhodod. Leaves in 4— 6-merous pseudowhorls at the upper 

(1917) 258; Merr. En. Born. (1921) 463, p.p. 2-4 nodes, variable in shape and size in the same 

Sleum. Reinwardtia 5 (1960) 185; Blumea 11 specimen, oblong or elliptic-oblong to elliptic, not 

(1961) 126, non Low^ ex Hook./, in Hook. Ic. PI. rarely subovate- or subcordate-elliptic or -oblong, 

(1852) t. 884. apex subacutely to broadly subobtusely acuminate 

or obtusely attenuate, sometimes even rounded, 

/. verticillatum. base very broadly attenuate or mostly rounded to 

Generally epiphytic shrub; roots reported to be slightly cordate, thick-coriaceous, stiff, lepidote 



Jan. 1966] 



Ericaceae (Sleumer) 



611 



initially on both faces, early glabrescent above 
midrib excepted, remaining laxly lepidote for a 
long time beneath (scales minute, dark, stellate- 
dentate at the marginal zone, centre very small, 
slightly impressed), finely ± glabrescent on the 
intervenium, entire, slightly revolute at the margin 
in dry specimens, (5-)6-10(-l 1) by (2-)3-4 (some- 
times up to 6) cm, midrib broadened above the 
base, narrowed and flat or a little sunk distally, 
as thick and rugose as the petiole, much prominent 
beneath in the lower part, nerves 7-9(-10) curved- 
ascending pairs, somewhat impressed above, 
rather indistinctly raised beneath, often — obsolete 
on both faces; petiole rugose, flattened and groov- 
ed above, lepidote for a long time, (l-)1.5-2(-2.5) 
cm by 2-3 mm. Umbels compact, (8-) 10-1 5- 
flowered. Outer perulae ovate-subacuminate, inner 
ones oblong to spathulate, sericeous and minutely 
lepidote dorsally, lepidote at the :r reflexed apex 
internally, up to 2 by 1.5 cm. Bracteoles filiform 
(0.5 mm) below, subspathulate (up to 7 mm) 
distally, laxly short-hairy, up to 2 cm. Pedicels 
densely to laxly clad with rather long and sub- 
patent hairs, laxly clothed with small caducous 
scales, (3-)3.5-4.5 cm by 1-1.5 mm at anthesis. 
Calyx oblique, patellar, irregularly and shortly 
5-dented (one tooth occasionally more acutely 
acuminate and up to 2 mm long), c. 4 mm 0, 
pubescent outside. Corolla funnel-shaped, reddish 
crimson, red-pink or salmon, rather densely to 
laxly lepidote and occasionally set with some 
scattered hairs at the very base outside, subdensely 
hairy at the tube inside, (4-)5-5.5 cm long in all, 
tube c. 3 cm, base a little pouched, 5 mm 0, 
dilated in its upper third to c. 1 cm, lobes erecto- 
patent, broad-spathulate, 2-2.5 by 1.5-1.8(-2) cm. 
Stamens 10, subequal, attaining ± half the corolla 
lobes, or finally slightly exserted; filaments fili- 
form, rather laxly pubescent in the lower third, 
glabrous for the rest; anthers oblong, curved, 
2.5-3.5 by 1 mm. Disk densely white-pilose at its 
upper margin, glabrous below. Ovary subcylindric- 
conical, rather abrupt with the style, densely clad 
with rufescent subpatent hairs, which cover very 
small scales, 4—5 by 2-2.5 mm; style equalling or 
slightly exceeding the stamens in full anthesis, 
slender, hairy and lepidote for the lower '/j-Vs- 
glabrous above; stigma subcapitate. Immature 
capsule cylindric, 2.4-2.7 by 0.5 cm, patent-hairy 
and lepidote. 

Distr. Malesia: Borneo (Sarawak). 

Ecol. Epiphytic in moss-forest, but also found 
in moss on ground, apparently rare. Ft. Jan., Feb. 

/. velutinum (Becc.) Sleum. Reinwardtia 5 (I960) 
185. — R. velutinum Bfxc. Malesia I (1878) 204; 
Merr. fcn. Born. (1921) 463. 

Leaves with short, soft hairs beneath. Corolla 
besides the scales with short hairs all over the 
tube outside, or at least so at the base. 

Distr. Malesia: Borneo (Sarawak), in forest, 
700-1430 m. 

198. Rhododendron ruftosum Low ex H(K)K. /. in 
Hook. k. I'l. (1852) t. 885; Walp. Ann. 5 (1858) 



446; Staff, Trans. Linn. Soc. II, Bot.4 (1894) 197; 
GiBBS, J. Linn. Soc. Bot. 42 (1914) 104; Millais, 
Rhodod. (1917) 237; Merr. En. Born. (1921) 
463; Sleum. Reinwardtia 5 (1960) 185; Blumea 12 
(1963) 109; ibid. 12 (1964) 340. — R. kinabaluense 
Merr. J. Str. Br. R. As. Soc. n. 76 (1917) 108; En. 
Born. (1921) 461. 

var. rugosum. 

Shrub or small tree, in general knotty and rather 
straggly, 2-5 (-10) m, also epiphytic and pendent. 
Branches robust, divaricately branched. Branchlets 
terete, 2-3 (-4) mm 0, upper internodes with some 
scarious bracts, densely clothed with rufous, 
furfuraceous, deeply stellate-incised and ir stalked, 
early caducous scales, early glabrescent, remaining 
a little warty and rough to the touch, finally with 
greyish cork; internodes (l-)2-5(-8, very rarely 
up to 12) cm. Leaves in 3-5(-6)-merous pseudo- 
whorls, mostly at the upper l-2(-3) nodes, 
broadly ovate-elliptic or elliptic, more rarely 
oblong-elliptic, often variable in shape and size in 
the same pseudowhorl, rather coriaceous, apex 
shortly acuminate, subobtuse, base broadly 
attenuate into the petiole, rarely more obtuse, 
with a dense rufous, almost scurfy, lepidote 
tomentum on both faces initially, early glabrescent 
above, more persistently lepidote beneath (scales 
detersile, not rarely ± dendroid, i.e. small centre 
deepened in middle and extended into a shorter or 
longer foot, which stands within a minute epi- 
dermal depression, marginal zone narrowly 
irregularly stellate-incised, often so to nearly the 
centre), finally glabrescent beneath too, the 
remaining pits manifest, margin entire or faintly 
crenulate, mostly somewhat revolute, (5-)6-10 
(-13) by (2.5)-3-6 cm, midrib rather narrow, the 
subtriangular base excepted, much impressed 
above, as thick as the petiole and very prominent 
in the lower part beneath, nerves (5-)6-8 pairs, 
spreading and i curved, anastomosing before the 
edge, ± deeply impressed above, very prominent 
beneath, lamina thus bullate to various degree, 
veins subdensely reticulate, slightly impressed 
above, ± prominent, but often not much visible 
beneath; petiole subteretc, a little flattened, 
grooved above, 0.7-1.5(-2, rarely up to 2.5) cm 
by 2-3(-4) mm, densely stellatc-lepidote initially, 
rugulose in later stages. Umbels (8-)IO-l2(-l8)- 
flowered, flowers nodding initially. Outer perulae 
ovate-acuminate, firm, apex short-apiculate or 
submucronatc, laxly to subdensely lepidote in the 
upper half dorsally, often pruinose (as are the 
branchlets). inner ones larger, ovate-oblong to 
obovatc, apiculatc, lepidote in the upper, finely 
pubescent in the lower part outside, margin with 
caducous scales, up to 1.5 by 0.8 cm. Bracteoles 
filiform, laxly pubescent and lepidote, up to 2 cm. 
Pedicels rather stout ish, densely pubescent and 
lepidote. (l-)l.3-2.5 cm in anthesis, up to 4.5 cm 
by 1 mm in fruit. CuAx thick, disk-shaped, margin 
wavy and obtusely shortly 5-lobed, pubescent and 
lepidote dorsally, c. 4 mm 0. Corolla sweet-scented, 
tubular below, expanded at the lobes, slightly 
oblique, (2-)2.5-3 (rarely up to 3.5) cm long in 



612 



Flora Malesiana 



[ser. I, vol. 6^ 



all, densely short pubescent and less densely 
lepidote outside, pubescent at the lower half of 
the tube inside, waxy, pink to crimson or scarlet, 
even deep magenta, bright purple or blood-red, 
tube rather straight, (l.2-)1.5-2 cm long, the 
base a little 5-pouched, c. 5 mm 0, gradually 
dilated in the upper third to c. 1 cm at apex lobes, 
oblong-obovate to subspathulate, subretuse, (0.8-) 
1-1.5 by (0.6-)0-8-1.2 cm. Stamens 10, unequal, 
somewhat exserted from the corolla tube; fila- 
ments linear, with short spreading hairs in the 
lower third, glabrous upwards; anthers obovate- 
oblong, base of each cell protracted into a short 
apiculus, (2-)3 by 1 mm. Disk hairy at the upper 
margin, glabrous elsewhere. Ovary subcylindric, 
green-grey-velutinous, the hairs covering small 
scales, rather abrupt with the style, c. 4 by 2 mm; 
style stoutish, attaining half the corolla lobes, 
with some patent hairs at the base or more rarely 
up to the lower third, glabrous for the rest; stigma 
broad-capitate, distinctly lobed. Capsule subovoid- 
cylindrical, very shortly attenuate at the apex and 
base, deeply 5-furrowed, purplish, erect, sub- 
densely short-pubescent, laxly lepidote, 1.3-1.5 
by (0.4-)0. 5-0.6 cm, the valves faintly or not 
twisted when open. Seeds 2-2.5 mm including the 
tails. 

Distr. Malesia: North Borneo (Mts Kinabalu 
andTrus Madi). 

Ecol. In low mossy forest and open exposed 
ridge, also on edge of landslides, in widely different 
habitats from dense shade to full sun, locally 
abundant, (1525-)2100-2900 (rarely up to c. 
3300) m. Fl. fr. Jan.-Dec. 

Note. GiBBS found the lovely heads of waxy 
pink to red flowers one of the most vivid im- 
pressions in her ascent of Mt Kinabalu. 

var. coriifolium (Sleum.) Sleum. Blumea 12 (1964) 
340. — R. coriifolium Sleum. Blumea 12 (1963) 107. 

Leaves thick-coriaceous, stiff, more distinctly 
crenulate than in var. rugosum, apex and base 
obtuse, (2-)2.5^(-5.5) by 1.5-2(-2.6) cm; petiole 
3-4(-5) by 2(-3) mm. Umbels (5-)7-12-flowered. 
Corolla of a deeper red colour than usually found 
in var. rugosum. 

Distr. Malesia: North Borneo (East slope of 
Mt Kinabalu), in rather open low thickets along 
ridges, 3050-3200 m (and possibly still higher), 
rather local. 

Note. Replacing R. rugosum at higher altitudes 
and apparently subject to introgression by R. 
buxifolium var. robustum. 

199. Rhododendron keditii Sleum. Blumea 1 1 
(1961) 126; ibid. 12 (1963) 109. 

Terrestrial shrub. Branchlets robust; internodes 
1.5-3 by 0.3-0.5 cm, subglabrous. Leaves in 3-4- 
merous pseudowhorls at the upper nodes, oblong 
or oblong-elliptic, rarely subobovate-elliptic, apex 
shortly attenuate or subacuminate, — bluntish, 
base broadly attenuate or rounded, coriaceous, 
stiff, glabrous above at full age, subdensely lepidote 
beneath (scales minute, flat, variously dented in 
the thin early dissolute marginal zone, small 



centre much impressed), slightly convex above 
when dry, 6.5-10.5 by 3-6 cm, midrib slightly 
prominent to the base and becoming t flattish 
distally above, thick-prominent in the lower part 
and gradually decreasing upwards beneath, nerves 
6-8 irregular pairs, rather straight below, not 
rarely branched before the margin and anastomos- 
ing, distinctly immersed above in fully mature 
leaves, slightly prominent beneath, reticulation 
lax, faintly raised beneath only, or subobscure; 
petiole semiterete or flattened, rugulose, red, 
glabrous, (1-) 1.2-1. 6 cm long, (3-)4-5 mm wide, 
2-3 mm thick. Umbels ± 10-flowered. Flower buds 
ovoid-subglobular, 2(-2.5) by 1.5-2 cm; outer 
perulae chartaceous, ovate-subrotundate, dis- 
tinctly apiculate, more inner ones larger and 
thinner, broad-elliptic, very shortly apiculate, 
innermost ones oblong-obovate, obtuse, glabrous, 
shining, not fimbriate, l-2(-2.5) by 1-1.5 cm. 
Bracteoles linear below, elongate-subspathulate 
distally, glabrous, 1-1.5 cm by 1-3 mm. Pedicels 
rather slender, red, densely very shortly patent- 
hairy, laxly lepidote especially below the calyx, 
1.6-2.2(-2.5) cm. Calyx patellar, ± oblique, 
indistinctly 5-lobed, puberulous and very laxly 
lepidote dorsally, 2.5-3 mm 0. Corolla broadly 
funnel-shaped, deep pink, subdensely to more 
laxly short-hairy and laxly or not lepidote at the 
tube and the lower part of the lobes outside, hairy 
in the lower third of the tube inside, otherwise 
glabrous, 3-3.5 cm long in all, tube 4—5 mm at 
the slightly saccate base, dilated upwards to 1-1.3 
(-1.5) cm at apex, limb c. 2.5 cm 0, lobes suberect 
or somewhat spreading, ~ obovate, 1-1.5 by 
1-1.3 cm. Stamens 10, slightly exserted from the 
corolla tube; filaments linear and densely very 
shortly hairy below, narrower and glabrous to the 
upper 2/3; anthers broadly obovate-oblong, 2.5-3 
by 1 mm, cells slightly narrowed at the otherwise 
obtuse base. Disk prominent, markedly 10-lobed, 
glabrous. Ovary subconical-cylindric, densely 
short-hairy, not (or very early caducously) 
lepidote, 4-5 by 2-2.5 mm, rather abruptly 
narrowed to the style, the latter thickish, quite 
glabrous, 7-8 mm; stigma turbinate. 

Distr. Malesia: North Borneo (Mt Kinabalu), 
and in Sarawak (Mt Mulu), twice found. 

Ecol. In open sparse scrub on boulders or 
landslides, 1920-2010 m. Fl. Aug., fr. Dec. 

200. Rhododendron christi Foerster in Fedde, 
Rep. 13 (1914) 222; Schltr, Bot. Jahrb. 55 (1918) 
155, incl. var. loniceroides Schltr, I.e.; Diels, 
ibid. 62 (1929) 487; Sleum. Reinwardtia 5 (1960) 
185; Blumea 11 (1961) 127; ibid. 12(1963) 110.— 
Rhododendron sp., allied to R. wentianum, Stonor, 
Rhodod. Yearbook 6 (1951) 50, f. 54. 

Small, stiff, sparsely branched shrub, 0.3-1.2 m, 
terrestrial or epiphytic, with long virgate branches. 
Branchlets terete, 1.5-2 mm 0, upper internodes 
rather laxly lepidote, lower ones glabrescent, 
smooth, early covered with whitish cork; inter- 
nodes 4-9(-ll) cm. Leaves in 3-merous pseudo- 
whorls at the upper 2 or 3 nodes, ovate, subsessile, 
apex shortly subabruptly acutely acuminate, base 



Jan. 1966] 



Ericaceae (Sleumer) 



613 



cordate, (thin-)coriaceous, laxly lepidote initially, 
glabrescent above, laxly and ± persistently lepidote 
beneath (scales flat, small, thin marginal zone 
rather large, irregularly and — deeply dented or 
incised, darker centre small, slightly impressed), 
somewhat recurved at the margin in dry specimens, 
(4-)5-9(-ll) by 2.5-6.5 cm. midrib narrow and 
deeply grooved above, rather robust and prominent 
beneath, nerves 4-6 pairs, curved-ascending and 
joined, faintly or not impressed above, slightly 
prominent beneath, veins and veinlets rather 
densely reticulate, slightly raised on both faces; 
petiole flattened, l-2(-3) by 1-2 mm, lepidote. 
Umbels (2-)3-4-flowered. Outer perulae ovate, 
apex abruptly subulate, small, glabrous dorsally, 
inner ones larger, ovate, apiculate or obtuse, 
sericeous in the upper part externally, margin 
minutely erose, up to 2 by 0.8 cm. Bracteoles linear 
below, subspathulate-dilated (3-5 mm) towards 
the apex, laxly pubescent, up to 2 cm. Pedicels 
rather stoutish, red, very densely patently short- 
hairy, with few or no scales, (1.5-)2-2.5 cm at 
anthesis. Calyx disk-shaped, hairy, 3-4 mm 0, 
with 5 short (1 mm) obtuse ciliate lobes (of which 
some or other occasionally extends to 5 by 
1 mm). Corolla tubular below, expanded and 
oblique at the lobes, (3-)3.5-4(-4.5) cm long in all, 
at the basal part yellow or greenish yellow, 
otherwise red or orange-red, i densely clothed 
with rather longish hairs in- and outside which are 
whitish below, becoming reddish to purplish 
towards the lobes, no scales, tube slightly curved, 
2-2.5 cm, base c. 6 mm 0, slightly dilated to 8-10 
mm at apex, lobes obovate-spathulate or sub- 
orbicular, 1.3-1.8 by 1-1.5(-1.8) cm. Stamens 10, 
unequal, slightly exserted; filaments narrow-linear, 
subdensely patently hairy in the lower I4-V3, 
glabrous for the rest; anthers obovate, apex 
truncate, 2-2.5 mm. Disk densely pubescent in 
its upper half. Ovary cylindric, continuous with 
the style, short-tomentose, 5-7 by 2.5-3 mm; 
style stout, i densely hairy to nearly the top, ± 
as long as the stamens; stigma oblique, subcapitate. 
Capsule subcylindric, widest in the upper third, 
slightly curved, laxly pubescent, 3-3.5 by c. 0.5 
cm, valves thin, ± twisted. 

Distr. Malesia: E. New Guinea (Mt Saruwaged 
and vicinity; Eastern, Western and Southern 
Highlands; Wharton Range). 

Ecol. Epiphyte in mossy bush and on grassland 
trccferns, or terrestrial in light Quercus-Engel- 
hardia forest, out of rock faces on clifl"s at the 
moss forest level, or in scrubby summit growth, 
1 20O-2800(-3000, perhaps still higher up) m. 
Fl. May-Oct. 

Vcrn. Kuswamp, Mcndi. 

201. Rhododendron curviHorum J. J. S. Bull. Jard. 
Bot. Bt/g II, « (1912) 50; Nova Guinea 12 (1914) 
141. t. 36 B; SLhUM. Rcinwardtia 5 (I960) 186. 
Large terrestrial or epiphytic shrub, up to 2.5 m. 
Branchicts subtcrctc, 2.5-4 mm 0, tips subdensely 
substcllatc-lcpidotc, older parts early glabrescent; 
inlcrnudcs 4-17 cm. Leaves in 5-mcrous pseudo- 
whorls at the upper 1-2 nodes, oblong- to ovate- 



elliptic, apex short acuminate, subacute, base ± 
rounded, stiff"-coriaceous, at full age quite glabrous 
above, subdensely lepidote beneath (scales small, 
irregularly and ± deeply stellate-incised at the 
pale marginal zone, centre dark, finally blackish, 
faintly impressed), edge not or but little revolute, 
10-13 by (4.7-)5-7 cm, midrib narrowly impressed 
above, continuing the robust petiole beneath, 
nerves 5-8 pairs, spreading, straight below, curved 
upwards and anastomosing, slightly impressed 
above, prominent beneath, reticulation rather 
dense, faintly impressed or raised above, much 
more distinct and prominent beneath; i>etiole 
somewhat flattened, grooved above, glabrescent, 
reddish when dry, 0.7-1.7 cm by 2-3 mm. Umbels 
5-9-flowered. Outer perulae small (± 1 cm), 
ovate, apiculate, inner ones gradually larger, 
oblong to spathulate, sericeous outside, glabrous 
inside, 2.5-4 by 1.5 cm. Bracteoles linear below 
(0.5 mm), spathulate distally (4 mm), 2.8-3.5 cm. 
Pedicels short-tomentose, elepidote, 2-3.3 cm by 
nearly 1 mm. Calyx disk-shaped, tomentose 
dorsally, 4-6 mm 0, lobes short-deltoid (1 mm), 
obtuse, ± reflexed at full anthesis. Corolla tubular, 
± oblique, unequally 5-lobed, lilac (type) or 
yellow (Brass 11536), (5-)5.5-6.5(-7) cm long in 
all, densely rather shortly pubescent outside 
margin of the lobes excepted, subdensely pubes- 
cent in the lower half of the tube inside, elepidote, 
tube curved, 3-4 cm, at the 5-pouched base 6-7 
mm 0, widened to 1-1 .2 cm at apex, lobes obovate- 
spathulate to suborbicular, 2-2.5(-3) by 1-1.5 
(-1.8) cm. Stamens 10, unequal, the longest 
reaching halfway the corolla lobes; filaments 
linear, densely to sparsely pubescent below, more 
slender and glabrous towards the top; anthers 
broad-oblong to obovate-oblong, c. 4 by 1.8-2 
mm, base obtuse. Disk prominent, densely pubes- 
cent above, less so below. Ova/->' elongate-conical, 
tomentose, 8-10 by c. 3 mm, continuous with the 
style, the latter ± as long as the shortest stamens, 
densely pubescent to nearly the top; stigma thick, 
capitate, much lobed. 

Distr. Malesia: W. New Guinea (Keyts Mts 
and Bele R.). 

Ecol. Epiphytic or terrestrial in secondary 
forest or in low serai growths on landslip, 1900- 
2200 m. Fl. Nov.-Dec. 

Note. The size of the corolla in Smith's 
description of 1914 differs from that in t. 36 B; 
unfortunately, the type material, conserved in 
formaline at Bogor, is practically useless. I have 
drawn a new description of it based on good and 
apparently conspecific specimens (Brass 11215 
and 11536) from Bclc R., which have yellow 
flowers instead of 'lilac' as said of the type 
material, which came from the Keyts Mts. Also 
the description of the perulae, not given in the 
original diagnosis, is made after the Brass collec- 
tions. 

202. Rhododendron villosulum J. J. S. Med. Rijks- 
herb. /(. 25 (191 5) 5; Nova Guinea 12 (1917) 507; 
ibid. 12 (1918) I. 201; ibid. 18 (1936) 94; Si.eum. 
Rcinwardtia 5 (1960) 186; Blumca 11 (196!) 127. 



614 



Flora Malesiana 



[ser. I, vol. 6* 



— R. franssenianum J. J. S. Med. Rijksherb. n. 25 
(1915) 5; Nova Guinea 12 (1917) 508; ibid. 12 
(1918) t. 202. 

Shrub, c. 1 m, with apparently weak, sub- 
pendent long branches. Branchlets terete, 2.5-4 
mm 0, upper internodes subdensely clothed with 
substellate brownish, rather early caducous scales, 
lower parts giabrescent and smooth; internodes 
(4-)8-18 cm. Leaves in 3(-5)-merous pseudo- 
whorls at the upper 1-2 nodes, unequal in size, 
elliptic or ovate-elliptic, sometimes ovate-lanceo- 
late, apex shortly to long (1-2 cm) acuminate, 
often a little curved, subacute to very acute, 
base broadly attenuate to rounded, coriaceous to 
subcoriaceous, completely glabrous above with 
age, ± persistently and subdensely lepidote 
beneath (scales rather small, flat, marginal zone 
variously and irregularly substellate-dented or 
-lobed, dark centre slightly impressed), edge 
slightly or not recurved, (3-)3.5-8.5 by (2-) 
2.5-4.5 cm, midrib narrow and much impressed 
above, rather robust and prominent, nerves 
(4-)5-8(-9) pairs, irregular, spreading and anas- 
tomosing, faintly or not impressed above and 
raised beneath, reticulation rather dense and 
prominent on both faces; petiole a little flattened 
and grooved above, lepidote, (2-)3-5 by 1-1.5 mm. 
Umbels (2-)3-6-flowered. Flower buds ovoid, c. 
2.5-3 by 1.5 cm; outer perulae broad-ovate, apex 
abruptly narrowed, apiculate, subglabrous, inner 
ones larger, ovate to obovate-spathulate, obtuse, 

— densely sericeous dorsally especially in the 
upper half, up to 2.5 by 1 cm. Bracteoles linear, 
pubescent, up to 2 cm by 1 mm. Pedicels very 
densely patent-pubescent, elepidote, (1.5-)2-3 cm 
by c. 1 mm at full anthesis. Calyx disk-shaped, 
obliquely 5-angular or shortly 5-dented, densely 
hairy or nearly villous, c. 4 mm 0. Corolla tubular 
below, expanded towards the limb, oblique, dark 
to light red, 3-4(-4.5) cm long in all, more laxly 
to densely hairy all over, outside margin of the 
lobes excepted, and at the lower half of the tube 
inside, elepidote, tube curved, 2-2.5 cm, at the 
5-pouched base 4—6 mm 0, widened to 7-10 mm 
at apex, lobes broad-obovate to suborbicular, 
slightly retuse, 1.2-1.5(-1.8) by 0.8-1. 2(-l. 6) cm. 
Stamens 10, very unequal, the longest reaching 
nearly halfway the corolla lobes; filaments linear 
and pubescent in the lower, nearly filiform and 
glabrous in the upper half; anthers obovate, apex 
obliquely truncate, 2.5-3 by c. 1.5 mm. Disk 
densely pubescent at the upper margin only. 
Ovary subovoid-cylindric, continuous with the 
style, whitish tomentose, apparently elepidote, 6-7 
by 2.5-3 mm; style — as long as the stamens, 
densely hairy below, less so towards the top; 
stigma broad-capitate, distinctly lobed. 

Distr. Malesia: New Guinea (Wissel Lake 
region, Oranje and Nassau Mts, Eastern High- 
lands). 

Ecol. Generally epiphytic in moss forest, 
(1755-)2400-3000 m. 

Note. Certainly much related to R. christi, 
distinguished mainly by the non-cordate, more 
distinctly petioled leaves. 



203. Rhododendron glabriflonim J. J. S. Med. 

Rijksherb. n. 25 (1915) 6; Nova Guinea 12 (1917) 
509; ibid. 12 (1918) t. 200, b-f, t. 204 a; Sleum. 
Reinwardtia 5 (1960) 186. 

Shrub. Branchlets terete, c. 4 mm 0, upper 2 
internodes subdensely lepidote, pruinose, minutely 
lenticellate or verruculose, older parts giabrescent; 
internodes 4-5 cm. Leaves in 6-7-merous lax 
pseudowhorls at the upper 2 nodes, oblong, 
apex short-acuminate, subacute, base broad- 
cuneate into the petiole, coriaceous, stiff", glabrous 
and finally rugulose above, rather laxly lepidote 
beneath (scales small, flat, thin marginal zone 
relatively large, irregularly angled or substellate- 
lobed or -dented, dark centre minute, hardly im- 
pressed), margin narrowly cartilaginous, faintly 
revolute, 6-9 by 2-4 cm, midrib narrow and rather 
deeply impressed above, robust and much promi- 
nent (being the continuation of the thick petiole) 
in the lower part beneath, gradually decreasing 
towards the apex, nerves 8-10 pairs, the lowest 
from the base of the blade, steeply ascending, 
anastomosing, slightly impressed above, raised 
beneath, reticulation rather dense, not or but 
faintly impressed above, distinct and prominent 
beneath; petiole a little flattened, grooved above, 
lepidote, 0.8-1.2 cm by 2-2.5 mm. Umbels ^z 
9-flowered. Pedicels stoutish, very densely lepidote, 
epilose, 1.1-1.8 cm. Calyx disk-shaped, but 
slightly oblique, obscurely and very obtusely 
5-lobed, lepidote dorsally, c. 3.5 mm 0. Corolla 
tubular-campanulate, rather fleshy, red, the tube 
orange towards the base, 4.5-5.2 cm long in all, 
glabrous in- and outside, tube straight, c. 3.5 cm, 
at base 6-7 mm 0, widened gradually to c. 1.5 cm 
at apex, lobes erect or somewhat spreading, sub- 
orbicular, 1.4—2 cm 0. Stamens 10, c. 3.5 cm; 
filaments linear, glabrous; anthers obovate- 
obcordate, much narrowed towards the base, 
each cell ending there in a narrow apiculus, 4—4.5 
by 2-2.2 mm. Disk glabrous. Ovary subovoid- 
conical, 6-7 by c. 3 mm, densely lepidote, rather 
suddenly narrowed to the style, the latter stoutish, 
c. 2 cm, lepidote in its lower third; stigma capitate- 
truncate. 

Distr. Malesia: W. New Guinea (Wichmann 
Mts), at 3100 m. Once found. Fl. Feb. 

204. Rhododendron pachycarpon Sleum. Rein- 
wardtia 5 (1960) 186; Blumea 12 (1963) 110. 

Shrub, 1.2-1.5 m. Branches greyish corticate, 
5-6 mm 0. Branchlets terete, c. 4 mm 0, ends 
laxly lepidote in flowering, glabrous and pruinose 
at fruiting time; internodes 3-6 cm. Leaves either 
scattered or in 3-merous pseudowhorls at the 
upper nodes, obovate, apex obtuse or rounded, 
sometimes slightly emarginate, base cuneate, 
coriaceous and rigid, glabrous above at maturity, 
subdensely lepidote beneath (scales small, nearly 
round, rather large, marginal zone variously 
angular or lobed, centre minute and ~ distinctly 
impressed), margin narrowly cartilaginous and 
subserrulate-crenulate, flat, 5-6.5 by (2.5-)3-4.5 
cm, midrib narrowly impressed above, thick, 
flattened and obtusely prominent beneath, nerves 



Jan. 1966] 



Ericaceae (Sleumer) 



615 



6-8 pairs, lower 2 ones from near the base, the 
others at an angle of c. 45 ^ all high-ascending 
and curved-anastomosing before the edge, slightly 
impressed above and raised beneath, reticulation 
subinconspicuous, grooved above, lepidote, 3-5 
by 2.5-3.5 mm. Umbels {3-)4~S-f\o'weTed. Pedicels 
zn densely lepidote, epilose, 1.5 cm by c. 1 mm. 
Corolla tubular below, funnel-shaped-expanded at 
the lobes, thin-fleshy, of a pure sulphur or lemon 
yellow, f. 5 cm long in all (6-7 cm when fresh), 
nodding, showy, quite glabrous in- and outside, 
tube c. 3 cm long, straight, base 5-7 mm o, c. 1 cm 
at apex, lobes erecto-patent, obovate-suborbicular, 
c. 1.5 cm 0. Stamens 10, nearly equalling the 
corolla tube; filaments linear, glabrous; anthers 
obovate-obcordate, much narrowed towards the 
base, c. 3 by 1.5 mm, each cell protracted into a 
basal appendix. Disk prominent, glabrous. Ovary 
thick-obovoid-cylindric in later stages, very 
densely lepidote, 6-7 by c. 3.5 mm, abrupt with 
the style, the latter c. 3 cm, lepidote at the base, 
otherwise glabrous; stigma obliquely capitate, 
shortly lobed. Capjw/^ obliquely obovoid-cylindric, 
short-attenuate at the base, rather abruptly so at 
the apex, lepidote, 2-2.5 by 0.8-0.9 cm, crowned 
by the rest of the style; pedicel 2-3 cm by 1.5 mm. 

Distr. Malesia: E. New Guinea (Mt Saruwaged 
area: Mt Enggan; Rawlinson Range in Morobe 
Distr.). 

Ecol. In OF>en subalpine forest on limestone, 
2135-3655 m. Fl. March-June. 

205. Rhododendron celebicum (Bl.) DC. Prod. 7 
(1839) 725; Miq. Fl. Ind. Bat. 2 (1859) 1058; 
Ann. Mus. Bot. Lugd.-Bat. 1 (1863) 44, t. 2 B; 
Bl. Fl. Jav. PI. ined. (1863-83) t. 6; Sarasin, 
Reisen in Celebes 1 (1905) 235; Sleum. Rein- 
wardtia 5 (1960) 187. — Vireya celebica Bl. Bijdr. 
(1826) 855; G. Don, Gen. Syst. 3 (1834) 848. — 
R. lubulosum Reinw. Reis. I. A. (1858) 576, nom. 
nud. — R. javanicum [non (Bl.) Benn.] Koord. 
Minah. (1898) 514; Koord. -Schum. Syst. Verz. 3 
(1914) 100. — /?. discolor Warb. in Sarasin, Reisen 
in Celebes 1 (1905) 28, in text, nom. subnud., nun 
Franch. 1895. 

Epiphytic shrub, 1-1.5 m; roots said to be 
swollen and tuber-like at the ends. Branchlets 
terete, 1.5-3 mm 0, upper internodes subdensely 
substellate-lepidote, older parts glabrous, smooth, 
often pruinosc; internodes 1.5-6 (rarely up to 
12) cm. Leaves opposite or in 3-mcrous pseudo- 
whorls at the upper 1-3 nodes, elliptic to lanceolate 
elliptic, apex short-acuminate, subacute or blunt, 
base broadly attenuate to the petiole, coriaceous, 
glabrous and slightly rugo.sc above at full age, 
subdensely lepidote beneath (scales small, flat, 
marginal zone thin, rather large, variously angled 
or substellatc-lobcd to -dented, centre small, not 
impressed), edge but slighllyand narrowly rcvolutc, 
(3-)4-7.5 by (1.5-)2-3(-3.5) cm, midrib rather 
narrow and slightly impressed above, robust and 
prominent beneath, nerves 5-7 pairs, subpatent. 
anastomosing, faintly sunk above, very slightly or 
not raised beneath, no proper reticulation visible; 
petiole ; flattened, lepidote, 4-IO( 12) by 1-1.5 



mm. Umbels (2-)3-5(-7)-flowered. Ferulae ovate 
to obovate, glabrous dorsally, margin with — 
early caducous stellate scales, 0.5-1.2 by 0.4-0.7 
cm. Bracteoles linear below, subspathulate-dilated 
and irregularly dented distally, glabrous, c. 1.5 cm 
by 1-2 mm. Pedicels slender, very densely stellate- 
lepidote and papillose, not properly hairy, 1.5-2 
cm. Calyx disk-shaped, oblique, shortly obtusely 
5-lobed, densely lepidote outside, 2.5-3 mm 0. 
Corolla tubular below, expanded at the lobes, thin, 
red to carmine, 3-3.5(-4) cm long in all, laxly to 
very laxly lepidote outside, glabrous inside, tube 
cylindric, straight, c. 2.5(-3) cm long, base angular 
by 5 pouches, 5-6 mm 0, widened to 8-10 mm at 
apex, lobes suborbicular, 6-10 by 6-8 mm. 
Stamens 10, unequal, the longest mostly reaching 
~ halfway the corolla lobes, sometimes equalling 
the corolla; filaments linear, glabrous; anthers 
broad-oblong, 2-2.5 by 1 mm, base rounded. 
Disk prominent, glabrous. Ovary subcylindric, 
very densely stellate-lepidote, 5-6 by 2 mm, 
continuous with the style, the latter quite glabrous 
or lepidote at the base only, i as long as the 
corolla; stigma thick-capitate. Submature capsule 
narrow subobovoid-cylindric, 2.5-3.8 by 0.35 cm, 
lepidote, pedicel up to 3 cm. 

Distr. Malesia: Celebes (Minahasa: Mt Klabat, 
Poso Lake area; Central part). 

Ecol. Mossy forest, 1800-2600 m. Fl. Jan., 
June, Nov. 

206. Rhododendron sayeri Sleum. Reinwardtia 5 
(1960) 188. 

Shrub, c. 1.8 m. Branchlets divaricate, terete, 
1.5-3 mm 0, upper internodes very densely 
rufous-stellate-lepidote, early glabrescent and 
smooth lower down; internodes 3-7 cm. Leaves in 
3(-5)-merous pseudowhorls at the upper 1-3 
nodes, broad-elliptic, sometimes obovate-elliptic, 
more rarely obovate, apex (very) broadly attenuate, 
obtuse or sometimes rounded, broadly attenuate 
or nearly rounded towards the base, coriaceous, 
mature ones quite glabrous above, subpcrsistently 
densely rufous-lepidote beneath (scales small, ± 
deeply stellate-dented or -lacerate in the marginal 
zone, minute centre dark and very slightly im- 
pressed), margin a little revolute to base or quite 
flat, (3.5-)4.5-6.5 by (2-)2.5-4 cm. midrib slightly 
impressed above, markedly prominent especially 
in the lower part, nerves 6-8 pairs, irregular, 
spreading, anastomosing before the edge, some- 
what impressed or raised as are the midrib and 
veins above, prominent beneath, veins forming a 
lax, well visible network on both faces, vcinlets 
less conspicuous beneath; petiole flattened and 
grooved dorsally, lepidote, 4-7 by 1-1.5 mm. 
Umbels (l-2-)3-4-flowcred. Ferulae ovate to 
obovate. obtuse, glabrous dorsally, very shortly 
subglandular-fimbriate, up to I by 0.8 cm. 
Bracteoles linear below, subspathulate-dilalcd 
distally. up to 1.2 cm by 0.5-1.5 mm. Pedicels 
very densely rufous-stcllatc-lepidotc, 1.5 2.3 cm 
by c. 0.6 mm. Calyx obliquely disk-shaped, irregu- 
larly 5-dentcd. lepidote at the base dorsally, teeth 
deltoid, subacute, 0.5 2 (rarely up to 3) mm, ± 



616 



Flora Malesiana 



[ser. I, vol. 6* 



spreading, subglabrous. Corolla tubular below, 
lobes expanded, suboblique, 4-4.5 cm long in all, 
± laxly stellate-lepidote outside, laxly and longish 
patent-hairy at the tube inside, tube • curved, c. 
3 cm, the 5-saccate base 4-5 mm 0, ampliate to 
7-8 mm at apex, lobes suborbicular, 0.9-1.3 
cm 0. Stamens 10, ± as long as the corolla tube; 
filaments linear, laxly patently long-hairy in the 
lower 2/3; anthers subobovate-oblong, 2.5 mm by 
c. 1.2 mm, base obtuse. Disk prominent, sparsely 
hairy in the upper margin. Ovary subcylindric, 
rather abruptly narrowed to the style, very densely 
stellate-lepidote, epilose, 5-6 by 2-2.5 mm; style 
ifc as long as the stamens, laxly stellate-lepidote in 
its lower third, glabrous for the rest; stigma thick- 
capitate. 

Distr. Malesia: New Guinea (Central Distr.: 
Mt Obree), growing in profusion on brink of 
precipitous cliffs at Howers Lake, c. 2135 m. 
Once found. Fl. Aug. 

207. Rhododendron pachystigma Sleum. Blumea 12 
(1963) 110. 

Terrestrial shrub, 1.5 m. Branchlets subterete, 
tips densely lepidote, older parts striate and prui- 
nose; internodes 4— 10(-13) cm by 3-5 mm. Leaves 
at the apex of or at the upper 2 internodes or al- 
most at the nodes, obovate or elliptic-obovate, 
apex rounded, sometimes slightly retuse, base ± 
broadly cuneate to the petiole, when young 
densely lepidote on both faces, mature ones 
coriaceous, firm, rather early glabrescent (scales 
small, rufescent, sessile and flattish, detersile, 
marginal zone irregularly substellate-dented, centre 
minute, hardly or not impressed), entire, (5-) 
6-8.5(-9) by (2.5-)3^.5(-5) cm, subrevolute 
towards the base, midrib thick and grooved in the 
lower part above, as strong as the petiole at 
the base beneath, nerves 8-9 pairs, straight, at an 
angle of c. 45° and subparallel at the base, obtusely 
curved-anastomosing before the edge, faintly im- 
pressed above, a little raised beneath, no reticula- 
tion; petiole densely lepidote initially, l-2(-2.5) 
cm by c. 2 mm. Umbels 2-3-flowered. Immature 
perulae laxly lepidote dorsally, no hairs. Bracte- 
oles filiform, glabrous, seen up to 1.5 cm. Pedicels 
with dense, small rusty scales, furthermore with 
fine short hairs in the lower % , and upwards more 
densely with longer hairs, as on the calyx, (8-) 
11-14 by 1.5-2 mm. Calyx obliquely patellar, 
6-7 mm 0, lobes irregular, spreading, deltoid to 
elongate-deltoid, fimbriate and caducously lepi- 
dote along the margin, 2-3 (-4) mm. Corolla 
funnel-shaped, deeply (6-)7-lobed, light orange 
at the tube outside, white at the lobes, fleshy, 
(7-)7.5-8 cm long in all, sparsely lepidote or 
glabrous outside, densely white-hairy in the lower 
part of the tube inside, tube straight, (2.5-)3 cm 
long, base 6-7-pouched, 8-9 mm 0, gradually 
widened to 2-2.5 cm, lobes obovate-spathulate, 
spreading, 4. 5-5 by (2.5-)3-3.5 cm. Stamens 12-14, 
unequal, well exserted; filaments linear, 1 mm wide 
below, denjely white-hairy in the basal part, 
filiform an<i glabrous in the upper part; anthers 
versatile, elongate-oblong, base obtuse, c. 8 by 



1-1.2 mm. Disk prominent, densely white-hairy 
on the margin. Ovary thick-cylindric, 8-9 by c. 4 
mm, red-brown stellate-scaly, no hairs, tapering 
to the style, the latter glabrous, c. 4 cm by 1-1.5 
mm; stigma subpeltate, 6-8(-9) mm 0, with 6-7 
thick, subglobular lobes. 

Distr. Malesia: W. New Guinea (Balim R. 
Valley near Wiligimaan), once collected. 

Ecol. Young secondary forest, on stony clay, 
c. 1800 m. Fl. June. 

Vern. Poroom, Dani. 

208. Rhododendron angulatum J. J. S. Bull. Jard. 
Bot. Btzg II, 8 (1912) 50; Nova Guinea 12 (1914) 
140, t. 36 A; Sleum. Reinwardtia 5 (1960) 188. 

Terrestrial shrub, c. 75 cm. Branchlets rather 
robust, terete, laxly lepidote. Leaves in i 5-merous 
pseudowhorls at the upper nodes, ovate-lanceolate, 
apex gradually long-acuminate, very acute, base 
obtuse-rounded or very slightly cordate, coriace- 
ous, glabrous above with age, subdensely lepidote 
beneath (scales stellate-dented in the marginal 
zone, centre ± impressed, leaving minute pits 
when the scale is gone), faintly or not recurved at 
margin, 11-18 by 3.5-5.8 cm, midrib narrowly 
grooved above, stout and much obtusely promi- 
nent beneath in the lower part, nerves 12-15 
pairs, ascending, anastomosing, very slightly 
raised or impressed above, obscure beneath, no 
reticulation visible; petiole robust, compressed 
and grooved dorsally, transversely rugulose, 
minutely lepidote, 1-1.2 cm by 3 mm. Umbels ± 
6-flowered. Pedicels laxly minutely lepidote, 
more densely so below the calyx, epilose 4—5 cm 
by 2.5 mm. Calyx annular, oblique, obtusely and 
rather inconspicuously 5-angular, ciliolate, c. 
4.5 mm 0. Corolla funnel-shaped, 5-lobed in the 
upper 2/3, rose red and rather fleshy, 5.5-6 cm 
long in all, sparsely minutely lepidote outside, laxly 
pubescent at the tube inside, tube distinctly 5- 
angular or pouched at the base outside, deeply 
5-grooved lengthwise in the lower part inside, 
the ribs extending upwards to the median of the 
corolla lobes, c. 2 cm long, c. 8 mm at base, 
hardly widened distally, lobes subovate-elliptic, 
obtuse, spreading, 3.5-4 by c. 2 cm. Stamens 10, 
nearly as long as the corolla; filaments linear and 
shortly patently pubescent below, narrower and 
glabrescent upwards; anthers linear-oblong, 5.5-6 
by 1 mm, obtuse at the base. Disk much promi- 
nent, practically glabrous. Ovary conical-cylindric, 
densely clothed with thickish nearly roundish 
scales, no hairs, c. 10 by 3 mm, continuous with 
the style, the latter lepidote and sparsely minutely 
pubescent (or nearly papillose) at the base, 
glabrous for the rest, c. 4.8 cm; stigma thick- 
capitate. 

Distr. Malesia: W. New Guinea (Oranje Mts: 
Mt Goliath), common in shadow^y swampy 
humous places at 1500 m. Once collected. Fl. 
April. 

209. Rhododendron loerzingii J. J. S. in K. & V. 

Bijdr. 13 (1914) 105, 107; Amsh. in Back. Bekn. 
Fl. Java (em. ed.) 7 (1948) fam. 162, p. 3; Sleum. 



Jan. 1966] 



Ericaceae (Sleumer) 



617 



Reinwardtia 5 (1960) 189; Steen. Endeavour 21 
(1962) 189, t. 4. 

Erect, much-branched, terrestrial shrub, c. 2.5 
m. Branchlets divaricate, terete, tips subdensely 
clothed with flat substellate scales, lower down 
glabrescent and smooth; internodes 2-8(-10) cm. 
Leaves very close together or in 3-5-merous 
pseudowhorls at the upper nodes, mostly oblong- 
elliptic, more rarely elliptic, apex short-acuminate, 
subacute, rarely obtuse, base broadly and ± 
acutely attenuate to the petiole, coriaceous, 
initially lepidote on both faces, glabrescent above, 
more persistently lepidote beneath (scales dense 
though distant from each other, small, rufescent, 
the rather narrow marginal zone variously angled, 
lobed or substeilate-dented, centre dark, distinctly 
impressed, pits blackish with age), edge narrow- 
cartilaginous and somewhat revolute, 6-8.5(-12) 
by 2.3-3.5(-4.4) cm, midrib narrow and slightly 
impressed above, rather stout and obtusely 
prominent beneath, nerves 8-10 pairs, spreading, 
anastomosing before the edge, mostly faintly 
impressed above, slightly raised beneath, veins 
subinconspicuous; petiole compressed and grooved 
dorsally, 4-8(-10) by 1.5-2 mm. Umbels (3-)5-9- 
flowered. Flower buds ovoid, outer perulae ovate, 
inner ones obovate, obtuse or mostly apiculate, 
glabrous except some scales on the upper part of 
the dorsal mediane or the very tips, margin laxly 
set with caducous scales, 0.4-1.7 by 0.3-0.8(-l.l) 
cm. Bracteoles linear below, spathulate at the 
apex, glabrous, 0.7-1 cm by 0.5-1 mm. Pedicels 
rather slender, very densely stellate-lepidote, 
epilose, 2-2.5 cm. Calyx small, obliquely annular 
or nearly cup-shaped, lepidote dorsally, c. 3 mm 
0, teeth deltoid, unequal, subpatent, acute, 
glabrous apically, 0.5-2 (occasionally up to 3) mm. 
Corolla tubular in outline, expanded at the lobes, 
scarlet, membranous, sparsely stellate-lepidote 
outside, pubescent in lower half of the tube in- 
side, (2.2-)2.5-3 cm long in all, tube straight, 
1.5-2.2 cm long, at base 4-5 mm 0, widened to 7-8 
mm at apex, lobes obovate, subequal, 0.9-1.3 by 
0.6-1 cm. Stamens 10, :•. as long as the corolla 
tube; filaments linear, subdensely clad with longish 
patent hairs in the lowest 3-5 mm, glabrous for 
the rest; anthers obovate, apex truncate, base 
subacute, 1-1.5 mm. Disk prominent, glabrous. 
Ovary subcylindrical-conical, much abrupt with 
the style, densely stellate-lepidote, or with some 
very short hairs in especially the lower part between 
the scales, i.e. practically epilose, 5-7 by c. 3 mm; 
style rather thick, glabrous, nearly as long as the 
stamens; stigma capitate, with thick lobes. Capsule 
oblique cylindric, 5-angular, densely lepidote, 
2-2.3 by 0.5 cm. Seeds c. 6 mm including the tails; 
pedicel to 4.8 cm. 

Distr. Malesia: Central Java (Kcdu: Mts 
Tlcrep and Sumbing). 

Ecol. In scrub or grassland, on rather dry but 
fertile soil, very rich in humus, apparently rare, 
1«0(X-2(XXJ m. //. March-Nov../r. Aug.-Nov. 

Vcrn. Pur wo yiUii, J. 

210. Khododcndrun imprcsHopunctatum J. J. .S. in 



Fedde, Rep. 30 (1932) 164; Slelfm. Reinwardtia 5 
(1960) 189. 

Terrestrial shrub, c. 1.2 m. Branchlets subterete, 
upper 1-2 internodes ± densely clad with stellate 
scales, lower ones quite glabrous, striate, dark, 
later with thin grey cork; internodes 2-5(-10) cm. 
Leaves in 4— 5-merous pseudowhorls at the upper 
1-3 nodes, oblong-elliptic, apex short-acuminate, 
subacute or rarely bluntish, base broad-attenuate 
to nearly rounded, coriaceous, subdensely set 
with small, flat, non-impressed and early decolour- 
ed scales for a long time, especially on the midrib, 
finally glabrescent above, more densely and ±, 
persistently lepidote beneath (scales dense, but 
for their greater number distant from each other, 
rust-coloured, rather small, marginal zone ± 
irregularly lobed or dented, dark centre distinctly 
impressed), margin slightly recurved, 6.2-8.3 by 
2.4—3.2 cm, midrib grooved above, strongly and 
obtusely prominent beneath, nerves 8-10 pairs, 
spreading, but obscurely anastomosing, very 
slightly impressed above, not much raised beneath, 
reticulation inconspicuous; petiole very densely 
lepidote initially as is the lower part of the midrib 
beneath, i.e. the scales ± touching each other 
there, semiterete, much grooved above, 8-10 by 
1.5-2 mm. Umbels 5-7-flowered. Pedicels sparsely 
lepidote, 1-1.9 cm by 0.7 mm. Calyx obliquely 
disk-shaped, unequally 5-angular or very shortly 
dented, c. 4 mm 0. Corolla tubular-funnel-shaped, 
red, 3-3.5 cm long in all, quite glabrous in- and 
outside, tube straight, 2-2.3 cm, at the somewhat 
saccate base c. 5 mm 0, slightly dilated upwards, 
lobes expanded, obovate to suborbicular, 1-1.3 
by (0.7-)0.9-1.2 cm. Stamens 10, somewhat exsert- 
ed from the corolla tube; filaments linear, densely 
patently hairy in the lowest 5-7 mm; anthers 
oblong, base obtuse, c. 3.5 by 1 mm. Disk promi- 
nent, glabrous. Ovary conical-subcylindric, very 
densely stellate-lepidote, epilose, 5-6 by 2-2.5 mm, 
rather abrupt with the style, the latter ± as long as 
the stamens, glabrous; stigma capitate. 

Distr. Malesia: Moluccas (Buru), once found 
in peatmoor at Kunturun, 1075 m. Fl. Feb. 

Note. Size and colour of the scales suggest, 
that R. impressopunctatum is a hybrid of R. 
malayanum with another species of the Javanicum 
group, presumably R. seranicum; these three 
species grow together at Kunturun. 

211. Rhododendron xanthopctalum Merr. Publ. 
Gov. Lab. Philip. /;. 29 (1905) 41; Philip. J. Sc. 
1 (1906) Suppl. Ill; ibid. 3 (1908) Bot. 380; 
M1LLAI.S, Rhodod. (1917) 263; Mirr. En. Philip. 
3 (1923) 246; Millaks, Rhodod. scr. 2 (1924) 261; 
CoPEL./. Philip. J. Sc. 40 (1929) 169, t. 16. f. 1-2; 
Sleum. Reinwardtia 5 (1960) 189. 

Epiphytic shrub, 80 cm or less, with a stout 
simple or but slightly branched stem. 1 cm below, 
covered with light grey bark. Branchlets reddish 
brown, glabrous. Leaves scattered along the upper 
internodes, oblong-elliptic, apex slightly acuminate 
or merely acute, base acute, coriaceous, glabrous 
above, lepidote. margin revolute, 9-12 by 3.5-5 
cm, midrib very stout and prominent; petiole very 



618 



Flora Malesiana 



[ser. I, vol. 6'' 



robust, 1.5 cm. Umbels 3-4-flowered. Pedicels 
glabrous, 1.5 cm. Calyx reduced to an obscure 
disk. Corolla campanulate, yellow, 3-3.4 cm long 
in ail and about as wide at the limb, tube 1.5 cm 
long, c. 5 mm at base, lobes rounded, 1.5 cm. 
Stamens 10; filaments pubescent below, 1.8 cm; 
anthers linear, without any basal appendix, 4 mm. 
Disk crowned with abundant hairs. Ovary lepidote, 
8 mm; style glabrous, abrupt, 1.2 cm. 

D i s t r . Malesia: Philippines (Luzon; ? Mindoro). 

Ecol. Epiphyte in moss forest, 1200 m, appar- 
ently rare. 

Note. An imperfectly known species of which 
I have seen not more than a single leaf. After 
Copeland/., the occurrence in Mindoro remains 
uncertain. 

212. Rhododendron hirtolepidotum J. J. S. Nova 
Guinea 12 (1914) 135, t. 32; Kaneh. &. Hatus. 
Bot. Mag. Tokyo 56 (1942) 479; Sleum. Rein- 
wardtia 5 (1960) 189; Blumea 12 (1963) 110. 
Terrestrial or occasionally epiphytic shrub, 
(0.5-)l-2(-2.5) m, not much branched; bark 
brown. Branchlets terete, (2-)3-5 mm 0, upper 
internodes or very tips only subdensely clothed 
with early caducous, substellate, rufescent scales, 
lower parts glabrous and smooth, early covered 
with cork; internodes (2-)4-18 cm. Leaves in 
4-6(-8)-merous pseudowhorls at the upper 1-2 
(-3) nodes, oblong-elliptic, sometimes obovate- 
elliptic, apex broad-attenuate or shortly sub- 
acuminate, obtuse to nearly rounded, base broadly 
narrowed, subtruncate to rounded, rarely sub- 
cordate, coriaceous, glabrous above at full age, 
rather densely and less fugaciously lepidote be- 
neath (scales small, variously substellate-lobed or 
-dented in the marginal zone, dark centre but 
slightly impressed), finally completely glabrescent, 
the remaining pits blackish and shallow, edge 
slightly revolute, (4-)5-7.5(-10.5) by (2-)2. 5-4.2 
cm, midrib slightly impressed above, stout and 
much prominent in the lower part beneath, nerves 
6-8(-10) pairs, spreading, rather straight below, 
curved and ± indistinctly anastomosing before the 
edge, slightly raised on both faces or a little im- 
pressed above, veins lax, somewhat prominent 
above, mostly subinconspicuous beneath, no 
proper reticulation; petiole semiterete, grooved 
above, lepidote and rugulose, (4-)6-12(-16) by 
1.5-2(-2.5) mm. Umbels 5-9-flowered. Perulae 
ovate to obovate, obtuse, firm-membranous, with 
some sparse scales at the top dorsally or quite 
glabrous, up to 1.5 by 0.8 cm. Bracteoles linear 
below, subspathulate-dilated apically, glabrous, 
up to 2.2 cm by 1.5 mm. Pedicels sparsely stellate- 
lepidote below, more densely so distally, 3^ cm 
by 0.6-0.8 mm. Calyx very oblique, disk-shaped, 
5-angular, c. 4 mm 0, dorsally densely lepidote. 
Corolla tubular below, suboblique, rose-red to 
carmine, membranous, rather laxly to sub- 
densely lepidote outside, short-hairy at the lower 
half of the tube inside, 4-^.5(-5) cm long in all, 
tube (2-)2.5-3 cm long, at base 5-saccate, 6-7 mm 
0, slightly dilated to apex, lobes ± obliquely 
obovate-spathulate, (0.9-)1.3-1.7 by 1-1.2 cm. 



Stamens 10, slightly shorter than the corolla; 
filaments linear, densely pubescent below, less so 
upwards; anthers oblong, obtuse at the base, 
3.5-4 by 1 mm. Disk short-pubescent at the upper 
margin, glabrous below. Ovary subcylindric, 
densely stellate-lepidote, with some spreading 
hairs on the very top between the scales, c. 8 by 
2.5 mm, rather gradually tapering to the style, 
the latter stoutish, laxly lepidote and subdensely 
patent-pubescent in the lower 1-1.5 cm, glabrous 
for the rest, ±_ as long as the corolla tube; stigma 
capitate, deeply 5-lobed. Capsule subcylindric, 
subdensely set with red-brown scales, 3.5-4 by 
0.4—0.5 cm; pedicel to 6 cm by 1 .3 mm. 

Distr. Malesia: New Guinea (Vogelkop 
Peninsula: Arfak Mts, Nettoti Range, Mt Watje- 
toni). 

Ecol. In light mossy forest in poor humous or 
sandy soil, or in swampy places with Sphagnum 
(1200-) 1750-2000 m. Fl. April, Nov.-Dec. 

Vern. Krobua, Kebar. 

213. Rhododendron pseudomurudense Sleum. Rein- 
wardtia 5 (1960) 189. — R. murudense (non Merr.) 
J. J. S. Bull. Jard. Bot. Btzg III, 13 (1935) 449. 
Epiphytic shrub. Branchlets very robust, upper 
internode obtusangular, c. 5 mm 0, rather densely 
clothed with flat, appressed, rufescent scales, 
lower ones terete, 6-8 mm 0, glabrescent and 
smooth; internodes 5-16 cm, occasionally longer. 
Leaves spirally arranged, all or most of them (3-5) 
in the upper half of the upper 1-2 internodes, 
oblong, sometimes subovate- or elliptic-oblong, 
apex very broadly attenuate, obtuse, base mostly 
obtuse to rounded, or sometimes cordulate, 
coriaceous, thick and stiff, glabrous above at 
full maturity, subdensely lepidote for a long time 
beneath (scales minute, variously and irregularly 
substellate-dented at the marginal zone, which 
disappears early, centre dark, faintly or not 
impressed, leaving dark points when the proper 
scales are gone), margin flat, 12-18 by 4—7 (-8) 
cm, midrib as robust as the petiole, obtusely 
prominent on both faces in the lower part, nerves 
(15-) 17-20 pairs, very distinct and subparallel 
from the midrib at c. 90', i straight below, 
curved and anastomosing before the edge, promi- 
nent on both sides as is the rather dense reticula- 
tion; petiole red brown in dry specimens, subterete, 
not grooved above, glabrous, (1. 3-) 1.5-2.1 cm 
by 3-4(-5) mm. Umbels 12-25-flowered. Perulae 
ovate to ovate-oblong, glabrous (immature). 
Bracteoles linear or nearly filiform, glabrous, 
c. 2 cm by c. 1 mm. Calyx oblique, disk-shaped, 
obscurely lobed, glabrous, c. 4.5 mm 0. Corolla 
broad-funnel shaped, dark red to salmon, 2.5-3 
cm long in all, glabrous in- and outside, tube 
cylindric below, widened upwards, 1.2-1.5 cm 
long, the 5-saccate base c. 4 mm 0, widened to c. 

1 cm at apex, lobes oblong-obovate to subspathu- 
late, 1.2-1.5 by 0.8-1 cm. Stamens 10, unequal, c. 

2 cm long; filaments linear, quite glabrous; 
anthers obovate-elliptic, base obtuse, 3-3.5 by 1.5 
mm. Disk thick-prominent, glabrous. Ovary 
cylindric, c. 4 by 2.5 mm, very abrupt with the 



Jan. 1966] 



Ericaceae (Sleumer) 



619 



style, quite glabrous, somewhat rugulose, blackish 
when dry; style (4— )5-6 mm, glabrous; stigma 
capitate, distinctly lobulate, c. 2 mm o. Capsule 
(immature) obliquely subcylindric, 1.5 by 0.5 
cm; pedicel up to 5 cm. 

Distr. Malesia: Central E. Borneo (Mt Kemul), 
1600-1850 m (summit), common in primary 
forest. Fl. Oct. 

214. Rhododendron seranicum J. J. S. in Fedde. 
Rep. 30 (1932) 165; Bot. Jahrb. 68 (1937) 202, 
in obs. {'seranense'); Sleum. Reinwardtia 5 (1960) 
189. 

Shrub, up to 5 m. Branches 5-6 mm o, with ashy 
cork. Branchlets 2.5-4 mm 0, terete, upper inter- 
node laxly lepidote, often pruinose, smooth; 
internodes 1.5-9(-lI.5) cm. Leaves scattered 
along the upper 1-2 internodes, the upper 2-3 
often close together near the node, elliptic, 
subovate- or oblong-elliptic, more rarely oblong, 
apex short-acuminate, subacute, base broadly 
attenuate to the petiole, or nearly rounded, 
coriaceous, quite glabrous above at maturity, 
rather laxly lepidote beneath for some time (scales 
small, flat, marginal zone irregularly lobed or 
dented, centre faintly or not impressed), finally 
— glabrescent beneath too, edge shortly and 
narrowly recurved, 5-8.5 by (2-)2.5^ cm, midrib 
a continuation of the thick and somewhat flattened 
petiole, much and bluntly prominent in the lower 
part on both faces, nerves 8-10 pairs, ir irregular, 
spreading below, curved-ascending and obscurely 
anastomosing towards the edge, slightly prominent 
on both faces, veins laxly reticulate, a little raised 
or often - inconspicuous; petiole 0.5-1 (rarely 
up to 1.5) cm by 2-3 mm. Umbels 7-1 1 -flowered. 
Inner perulae oblong to narrow-oblong, obtuse, 
membranous, glabrous, 3-3.5 by 0.5-1 cm. 
Bracteoles linear, glabrous, c. 2.5 cm by I mm. 
Pedicels thickened at apex, sparsely lepidote or 
practically glabrous, (1.5-)2-3 cm by c. 1 mm. 
Calyx very oblique, shortly cup- or disk-shaped, 
thickened, obscurely or not lobcd, laxly lepidote 
outside, c. 5 mm 0. Corolla broad-funnel-shaped, 
light violet, rose or brownish yellow (limb orange, 
salmon rose towards the border), or fiery red, 
glabrous in- and outside, f4-)4.5-5.5 cm long in 
all, tube 1.7-2.1 cm long, the saccate base 5-6 
mm 0, widened to 1-1.5 cm at apex, lobes unequal, 
oblong-obovate, c. 3 by 2 cm. Stamens 10, un- 
equal, 2.8-3.2 cm; filaments flat-linear, glabrous; 
anthers oblong, somewhat curved, base blunt or 
minutely apiculatc, 3.5-4 by 1 mm. Disk prominent, 
glabrous. Ovary oblong-conical, glabrous from 
the beginning, minutely rugulose, blackish when 
dry, c. 6 by 3 mm, rather abrupt with the style, 
which is glabrous and 1.5-2 cm long; stigma thick- 
capitate, manifestly 5-lobcd. Capsule oblique, 
thick-fusiform, 3.5-4 by 0.7-0.8 cm, glabrous; 
pedicel 3-4 cm by 1.5 mm. 

Distr. Malesia: Moluccas (Ceram, Buru, and 
Ambon). 

licol. Light thickets on open spots and in 
primary forest, 9(X)-I700 m, on limestone, locally 
abundant. H. Aug.-Feb. 



Note. A specimen slightly differing by salmon 
red, 3.5 cm long corollas and manifestly biappen- 
diculate anthers has been found in Central Celebes 
(Palopo-Rantepao road), in dry rocky hillside at 
900 m. 

215. Rhododendron loboense Copel./. Philip. J. Sc. 
40 (1929) 137, 172. t. 15, f. 3, t. 16, f. 5-6; Sleum. 
Reinwardtia 5 (1960) 190. 

Epiphytic shrublet, 30-40 cm. Branchlets terete, 
c. 5 mm 0, glabrous. Leaves scattered along the 
upper internode (lower internodes not known), 
elliptic to oblong-elliptic, apex shortly sub- 
acuminate, obtuse, base broadly subacutely 
attenuate, very coriaceous and stiff, glabrous 
above at maturity, rather laxly and ± persistently 
lepidote beneath (scales relatively small, irregu- 
larly lobed or dented at the marginal zone, centre 
faintly or not impressed), finally glabrous on 
both faces, margin but slightly recurved, 10-12 
by 4-6 cm, midrib robust, rather flattish in dry 
specimens, obtusely prominent on both sides, 
as broad as the petiole at the base, gradually 
narrowed upwards, nearly inconspicuous at the 
apex, nerves 12-15, rather dense pairs, at an acute 
angle, indistinctly anastomosing, ± markedly 
prominent at both faces, veins laxly reticulate, 
mostly visible beneath only; petiole somewhat 
flattened, (1-) 1.5-2 cm by 2.5-3 mm. Umbels ± 
5-flowered. Pedicels practically glabrous, 1-1.5 cm 
by c. 1.5 mm. Calyx disk-like, margin thickened, 
wavy and faintly lobed, glabrous. Corolla funnel- 
shaped, 3.5-4 cm long in all, yellow, glabrous on 
both sides, lobes unequal, oval, c. 2 cm. Stamens 
10; filaments linear, glabrous, c. 2.5 cm; anthers 
oblong, base obtuse, 4 by 1 mm. Disk glabrous. 
Ovary subobovoid-cylindric, quite glabrous from 
the beginning, c. 7 by 2.5 mm; style abrupt with the 
ovary, glabrous, 1.3 cm; stigma capitate, c. 3 mm 
0. Capsule subcylindric, 4 by c. 0.5 cm; pedicel 
1.5-2 cm. 

Distr. Malesia: Philippines (S. Luzon: Prov. 
Batangas, Lobo Mts). One collection. Fl. April. 

Vern. Maldgos, Lobo. 

216. Rhododendron williamsii Merr. ex Copel. /. 
Philip. J. Sc. 40 (1929) 137, 163, t. 9, t. 11, f. 2; 
Sleum. Reinwardtia 5 (1960) 190. — R. schaden- 
bergii {non Warb.) Mfrr. Publ. Gov. Lab. Philip. 
n. 29 (1905) 40, p.p.\ Philip. J. Sc. 3 (1908) Bot. 
380, p.p.; En. Philip. 3 (1923) 245, p.p. 

Small tree, 4-6 m; branches light grey or brown- 
ish, terete. Branchlets subtcrete, upper internodes 
laxly lepidote at the tips, or early glabrescent and 
not rarely quite glabrous and somewhat pruinose; 
internodes 3-10 cm. Leaves in 3(-4)-merous 
pseudowhorls at or close to the upper 1 or 2 nodes, 
oblong-ovate or oblong-elliptic, rather thinly 
coriaceous, apex shortly but sharply acuminate, 
ih acute, base acutely attenuate into the petiole 
or blunlish. practically glabrous above with age, 
rather laxly lepidote beneath initially (scales 
small, flat, variously and irregularly lobcd in the 
very thin, early dis.solvcd marginal zone, centre 
slightly darker, but faintly impressed), finally 



620 



Flora Malesiana 



[ser. I, vol. 6* 



glabrescent beneath too, leaving shallow dark pits, 
little or not revolute, 8-11 by 3-5(,-6) cm, midrib 
broad (1.5-2 mm) at the base, much narrowed 
upwards, impressed in the upper % above, 
obtusely prominent below, nerves 6-1 0(-l 2) pairs, 
somewhat ascending, slightly impressed or raised 
above, more conspicuously prominent beneath, 
open or rather obscurely anastomosing before the 
edge, reticulation lax, slightly prominent on both 
sides, not rarely subinconspicuous; petiole 1-1.5 
(-2) cm by 1.5-2.5(-3) mm, somewhat flattened. 
Umbels 5-8-flowered. Outer perulae broad-ovate, 
mucronulate, 1.5 by 1.2 cm, inner ones broad- 
oblong or elliptic, obtuse up to 2 by 1 cm with 
some dorsal scales or glabrous, margin very shal- 
lowly dented by early caducous scales. Bracteoles 
linear, glabrous, up to 2 cm by 0.5 mm. Pedicels ± 
densely set with rufescent flattish scales, especially 
towards the apex, 2-2.5 cm by 1 mm. Calyx very 
oblique, disk-shaped, shortly irregularly 5-dented 
(up to 1 mm), 4-5 mm 0. Corolla funnel-shaped, 
white, (2.5-)3-4 cm long in all, sparsely lepidote 
or mostly glabrous outside, glabrous inside, tube 
1.8-2.3 cm, base 5-pouched, c. 7 mm 0, widened to 
c. 1 cm at apex, lobes obovate-suborbicular, sub- 
expanded, 1-1.5 cm 0. Stamens 10, as long as or 
slightly exceeding the tube; filaments linear, 
glabrous; anthers oblong, c. 3 by 1 mm, each cell 
with a short basal apiculus. Disk glabrous, 
prominent. Ovary subovoid-cylindric, initially 
with rather dense, flat, rufescent, stellate scales, 
glabrescent, subabrupt with the style, (5-) 6-8 
by 2.5-3 mm; style c. 2 cm, with some scales at the 
base, otherwise glabrous; stigma thick-capitate, 
3 mm 0. Capsule oblong-fusiform, 2.5-3 by 0.6-0.7 
cm, bearing the long persistent style, which may 
attain 2.5 cm; pedicel to 4 cm by 1.5 mm. Seeds 
5-6 by 0.2 mm including the tails. 

Distr. Malesia: Philippines (Luzon: Mountain 
and. Zambales Prov.). 

Ecol. Along streams, in ravines, 1500-2200 m. 

217. Rhododendron robinsonii Ridl. J. Fed. Mai. 
St. Mus. 4 (1909) 44; ibid. 5 (1914) 39; Fl. Mai. 
Pen. 2 (1923) 216; Renders. Mai. Nat. J. 6 (1950) 
261; Sleum. Reinwardtia 5 (1960) 190. — R. 
javanicum [non (Bl.) Benn.] Clarke in Hook. /. 
Fl. Br. Ind. 3 (1882) 463; Ridl. J. Str. Br. R. As. 
Soc. n. 23 (1891) 145; K. & G. J. As. Soc. Beng. 
74, ii (1905) 74; Ridl. J. Fed. Mai. St. Mus. 5 
(1914) 38; Fl. Mai. Pen. 2 (1923) 216; Burk. & 
Renders. Gard. Bull. S. S. 3 (1925) 390; Renders. 
Mai. Nat. J. 6 (1950) 262. — R. teysmannii {non 
MiQ.) Renders. J. Mai. Br. R. As. Soc. 5 (1927) 
256. 

Small, epiphytic or sometimes terrestrial shrub, 
1-2.5 m. Branchlets subterete, very tips laxly 
lepidote, lower parts glabrous, upper 1-2 inter- 
nodes dark, older ones with a thin, ashy cork; 
internodes 4— 8 (-10) cm. Leaves in 3-4(rarely up 
to 8)-merous pseudowhorls at the upper 2-3 nodes, 
and often 1-2 additional ones below the nodes 
(several scars in the lower part of the internodes 
from early caducous, much reduced bract-like 
leaflets), elliptic or mostly elliptic-oblong, some- 



times lanceolate, apex short-acuminate, mostly ± 
acute, base acutely narrowed to the petiole, ± 
coriaceous, glabrous and with numerous minute 
impressed darker points at maturity above, laxly 
lepidote beneath (scales variously irregularly lobed 
or dented in the marginal zone, dark centre but 
shallowly impressed), finally ± completely 
glabrescent beneath too, margin very slightly 
or not revolute, (5-)6-12(-16, rarely up to 17) by 
(2-)2.5-4(-6) cm, midrib as stout as the petiole, 
obtusely prominent in its lower part at both 
faces, ± impressed distally above, nerves 8-10 
(-12) pairs, curved-spreading and obscurely inter- 
arching at some distance from the edge, slightly 
prominent on both sides, reticulation obscure, or 
faintly visible beneath only; petiole rugulose, some- 
what flattened, (0.7-)l-1.5(-2) cm by 1.5-2(-3) 
mm. Umbels (4-)5-8(-12)-flowered. Flower buds 
ovoid, acuminate, 3-3.5 by 2 cm, outer perulae 
ovate-acuminate, subacute, initially laxly lepidote 
dorsally, but very early glabrescent, inner ones 
ovate-oblong, more obtuse, glabrous, up to 2.5 by 
1.3 cm. Bracteoles linear below, subspathulate at 
the apex, glabrous, up to 2 cm by 0.5-2 mm. 
Pedicels laxly lepidote, early glabrescent, 2-3.5 
(-4.5) cm by 0.5-1 mm at anthesis. Calyx oblique, 
disk-shaped or merely rim-like, obscurely lobed 
or wavy, glabrescent, 3-4 mm 0. Corolla funnel- 
shaped, golden to deep creamy yellow, flushed 
apricot-pink, sometimes yellowish salmon, bright 
red or diluted crimson, thin, 3-3.5 cm long and 
wide, glabrous in- and outside, tube c. 1.5 cm 
long, base distinctly 5-pouched, 5-6 mm 0, 
widened to 1-1.5 cm at apex, lobes ± expanded, 
broad-oblong-obovate, 1.3-1.8 by 1-1.3(-1.5) cm. 
Stamens 10, unequal, 2-2.5 cm; filaments linear, 
quite glabrous; anthers oblong, a little curved, 
base obtuse, 3-3.5 by 1-1.3 mm. Disk prominent, 
glabrous. Ovary ovoid-conical, initially with some 
early caducous substellate scales, or mostly gla- 
brous from the begin, often faintly verruculose, 
5-6 by 2-2.5 mm, ± abrupt with the style, the 
latter c. 1.5 cm, rather slender, glabrous; stigma 
large, capitate, thickly and markedly 5-lobed, 2-3 
mm 0. Capsule thick-fusiform, somewhat oblique 
and curved, glabrous, c. 3 by 0.6 cm. 

Distr. Malesia: Malay Peninsula (Main Range 
from Perak to Selangor, and in the Taiping Rills). 

Ecol. Epiphytic on lofty trees, also terrestrial 
on rocks by stream, in wooded hillside or swamp, 
locally common, but rare in general, 1035-1830 m. 
Fl. fr. Jan.-Dec. 

Vern. Nubang bintang, Selangor, akar, Pahang. 

218. Rhododendron rarilepidotum J. J. S. in Merr. 
Contr. Arn. Arb. 8 (1934) 126; Sleum. Rein- 
wardtia 5 (1960) 191. 

var. rarilepidotum. 

Shrub, 1.5-4 m, said to be often much branched. 
Branchlets subterete, c. 3 mm 0, upper internodes 
laxly lepidote, lower ones glabrescent, not rarely 
pruinose, smooth; internodes 4-11 cm. Leaves 3-6 
in a pseudo whorl at the upper 1-3 nodes, further 
sometimes 1-2 inserted 1-2 cm lower, elliptic or 



Jan. 1966] 



Ericaceae (Sleumer) 



621 



oblong-elliptic, not rarely oblong-lanceolate, apex 
short-acuminate, subobtuse or pointed, base — 
broadly and subacutely attenuate to the petiole, 
± thick-coriaceous, stiff, — densely lepidote at 
both sides on young shoots, quite glabrous above 
at full age (often minutely blackish punctate there), 
laxly to subdensely lepidote for a long time beneath 
(scales small, irregularly substellate-dented or 
-lobed in the marginal zone which disappears with 
age, dark centre rather large, minutely but well 
visibly sunk), finally — glabrescent beneath too, 
edge but little revolute, (4.5-)5-8(-15) by 1.8-3.7 
cm, midrib a continuation of the robust petiole, 
rather flat above, more pronounced and obtusely 
prominent beneath, nerves 6-10(-12) pairs, 
spreading, mostly faintly impressed or rather 
inconspicuous above, slightly raised beneath, 
reticulation lax, faintly prominent beneath only; 
petiole a little flattened, slightly grooved above, 
initially lepidote, 0.7-1.5 cm by 2-3 mm. Umbels 
(6_)8-12-flowered. Ferulae membranous, quite 
glabrous from the beginning, outer ones ovate, 
subobtuse, inner ones ovate-oblong, slightly 
keeled, all early caducous, up to 2.5 by 1.2 cm. 
Bracteoles linear below, subspathulate-dilated 
apically, glabrous, c. 2 cm by 0.5-1 .5 mm. Pedicels 
£: laxly, rarely subdensely lepidote, 1.5-2 cm by 1 
mm. Calyx very oblique, rim-like, wavy, c. 4 mm o. 
Corolla broadly funnel-shaped, 2-2.5 cm long in 
all, c. 3.5 cm at the limb, (dark) red to crimson, 
rarely orange-red, laxly lepidote or mostly gla- 
brous outside, glabrous inside, tube 1-1.3 cm, 
subcylindric in the lower half, obtuse base slightly 
5-saccate, 5-6 mm o, widened to 1-1.5 cm at 
apex, lobes obovate-suborbicular, spreading, 1-1.3 
by c. 1 cm. Stamens 10, unequal, reaching — 
halfway the corolla lobes; filaments linear, gla- 
brous; anthers subelliptic-oblong, 2.5-3.2 by 1.3 
mm, obtuse at the base. Disk very prominent, 
glabrous. Ovary subcylindric-conical, mostly quite 
glabrous from the beginning, rarely ± laxly 
lepidote at first, becoming glabrous during the 
anthesis, 4-6 by 2-2.5 mm, abrupt with the style, 
the latter rather slender, 1.2-1.8 cm; stigma sub- 
capitate, large, 2-3 mm 0, lobes distinct. Capsule 
subcylindric, oblique, slightly curved, glabrous, 
2.5-3 by 0.5-0.7 cm; pedicel up to 4 cm. 

Distr. Malesia: N. Sumatra (Gajo Lands; 
East Coast). 

Ecol. In subalpine scrub or bush, apparently 
always terrestrial, on slopes, (975-) 1800-2500 m. 
Fl. Jan. -Dec. 

Vern. Balingan, pariama, Karo. 

Note. R. rarilepidotum is closely related to R. 
rohinsonii from the Malay Peninsula and mainly 
differs by its smaller, generally red to dark red 
corollas. 

¥ar. ootrichum Sleum. Reinwardtia 5 (I960) 191. 

Differs by the white-tomcntose ovary, broader 
and more shortly acuminate leaves, more densely 
lepidote pedicels and slightly smaller pcrulac. 
Corolla scarlet, very laxly hairy inside. Capsule 
2-2.5 by c. 0.6 cm. 

Dittr. Malesia: Sumatra (East Coast: Mt 



Sibayak and Mt Pinto), 1750-2100 m. 

Note. Occurs together with R. sumatranum 
and is probably a local hybrid of that species with 
typical R. rarilepidotum. 

219. Rhododendron multicolor MiQ. Fl. Ind. Bat. 
Suppl. 1 (1860) 251, 586; Hook. /. Hot. Mag. 
(1884) t. 6769; Henslow, J. R. Hort. Soc. 13 
(1891) 263, f. 42 (/o/.), 43d (/?.), inch var. curtisii 
Henslow, I.e.; Ridl. J. Fed. Mai. St. Mus. 8 
(1917) 58; MiLLAis, Rhodod. (1917) 216; J. J. S. 
m Merr. Contr. Arn. Arb. 8 (1934) 124; Sleum. 
Reinwardtia 5 (1960) 191. — R. salicifolium Bl. 
Fl. Jav. PI. ined. (1863-83) t. 7 C, f. 2, cf. Steen. 
Bull. Jard. Bot. Btzg III, 17 (1948) 388. — R. 
curtisii T. Moore, Florist & Pomologist (1884) 
113, t. 615; Anon. Garten-Zeitung Berlin 3 (1884) 
469, fig.; Anon. 111. Gartenzeitung Stuttgart 30 
(1886) t. 10. — Azalea multicolor (MiQ.) O. Ktze, 
Rev. Gen. PI. 2 (1891) 387. — R. javanicum [non 
(Bl.) Benn.] Steen. Arch. Hydrobiol. Suppl. 11 
(1932) 318. 

Shrub, epiphytic or mostly terrestrial, 1-1.5 m. 
Branches lax. Branchlets slender, terete, upper 
l(-2) internodes reddish and c. 2 mm 0, laxly to 
subdensely set with flat, substellate scales, older 
parts glabrescent; internodes 1.5-4(-7) cm. Leaves 
in 3-5-merous pseudowhorls at the upper 1-2 
nodes, lanceolate to oblong-lanceolate, apex ± 
shortly acuminate, subacute, base ± broad- 
attenuate to the petiole, sometimes -: obtuse, 
lb subcoriaceous, subdensely lepidote initially on 
both faces, glabrescent above, lepidote for a long 
time beneath (scales small, irregularly lobed or 
dented at the marginal zone, dark centre minute, 
but shallowly impressed), edge flat, (3-)4-7 by 
(0.7-)i-i.5(-2, very rarely up to 2.5) cm, midrib 
rather robust and prominent in the lower part 
on both sides, nerves 6-8 pairs, rather straight and 
ascending, indistinctly anastomosing before the 
edge, very slightly raised on both faces, often 
almost obsolete; petiole flattened dorsally, 2-6 by 
1-2 mm. Umbels (3-)4-6-flowered. Ferulae ovate- 
oblong to oblong, membranous, obtuse, glabrous 
and shining, up to 2.2 by 0.7(-0.9) cm. Bracteoles 
linear, glabrous, up to 1.5 cm by 1 mm. Pedicels 
stoutish, very sparsely lepidote or glabrous, 
1.8-2.5(-3) cm. Calyx oblique, disk-shaped or 
rim-like, c. 4 mm 0, obtusely angled or very 
shortly lobed, sometimes acute and up to 1 mm. 
Corolla broad-funnel-shaped, 1.5-2(-2.5) cm long 
in all, sometimes sweet-scented, whitish cream to 
yellow, rose or fiery red, lobed ± halfway, gla- 
brous outside, short-pubescent at the tube inside, 
tube c. 5 mm at the 5-pouchcd obtuse base, 
c. I cm at apex, lobes somewhat expanded, 
broadly obovate-spathulate, 1-1.3 by 0.7-1 cm. 
Stamens 10, unequal, as long as or a little longer 
than the corolla tube; filaments linear, densely set 
with rather longish. spreading, white hairs in the 
lower Vi-'/j. glabrous for the rest; anthers broad- 
oblong to obovatc, 1.5-2 by 1 mm. Disk promi- 
nent, glabrous. Ovary obovoid-cylindric, glabrous, 
slightly rugose, r. 4 by 2 mm, abrupt with the style. 
the latter slender, glabrous. (7-)8-10 mm; stigma 



622 



Flora Malesiana 



[ser. I, vol. 6* 



obconical. Capsule subcylindric, 1.8-2.2 by 0.5 cm. 

Distr. Malesia: Sumatra. 

Ecol. Mostly terrestrial in open places, hillside 
or pastures, also near solfataras, sometimes 
epiphytic in mountain forest, at (900-) 1000-1 700 
(-2100) m, locally common, both on fertile soil 
and stony ground, volcanic tuff, on rocks. Fl. 
Jan. -Dec. 

Vern. Sarodang, Toba, karamunting-rimbo- 
-njang djanten, Batangbaru, M. 

220. Rhododendron brevitubum J. J. S. Ic. Bog. 4 
(1914) 253, t. 381; Merr. En Born. (1921) 460; 
Sleum. Remwardtia 5 (1960) 192. 

Epiphytic shrub. Branchlets terete, 3.5-6 mm 
0, glabrous, upper 2 internodes red brown, 3-5 
cm. Leaves mostly 2 subopposite at the upper 
2(-3) nodes, and 1 or 2 scattered on the inter- 
nodes, broad-elliptic or subovate-elliptic, apex 
broad-attenuate, obtuse, base rounded or r: 
distinctly cordulate, — stiff-coriaceous, finally 
glabrous above, laxly to subdensely lepidote 
beneath for a long time (scales minute, irregu- 
larly dented or lobed in the marginal zone, which 
is early discarded, dark centre slightly impressed), 
finally glabrescent underneath too, edge somewhat 
recurved, (7-) 8-12 by 3.7-7.5 cm, midrib robust, 
much broadened, usually much prominent on 
both faces, nerves 10-12 pairs, at an angle of c. 
70°, rather straight below, slightly curved distally 
and ± anastomosing before the edge, all i 
parallel, little but conspicuously raised on both 
sides, veins and veinlets forming a rather dense 
network; petiole much flattened and a little convex 
above, 7-12 by 3^ mm, sparsely lepidote or 
glabrous. Umbels 3-4-flowered. Ferulae ovate, 
obtuse, glabrous, up to 1.7 by 1 cm in an immature 
flower bud. Pedicels glabrous, c. 1 cm by 1.5 mm. 
Calyx obliquely and very shortly cup-shaped, 
subentire, wavy, glabrous, c. 6 mm 0. Corolla 
broad-funnel-shaped, c. 5.5 cm long in all, c. 
8 mm at the much expanded limb, rather 
fleshy, glabrous outside, densely pubescent at the 
tube inside along 5 vertical stripes, tube subcylin- 
dric, c. 1.5 cm long, c. 1.3 cm at the 5-saccate 
base, widened to c. 2 cm at apex, lobes broad- 
obovate-spathulate, 3.5-3.7 by 2.4-3.1 cm. 
Stamens 10, unequal, 2.2-3 cm; filaments filiform 
and glabrous in the upper, yellowish villous in 
the lower dilated half, 1.7-2.5 cm; anthers linear, 
curved, versatile, 0.6-0.8 by 1.3 mm. Disk puber- 
ulous on the upper margin, otherwise glabrous. 
Ovary cylindric-conical, 5-ribbed, c. 9 by 4 mm, 
rather abrupt with the style, the latter thickish, 
glabrous, c. 1.5 cm; stigma capitate, distinctly 
lobed. 

Distr. Malesia: Central Borneo (Liang Gagang 
and B. Mi'li). Fl. Nov. 

221. Rhododendron sessilifolium J. J. S. in Merr. 
Contr. Arn. Arb. 8 (1934) 125; Sleum. Rein- 
wardtia 5 (1960) 192. 

Terrestrial, or rarely epiphytic shrub, 1-3 m. 
Branchlets terete, brown when dry, 2-3 mm 0, 
upper 1-2 internodes sparsely to subdensely 



covered with flat, rather early caducous scales, 
smooth; internodes 4-10 cm. Leavej in i 5-merous 
pseudowhorls at the upper 1-2 nodes, sessile or 
subsessile, lanceolate to oblong-lanceolate, some- 
times subobovate-oblong, apex gradually or 
rarely abruptly acuminate, subacute or more 
bluntish, base broadly attenuate to rounded or 
subtruncate, occasionally slightly cordate, coria- 
ceous, stiff, finally glabrous above, lepidote for 
a long time beneath (scales rather small, irregularly 
stellate-dented at the marginal zone, dark centre 
shallowly or not impressed), finally glabrescent 
beneath too, but remaining brownish punctulate, 
very edge somewhat revolute, (8-)10-17.5 by 
2.4-4(-5) cm, midrib as thick as the petiole, obtuse 
or i flattened, c. 2 mm wide at the very base, 
nerves 10-13 spreading pairs, straight below, 
curved upwards, but indistinctly anastomosing, 
very slightly impressed above in old leaves, or 
mostly a little raised on both faces, not rarely 
obsolete, reticulation mostly obscure; petiole 0-4 
(rarely up to 5) by 2-3 (-4) mm. Umbels (4-) 
6-8(-10)-flowered. Flower buds ovoid, 2-2.5 by c. 
1.5 cm, outer perulae ovate, obtuse, inner ones 
ovate-oblong to oblong, obtuse, glabrous, up to 
2.5 by 1.2 cm. Bracteoles subfiliform, glabrous, 
up to 1.5 cm. Pedicels laxly stellate-lepidote, 
epilose, 2.5-3 cm by 1-1.5 mm. Calyx oblique, 
rim-like, obscurely lobed, c. 4 mm 0. Corolla 
broad-funnel-shaped, (3-)3.5-4 cm long in all, 
a clear to light orange yellow (possibly occasion- 
ally also red?), glabrous outside, very laxly 
and shortly pubescent inside, tube 1.5-2 cm long, 
base 5-saccate, 5-6 mm 0, c. 1.5 cm at apex, 
lobes obovate, 1.4-1.8(-2) by 1-1.5(-1.7) cm. 
Stamens 10, subequal, attaining i halfway the 
corolla lobes; filaments linear and densely sub- 
patent-hairy in the lower 14. glabrous above; 
anthers broad-oblong, c. 4 by 1.3 mm. Disk 
prominent, much lobed, glabrous. Ovary thick- 
-cylindric, a little narrowed towards the base, with 
some scales initially, early glabrescent, or glabrous 
from the beginning, 5-7 by 2.5-3 mm, rather 
abrupt with the style, the latter ± excentric, 
glabrous, — equalling the stamens; stigma sub- 
capitate. Capsule subcylindric, 2.5-3 by 0.6-0.7 
cm. 

Distr. Malesia: Sumatra (Tapanuli, East 
Coast, West Coast). 

Ecol. In light jungle or open places, on rim of 
crater, hills, 1100-1980 m. Fl. Jan.-Dec. 

222. Rhododendron beccarii Sleum. Reinwardtia 

5 (1960) 192. 

Epiphytic, apparently rather small shrub. 
Branchlets terete, 4-8 mm 0, uppermost internodes 
with lax substellate, appressed scales, lower parts 
glabrous, smooth, not rarely pruinose; internodes 
(5-)7-17 cm. Leaves in 5-8-merous pseudowhorls 
at the upper 2-3 nodes, oblong or lanceolate- 
oblong, apex gradually long-acuminate (2-3 cm), 
acute or nearly so, ± broadly attenuate towards 
the base, very base subtruncate, sessile or more 
rarely subsessile, coriaceous, stiff, quite glabrous 
with age, but remaining blackish punctulate above. 



Jan. 1966] 



Ericaceae (Sleumer) 



623 



rather laxly subpersistently lepidote beneath 
(scales small, irregularly substellate-dented or 
-lobed in the marginal zone, centre minutely im- 
pressed), finally glabrescent and punctate beneath 
too, margin slightly revolute, 10-16 by 3-5(-6) cm, 
midrib thick, 2-3 (-4) mm wide and obtusely 
prominent on both faces below, nerves 10-12 
obliquely ascending pairs, faintly interarching 
before the edge, little raised on both sides, some- 
times hardly visible, veins and veiniets subobscure; 
petiole compressed dorsally, 0-2 by 2-4 mm. 
Umbels 4-6-flowered. Flower buds ovoid-acumi- 
nate, 3-3.5 by 1-1.5 cm, outer perulae ovate, 
acuminate, inner ones oblong-ovate, all subacute, 
up to 2.5 (rarely up to 3) by c. 1 cm, glabrous 
dorsally, margin initially with caducous scales. 
Bracteoles filiform, glabrous, up to 2.5 cm by 0.5 
mm. Pedicels laxly lepidote, 2-3 cm. Calyx much 
oblique, disk-shaped, obscurely lobed, subglabrous, 
c. 4 mm 0. Corolla funnel-shaped, red or orange, 
(5-)5.5-6.5 cm long in all, glabrous outside, i 
densely shortly white-hairy in the lower half of the 
tube inside, tube 3-3.5 cm long, base thickened, 
5-saccate, 5-6 mm o, subcylindric in the lower, 
gradually widening in the upper part to 1.5-2 cm, 
lobes obovate-spathulate, 2-3 by 1.5-2 cm. 
Stamens 10, unequal, the longest nearly as long 
as the corolla; filaments linear and very densely 
subpatently white-hairy below, filiform and gla- 
brous distally; anthers oblong, base obtuse, c. 

4 by 1 mm. Disk prominent, glabrous. Ovary sub- 
cylindric, glabrous or very laxly lepidote, tapering 
to the style, c. 10 by 2.5 mm; style nearly as long 
as the corolla tube, glabrous; stigma subcapitate. 
Capsule subfusiform, red, slightly curved, glabrous, 
(4_)4.5_5.5(_6) by 0.6-0.7 cm; pedicel 3^ cm 
by 1-1.5 mm. 

Distr. Malesia: Sumatra (West Coast: Mt 
Singgalang, Mt Sago). 

Ecol. In mountain forest, mossy forest or 
thickets, 1400-2450 m. Fl. Jan.-Aug.,/r. June-Aug. 

223. Rhododendron kemulease J. J. S. Bull. Jard. 
Bot. Btzg III, 13 (1935) 448; Sleum. Reinwardtia 

5 (I960) 193. 

Small, crumpled tree, trunk c. 35 cm a. Branch- 
lets terete, 3-4 mm 0, red brown, with lax, small, 
substellate, flat scales at the upper internodes or 
the very tips, early glabrescent below; internodes 
2-8.5 cm. Leaves in 5-7-mcrous pseudowhorls at 
the upper 1-2 nodes, oblong- to lanceolate- 
elliptic, apex gradually attenuate or shortly 
acuminate, subacute or obtuse, base acutely 
narrowed to the petiole, coriaceous, densely 
minutely lepidote on both faces when very young, 
glabrous and • shining above at maturity, more 
persistently lepidote beneath (scales minute, 
marginal zone irregularly dented and early dis- 
solved, dark centre shallowly impressed), finally •: 
glabrescent beneath too, but remaining conspicu- 
ously punctulatc there, margin faintly recurved, 
7-12 by 2-4 (rarely up to 4.5) cm, midrib narrow 
and sunk above, as broad as the petiole and 
obtusely prominent in the lower part beneath, 
nerves 7-10 pairs, irregular, ascendmg, obscurely 



anastomosing, very slightly impressed above, 
slightly prominent or subinconspicuous beneath, 
no reticulation; petiole subterete, rugose, grooved 
above, minutely lepidote or glabrescent, (4-) 
8-15 by 2-2.5 mm. C/m6Wi 7-12-flowered. Bracte- 
oles linear, laxly pubescent, 2-3 cm by 0.5-1 mm. 
Pedicels laxly lepidote at both ends, otherwise 
very laxly short-pubescent, ± glabrescent, 2-2.5 
cm by nearly 1 mm. Cu/v.vsuboblique, disk-shaped, 
obscurely 5-angular, minutely lepidote, 4-5 mm 
0. Corolla narrow-funnel-shaped, salmon coloured, 
sparsely minutely lepidote outside, short-pubescent 
at the lower half of the tube inside, 5-6(-7) cm 
long in all, limb 3-4 cm o, tube rather straight, 
3-4 cm long, base 3-4(-5) mm 0, c. 1 cm at apex, 
lobes broadly oblong-obovate, 2-2.5(-3) by 1.5-2 
(-2.5) cm. Stamens 10, subequal, 4-5 cm long; 
filaments linear and densely patently short-hairy 
in the lower third, narrower to filiform and 
glabrous distally; anthers oblong, 3-4 by 1 mm, 
base obtuse. Disk puberulous at the upper margin, 
glabrous for the rest. Ovary subcylindric, lepidote 
especially at the base and apex, early glabrescent, 
c. 6 by 2.5 mm, continuous with the style, the 
latter nearly as long as the stamens, subdensely 
lepidote below, more laxly so up to the middle, 
otherwise glabrous; stigma capitate. 

Distr. Malesia: Central Borneo (Mt Kemul), 
in primary forest, said to be common locally, 
1800-1850 m. Fl. Oct. 

224. Rhododendron moultonii Ridl. J. Str. Br. R. 
As. Soc. /(. 63 (1912) 61; Merr. En Born. (1921) 
462; Sleum. Reinwardtia 5 (1960) 193. 

Small, terrestrial shrub with grey bark, c. 3 m. 
Branchlets 4-7 mm 0, upper internode obtus- 
angular and laxly to very laxly lepidote, lower ones 
terete and glabrous; internodes 10-20 cm. Leaves 
at the upper 1-2 internodes, 2(or 3) of them 
rz opposite or in a pseudowhorl, 2 or 3 others 
scattered along the internode, elliptic to elliptic- 
oblong, apex shortly (1-2 cm) acuminate or 
obtusely attenuate, base broad-attenuate to sub- 
truncate rounded, coriaceous, rigid, glabrous 
above at full age, laxly to rather subdensely 
lepidote beneath (scales small, irregularly sub- 
stellate-lobcd or -dented in the marginal zone, dark 
minute centre but shallowly impressed, leaving 
dark points on the undersurface, edge h flat, 
13-20(-25) by (4-)5-8 cm, midrib a continuation 
of the very robust petiole in its lower '/j, c. 4 mm 
wide, rather flat above, much and obtusely raised 
beneath, nerves (1 6-) 1 8-20 pairs, narrow and 
\: parallel, spreading at a wide angle, anastomos- 
ing near the edge, prominent on both faces, reticu- 
lation rather dense and distinctly raised on both 
sides; petiole somewhat flattened, initially lepidote, 
(0.8-)l-1.4 cm by 3-4 mm. Umbels 7 9-flowcred. 
Pedicels (very) laxly lepidote to glabrescent, 
2-3 (rarely up to 4) cm by 1.5 mm. Calyx sub- 
oblique, thick, disk-shaped, c. 5 mm 0, lobes very 
short, rounded, often hardly distinguished. 
Corolla funnel-shaped, yellowish orange or yellow, 
flushed pink outside on the lobes, 4 4.5 cm long 
in all. 5-lobcd to '/§-*/». glabrous outside, equally 



624 



Flora Malesiana 



[ser, I, vol. 6* 



laxly short-hairy at the lower part of the tube 
inside, tube 1.5-2.5 cm long, base 5-pouched, 5-6 
mm 0, widened to c. 1 .5 cm at apex, lobes obovate- 
spathulate, 1.5-2.5 by c. 1.5 cm. Stamens 10, c. 
3 cm; filaments linear and shortly patent-pubescent 
below, filiform and glabrous towards the top; 
anthers oblong, slightly curved, 5-6 by 1.2 mm. 
Disk prominent, glabrous. Ovary subcylindric- 
conical, suddenly narrowed to the style, glabrous, 
finely rugose, c. 6 by 2.5 mm; style glabrous, 
rather thick (c. 0.5 mm), c. 1.4 cm long; stigma 
capitate-obconical. 

Distr. Malesia: Borneo (Sarawak). 

Ecol. Growing on the ground in mossy forest, 
1220-1525 m. Fl. May, Oct. 



(1891) 262, 281, 282, f. 42 (/o/.), 43 e (/?.); KooRD. 
Teysmannia 10 (1899) 450; Nat. Tijd. N.I. 60 
(1901) 264; Wigman, Teysmannia 20 (1909) 112, 
t.; KooRD. Jungh. Gedenkb. (1910) 184; Exk. Fl. 
Java 3 (1912) 3, f. 1 ; Koord.-Schum. Syst. Verz. 1 
(1912) fam. 233, p. 106; J. J. S. in K. & V. Bijdr. 
13 (1914) 96; Ridl. J. Fed. Mai. St. Mus. 8 (1917) 
57; MiLLAis, Rhodod. (1917) 196; Koord. Fl. 
Tjib. 3 (1918)fam. 233,p. 2;Sp. Moore, J. Bot. 63 
(1925) Suppl. 57; Hochr. Candollea 2 (1925) 490, 
incl. var. gymnocarpum Hochr. I.e.; Steen. Bull. 
Jard. Bot. Btzg III, 12 (1932) 177; ibid. Ill, 13 
(1933) 50; Amsh. in Back. Bekn. Fl. Java (em.ed.) 
7 (1948) fam. 162, p. 2; Sleum. Reinwardtia 5 
(1960) 194; Steen. Endeavour 21 (1962) 189, t. 1; 




Fig. 38. Rhododendron javanicum (Bl.) Benn. Mt Gedeh, W. Java (Bruggeman). 



225. Rhododendron javanicum [Reinw. Verh. Bat. 
Gen. 9 (1823) 30, nom. nud.; in Bl. Cat. (1823) 72, 
nom. nud.] (Bl.) Benn. in Benn. & Br. PI. Jav. Rar. 
(1838) 85, excl. t. 19, which is var. teysmannii; DC. 
Prod. 7 (1839) 721; Hassk. Cat. Hort. Bog. 
(1844) 161, incl. var. citrinum Hassk. I.e.; Moritzi, 
Syst. Verz. (1846) 42; Paxton, Mag. Bot. 15 
(1849) n. Ill; Planch. Fl. Serres 5 (1849) 477-480; 
HouTTE, ibid. 6 (1850-51) 135, t. 576; Zoll. Syst. 
Verz. 2 (1854) 137; Miq. Fl. Ind. Bat. 2(1859) 1057; 
Suppl. 1 (1860) 251, 585; Ann. Mus. Bot. Lugd.- 
Bat. 1 (1863) 42, t. 1 B; Bl. Fl. Jav. PI. inM. 
(1863-83) t. 1; Henslow, J. R. Hort. Soc. 13 



Sleum. Blumea 12 (1963) 111. — Vireya javanica 
Bl. Bijdr. (1826) 854, incl. var. I.e. 855; G. Don, 
Gen. Syst. 3 (1834) 848. — Azalea javanica 
(Bl.) O. Ktze, Rev. Gen. PI. 2 (1891) 386. — 
Fig. 38. 

KEY TO THE VARIETIES 

1 . Ovary very sparsely lepidote and early glabres- 

cent, or glabrous from the beginning. 
2. Leaves predominantly in pseudowhorls at or 
in part inserted a little below the nodes, 
occasionally some additional scattered ones 



Jan. 1966] 



Ericaceae (Sleumer) 



625 



in the upper third of the internodes, apex ± 
acuminate. Anthers (2.5-)3-3.5 (rarely up to 

4) mm 1. var. javanicum 

2. Leaves exclusively scattered along the inter- 
nodes, apex — obtuse. Anthers 5-6(-7) mm. 
2. var. schadenbergii 
1. Ovary — densely pubescent 3. var. teysmannii 

1. var. javanicum. 

Shrub, 1-2 m, or treeiet, 2-3(-5) m, trunk up to 
8 cm 0, erect or writhed, terrestrial or epiphytic, 
generally not much branched; bark ashy. Branch- 
lets 3-5 mm o, young shoots obtusangular and 
laxly substellate-lepidote, older parts terete, 
glabrescent, smooth; internodes 4-15 cm. Leaves 
in (4— )5-7(-8)-merous pseudowhorls at the upper 
l-2(-3) nodes, occasionally not strictly so, 
broadly oblong- to subovate- or subelliptic- 
oblong (occasionally ovate- or obovate-elliptic), 
apex mostly acuminate, subacute, more rarely 
bluntish, base mostly ± acutely and rather 
shortly attenuate, ± coriaceous, initially ± 
densely lepidote on both faces, early glabrescent 
above, laxly to subdensely lepidote beneath for a 
certain time (scales small, flat, variously lobed or 
dented at the marginal zone, dark centre but 
faintly impressed), brownish blackish punctulate 
beneath after the scales are gone, margin dz flat, 
4-1 5(-20) by (1 .8-)2.5-6(occasionally up to 8) cm, 
midrib — a continuation of the robust petiole, 
broad and obtusely prominent to various degree 
in the lower part on both faces, ± narrowly 
impressed above, nerves 7-10 pairs, at an angle 
of c. 45', slightly prominent on both sides, rather 
indistinctly anastomosing before the edge, veins 
and veinlets a lax network, faintly raised or often 
obsolete on both sides; petiole compressed 
dorsally, not or hardly grooved above, brownish 
reddish as is often the midrib, initially lepidote, 
early glabrescent, (0.4-)0.7-2(-2.3) cm by 2-3 mm. 
Umbels 4-7(-12)-flowered. Flower buds ovoid, 
subacuminate, c. 3-3.5 by 1.5-2 cm. Outer 
perulae broad-ovate, shortly subulatc-mucronate, 
inner ones ovate-elliptic to oblong, obtuse, all 
membranous, glabrous, margin with minute 
early caducous scales, up to 3.5 by 2.2 cm. 
Bracteoles linear, glabrous, up to 3 cm by 0.5-1.5 
mm. Pedicels laxly stellate-lepidote especially up- 
wards, epilose, glabrescent, (l.5-)2-4 cm by c. 1 
mm. Calyx oblique, disk-shaped, obtusely 5- 
angular, c. 4 mm 0, glabrous or very laxly lepidote. 
Corolla funnel-shaped, rather fleshy, occasionally 
yellow or red (even scarlet), mostly ± orange, 
the lube often reddish or rose inside or with a 
ring of deep pink round the throat, scentless, 
(3-)4-5(-5.5, rarely up to 6) cm long and wide in 
all, - halfway 5-lobed, glabrous or nearly so 
outside, shorl-pubcsccnt to various degree at the 
tube inside (occasionally glabrous there too), 
tube (1.7-)2-2.5 cm long, obtusely 5-angular, 
base 5-saccatc, 5-6 mm a, widened to 1-1.5 cm 
at apex, lobes slightly unequal, expanded, obo- 
vatc to subspathulatc. 2.5-3.5 by 1 .7-2.5 (rarely up 
to 3) cm. .Sfameni 10, alternately slightly unequal, 
the longer ones reaching ; half the corolla lobes; 



filaments purplish, linear, ± densely pubescent in 
the lower third, more filiform and glabrous to- 
wards the top; anthers oblong to broad-oblong, 
sometimes more obovate-oblong, (2.5-)3-3.5 
(rarely up to 4) by c. 1.3 mm. Disk prominent, 
puberulous at the upper margin only. Ovary 
obliquely conical-cylindric, 5-angled, glabrous or 
very sparsely lepidote (and early glabrescent then), 
7-9 by c. 3 mm, but little abrupt with the style, 
the latter glabrous, ± as long as the shorter 
stamens; stigma capitate, c. 3 mm 0. Capsule 
elongate-cylindric, shortly attenuate at both ends, 
5-angled, glabrous, (3-)3.5-4.5 (rarely up to 5) 
by 0.7-0.8 (rarely up to 0.9) cm; pedicel up to 
5 cm by 1.5-2 mm. Seeds including tails 6-7 mm 
long. 

Distr. Malesia: Sumatra, Natuna I., Lingga 
Arch., Java, Bali. 

Ecol. Mostly epiphytic and in primary and 
secondary forest, also near craters, on narrow 
ridges, in rather dry ground, between rocks, on 
steep slopes, once found on stony mud-flow in 
S. Sumatra, (30O-)80O-24O0(-2550) m, locally 
common. Fl.fr. Jan.-Dec, with preference for the 
dry season. 

Vern. KSmbang gSni, k. klpiting, k. sademo, 
lumpeni, purwa gSni, soloko baros, soro sari, 
tjSpoko gSni, J, areuj gandarusa, djudjugan, hi 
badak, loda dindSr, I. dlnlr, songgoni tangkal, 
tjantigi bSsar, rjawareni sore, tjonkok, S, bunga sa 
rudang, b. tempa barSn, M. 

2. var. schadenbergii (Warb.) Sleum. Reinwardtia 
5 (1960) 195. — R. schadenbergii Warb. in Perk. 
Fragm. Fl. Philip. 3 (Feb. 1905) 172; Merr. 
Philip. J. Sc. 3 (1908) Bot. 380, p.p.; Millais, 
Rhodod. (1917) 239; Merr. En. Philip. 3 (1923) 
245, p.p.; CoPEL. /. Philip. J. Sc. 40 (1929) 170, 
t. 15, f. 2, t. 16, f. 3^. — R. spectabile Merr. 
Publ. Gov. Lab. Philip, n. 29 (Sept. 1905) 42; 
Philip. J. Sc. 3 (1908) Bot. 381; Elm. Leafl. 
Philip. Bot. 3 (1911) 1106; Millais, Rhodod. 
(1917) 246; Merr. En. Philip. 3 (1923) 245; 
Millais, Rhodod. ser. 2 (1924) 241; Copel. /. 
Philip J. Sc. 40 (1929) 168, t. 13, f. 4-5, t. 14. — 
R. sarasinorum Warb. in Sarasin, Reisen in 
Celebes 2 (1905) 386 {index), cf. ibid. 1 (1905) 62 
(text), nom. nud. — R. dementis Merr. Philip. 
J. Sc. 3 (1908) Bot. 160, 381; En. Philip. 3 (1923) 
243; Millais, Rhodod. ser. 2 (1924) 113; Copel./. 
Philip. J. Sc. 40 (1929) 167, t. 12, t. 13, f. 1-3. 

Leaves exclusively scattered along the upper 
internodes, apex h obtuse. Corolla red, orange 
or salmon pink, very rarely yellow. Anthers 5-6 
(-7) mm. 

Distr. Malesia: NE. Celebes (Mcnado), 
Philippines (Luzon; Camiguin dc Misamis; Jolo; 
Mindanao; Palawan). 

Ecol. As in var. javanicum. 

Vern. Philippines: malagos. Bag., lasatan, 
Bon., luhung di kamaduyung. If. 

Uses. Flowers said to be edible. 

3. var. tcywrnannii (Mi<j.) K. & G. J. As. Soc. Beng. 
74. li (1905) 75; J. J. S. in K. & V. Bijdr. 13 (1914) 



626 



Flora Malesiana 



[ser. I, vol. 6* 



99; Steen. Bull. Jard. Bot. Btzg III, 13 (1933) 
50; ibid. Ill, 17 (1948) 388; Sleum. Reinwardtia 
5(1960) 195; Blumea 12(1963) \\\. — R. javani- 
cum (Bl.) Benn. in Benn. & Br. PI. Jav. Rar. 
(1838) pro t. 19; Hook. Ic. PI. (1847) t. 4336; 
Lemaire, FI. Serres 3 (1847) 293, c. icon.; Bl. F1. 
Jav. PI. in^d. (1863-83) t. 2; HocHR. Candollea 
2 (1925) 490 {var. typicum). — R. teysmannii MiQ. 
FI. Ind. Bat. Suppl. 1 (1860) 251, 585; Ann. Mus. 
Bot. Lugd.-Bat. 1 (1863) 42, t. 1 A; Clarke in 
Hook./. FI. Br. Ind. 3 (1882) 463; Henslow, J. 
Hort. See. 13 (1891) 263, f. 42 (/o/.), 43 f (^.); 
RiDL. J. Str. Br. R. As. Soc. n. 23 (1891) 146; 
J. Fed. Mai. St. Mus. 4 (1909) 42; /6/^. 7 (1916)46; 
ibid. 8 (1917) 58; Millais, Rhodod. (1917) 251; 
Henders. Mai. Nat. J. 6 (1950) 261. — Azalea 
teysmannii (MiQ.) O. Ktze, Rev. Gen. PI. 2 (1891) 
387. — R. basirotundatiim J. J. S. in Merr. Contr. 
Arn. Arb. 8 (1934) 124, p.p. typ. 

Differs from \ar. javanicum only by the ± 
densely pubescent ovary. Pedicels nearly always 
pubescent. 

Distr. Malesia: Sumatra, Malay Peninsula, 
Java, Bali. 

Ecol. As in var. javanicum, apparently occurring 
rarely together with it on the same spot. 

Vern. Areng marenang, kakuran, kembang 
tjulang, S. 

Note. Part of the specimens collected in North 
Sumatra bear capsules 5.8-8 cm long, thus 
distinctly longer than stated above for R. javanicum 
sens. lat. 

In these long capsules they agree with R. 
beccarii, also from Sumatra (East Coast), which, 
however, has a practically glabrous (or certainly 
not hairy) ovary. 

It is possible that these North Sumatran speci- 
mens correspond with R. basirotundatum described 
from Atjeh (Bur ni Telong) without fruits and 
might represent a fourth variety. Material at 
hand is not sufficient to decide this question. 

R. teysmannii was figured by Blume and Miquel 
as having yellow flowers. This, I presume, is an 
error by an omission of the draughtsman, similarly 
as was made for R. album; there is no difference 
in colour with R. javanicum. — Ed. 

226. Rhododendron perplexum Sleum. Reinwardtia 
5 (1960) 197. 

Shrub, probably epiphytic. Branchlets terete, 
c. 4 mm 0, red-brown, glabrous; the one known 
(upper) internode 10 cm. Leaves in a ± 4-merous 
pseudowhorl, ovate to oblong-ovate, gradually 
acuminate towards the subacute apex, base 
rounded to slightly cordate, or a little inequi- 
lateral, thick-coriaceous, glabrous, subdensely 
lepidote beneath (scales minute, irregularly 
substellate-lobed or -incised in the marginal 
zone, much darker centre slightly impressed), 
margin a little revolute, 8.5-13.5 by 4-5.5 
cm, broadest slightly below the middle, midrib 
very stout, striate lengthwise, being a conti- 
nuation of the thick petiole, obtusely and much 
prominent in the lower part above, broad and 
rather flat beneath, nerves 12-15 pairs, at an 



angle of nearly 90°, ± parallel, obscurely in- 
arching before the edge, very slightly immersed 
above, not much raised beneath, reticulation lax, 
subtle on both faces; petiole slightly flattened and 
grooved dorsally, initially lepidote, 4-6 by c. 3 
mm. Umbels \- 9-flowered. Pedicels rather robust, 
laxly or very laxly lepidote, more densely so 
immediately below the calyx, not hairy, 2-2.7 cm. 
Calyx disk-shaped, obscurely lobed or crenate, 
thick, glabrous, 4-5 mm 0. Corolla funnel-shaped, 
2.5-2.7 cm long in all, glabrous outside, densely 
shortly grey-pubescent at the lower 2/3 of the tube 
inside, tube 1-1.3 cm long, at base c. 5 mm 0, 
widened to c. 1 cm at apex, lobes oblong-obovate, 
1.3-1.5 by 1-1.3 cm. Stamens 10, subequal, nearly 
as long as the corolla; filaments linear and very 
densely subpatently white-hairy in the lower half, 
more filiform and glabrous upwards; anthers 
broad-oblong, 3.5 by 1.3 mm. Disk prominent, 
glabrous. Ovary subcylindric, glabrous, 5-6 by c. 

2 mm, tapering to the style, the latter columnar, 
glabrous, c. 6 mm; stigma subcapitate. 

Distr. Malesia: Sumatra (West Coast: Kerin- 
tji) at c. 1000 m. One collection. 

227. Rhododendron crassifollum Staff, Trans. 
Linn. Soc. II, Bot. 4 (1894) 195; Gibbs, J. Linn. 
Soc. Bot. 42 (1914) 102; Merr. En. Born. (1921) 
460; Sleum. Reinwardtia 5 (1960) 198; Blumea 
11 (1961) 127; ibid. 12 (1963) 111; Jahrb. Deutsch. 
Rhodod. Ges. Bremen (1965) f. 19. — R. crassi- 
nervium Ridl. J. Str. Br. R. As. Soc. n. 63 (1912) 
61; Merr. En. Born. (1921) 461. — R. murudense 
Merr. Sarawak Mus. J. 3 (1928) 542. 

Mostly epiphytic, sometimes terrestrial shrub, 
(0.5-)1.2-3.5 m; bark fawn, smooth. Branchlets 
(3-)4-6 mm 0, upper 1-2 internodes dark red- 
brown, initially scantily lepidote, early glabrescent; 
internodes (1.5-)3-15 cm. Leaves opposite or 
mostly in 3(-4)-merous pseudowhorls at the 
upper 1-3 nodes, often not strictly so and inter- 
nodes often with scars of scattered reduced and 
very early caducous leaves, elliptic, oblong- 
elliptic or oblong, occasionally obovate-elliptic, 
apex in general broadly attenuate and obtuse or 
rounded, sometimes shortly obtusely acuminate, 
base rounded or subtruncate, not rarely slightly 
subauriculate-cordate, very rarely cuneate, thick- 
coriaceous, stiff, initially lepidote on both faces, 
glabrous and blackish punctulate with age above, 
lepidote for a certain time beneath (scales rather 
lax, small, flat, irregularly dented or lobed in the 
marginal zone, centre minute, faintly or not sunk), 
glabrescent with age beneath too, the scales 
leaving dark points, flat, 8-14(-18) by 4-8(-9) cm, 
midrib continuing the petiole, mostly much and 
± bluntly prominent and hardly grooved above, 
more flatfish and hardly raised, or even a little 
sunk towards the apex of the lamina beneath, c. 

3 mm wide at the base, often a little rugose, nerves 
8-12 pairs, subparallel at a wide to almost right 
angle, anastomosing before the edge, distinctly 
raised on both faces, reticulation rather dense, 
slightly prominent above, less or obscurely so 
beneath; petiole somewhat flattened, 1.2-1.8 cm 



Jan. 1966] 



Ericaceae (Sleumer) 



627 



by 3-4 mm. Umbels compact, (8-)10-14 (some- 
times up to 20)-flowered.. Ferulae ovate-acuminate, 
glabrous, up to 2.5 by 1 cm. Bracteoles linear, 
glabrous, up to 2 cm. Pedicels — glabrous. 3-4.5 
by 1(-1.5) mm. Calyx very oblique, patellar, 
obtusely and very shortly lobed or crenate, gla- 
brous, 3-4 mm o. Corolla funnelform-sub- 
campanulate, rather fleshy, flame red or scarlet, 
orange or apricot, salmon or bright shell pink, 
margin and ape.x of the lobes not rarely deep rose. 
{2-)2.5-3 cm long in all, glabrous outside, sparsely 
hairy inside the tube, tube 1.2-1.5 cm, at base dis- 
tinctly 5-pouched, 3-4 mm o, gradually widened 
to (l-)l.2-1.5 cm at apex, lobes obovate-oblong, 
(l-)1.2-l.5 by 0.8-l(-1.3) cm. Stamens 10, 
alternately a little unequal, (1. 2-) 1.4-2 cm; 
filaments linear and densely subpatently hairy in 
the lowest V4-\'5- filiform and glabrous for the 
rest; anthers subelliptic, c. 2-2.5 (rarely up to 2.8) 
by 1.3 mm. Disk prominent, glabrous. Ovary 
subcylindrical-conical, glabrous or initially 
sparsely lepidote, 3-4 by 2 mm, rather abrupt 
with the style, the latter columnar, glabrous, as 
long as the ovary; stigma thick, dilated-capitate, 
2 mm 0. Capsule thick-cylindric, 1.8-2.5 by c. 0.5 
cm. Seeds very narrow, 4-5 mm including the 
tails. 

Distr. Malesia: Borneo. 

Ecol. In moss forest, also in more open sunny 
places (landslides), not rare locally, 1220-2135 
(-2745) m. Fl. Jan. -Dec. 

228. Rhododendron rhodopus Sleum. Reinwardtia 
5 (1960) 199. 

Shrub, apparently terrestrial. Branchlets terete, 
4-5 mm 0. laxly substellate-lepidote at the upper, 
glabrous and pruinose at the lower internodes 
(13-15 cm). Leaves in — 5-merous pseudowhorls, 
some additional reduced ones on the internode, 
lanceolate, subinequilateral, apex rather long-acu- 
minate, acute, base broadly cuneate, coriaceous, 
glabrous above with age, laxly lepidote beneath 
(scales small, flat, variously, mostly shortly dented 
or lacerate at the marginal zone, minute centre 
hardly impressed), 9-l3(-l7) by 2-3.5(-5) cm, 
midrib above dilated and slightly raised at base, 
narrowed and impressed upwards, beneath thick 
and obtusely prominent below, nerves c. 8 pairs, 
at an acute angle, high-ascending, indistinctly 
anastomosing, slightly depressed above, not much 
raised beneath, veins subobscure; petiole olivace- 
ous, laxly lepidote, subterctc, 1.5-2 cm by 2(-3) 
mm. Umbels 8-9-flowcred. Outer pcrulae broadly 
ovate-oblong, densely and very shortly sub- 
strigosc-hirsutulous outside, glabrous inside, c. 
4 by 2 cm. Bracteoles linear, laxly pubescent 
towards the apex, c. 2 cm by I mm. Pedicels laxly 
lepidote, not hairy, 1 .5-2 cm by c. I mm at anthcsis. 
Calyx disk-shaped, wavy, hardly lobed, lepidote 
dorsally, c. 4 mm 0. Corolla tubular, lobed in 
tnc upper third, white, scented, 6.5 7 cm long in 
all, glabrous outside or here and there with a 
solitary (caducous) scale, subdenscly pubescent at 
the tube inside, tube straight, 4 5 cm, at base 
slightly 5-saccatc, r. 5 mm 0, dilated to r. 8 mm 



at apex, lobes spreading, suborbicular, 2-2.5 by 
1.5-2 cm. Stamens 10, subequal, reaching halfway 
the corolla lobes; filaments linear and subvillous 
in the lower -/j, filiform and glabrous for the rest; 
anthers oblong, 4-5 by c. 1.3 mm. Disk glabrous. 
Ovary cylindric, yellowish tomentose, minutely 
lepidote, c. 10 by 2.5 mm, gradually narrowed to 
the style, the latter nearly as long as the corolla 
tube, yellowish tomentose in the lower y^, elepi- 
dote, glabrous below the capitate stigma. Sub- 
mature capsule thick-cylindric, attenuate at the 
base, mostly curved, shortly yellowish villous and 
minutely lepidote, 5-7 cm by 7-8 mm, crowned 
by the villous style. 

Distr. Malesia: Central Celebes (Latimodjong 
Range: Pokapindjang; Luwuk), in Latimodjong 
at 2800-3000 m. Twice found. Fl. June, //•. 
Sept. 

229. Rhododendron baenitzianum Laut. in K. 
Sch. & Laut. Nachtr. (1905) 337; Schltr. Bot. 
Jahrb. 55 (1918) 156; Sleum. Reinwardtia 5 
(1960) 200; Blumea 12(1963) \12. — R. maboroen- 
se Schltr, Bot. Jahrb. 55 (1918) 156. 

Terrestrial shrub, 1.5-2 m. Branchlets brown, 
terete, 2-4 mm 0, tips densely red-brown sub- 
stellately scaly, lower parts glabrescent; internode 
3-10 cm. Leaves opposite or in 3-5-merous 
pseudowhorls at the upper 1-2 nodes, erecto- 
patent, subsessile or shortly petioled, oblong to 
lanceolate or elliptic-lanceolate, sometimes in 
part ovate-lanceolate, apex long (3^ cm) sub- 
caudate-acuminate, acute, base broadly obtusely 
or subacutely attenuate, rarely rounded, :; in- 
equilateral, subcoriaceous, glabrous above at 
maturity, laxly lepidote beneath (scales small, 
flat, irregularly and not too deeply lobed or dented 
at the marginal zone, centre small, faintly or not 
impressed), glabrescent finally beneath too 
leaving minute, dark spots, edge h flat, 12-17 
(-25) by (2.5-)3.5-6.5(-8.5) cm, midrib narrow 
and impressed above, broader and bluntly 
prominent beneath, nerves 8-IO(-12) pairs, as- 
cending, curved-anastomosing before the edge, 
slightly impressed or raised above, distinctly 
prominent beneath, reticulation whether or not 
finely raised on both faces; petiole semiterete, 
grooved above, lepidote, (3-)4-7 by 1.5-2 mm. 
Umbels (3-)4-5(-7)-flowered. Outer pcrulae ovate 
to obovate, cuspidate or shortly acutely acuminate, 
laxly caducously lepidote externally and along the 
margin, moreover very shortly appresscd- 
pubescent in the middle or towards the apex, 
inner ones subspathulate, acuminate, dorsally 
laxly lepidote, but not hairy, up to 2.5 by I cm. 
Bracteoles filiform to linear-subspathulate, sub- 
glabrous, up to 2 cm. Pedicels laxly lepidote 
especially at base and apex, • glabrescent, 
epilose. (2.5-)3.5-4 cm by 1.5 mm. Calyx sub- 
oblique, rim-like, r. 4 mmo. Corolla funncl-shupcd, 
7.5_9(-l()) cm long in all, tube yellow, orange at 
lobes, laxly lepidote outside, papillose-pubescent 
at the tube inside, tube 4-6 cm. at base 5-pouched, 
6-8 mm (pouches including or covering the 
calyx), c. 2 cm at apex, lobes spreading, broad- 



628 



Flora Malesiana 



[ser. I, vol. 6* 



obovate to suborbicular, 3 (-4) by 2.5-3 cm. 
Stamens 10, subequal, reaching halfway the 
corolla lobes; filaments linear, short-patent- 
hairy in the lower half, glabrous above; anthers 
oblong, 4 by 1.2 mm. Disk pubescent at the upper 
margin, glabrous below. Ovary subcylindric, 
shortly and densely covered with subappressed 
forward-directed hairs and with scales, 10-12 by 
2-2.5 mm, continuous with the style, the latter 
equalling the stamens at full anthesis, densely set 
with subpatent hairs and with scales to near the 
top; stigma thick-capitate, 5-lobed. Capsule 
elongate-fusiform, somewhat curved, laxly hairy 
and lepidote, 15-20 cm by 4-5 mm. 

Distr. Malesia: E. New Guinea (Torricelli 
Mts; Maboro Range: Waria R. region). 

Ecol. Along rivulets or in rocky places, 
215-1200 m. Fl. April, June, Aug., /r. Aug. 

Vern. Tindili, Orne: Wantipi. 

230. Rhododendron englerianum Koord. Nova 
Guinea 8 (1909) 186; ibid. 8 (1912) 877, t. 152; 
J.J.S. ibid. 12 (1914) 140, nota sub R. angulatum; 
Kaneh. & Hatus. Bot. Mag. Tokyo 56 (1942) 
478; Sleum. Reinwardtia 5 (1960) 199. — R. 
lowii {non Hook./.) F.v.M. Trans. R. Soc. Vict. 
1 (1889) 22. — R. megalostigma F.v.M. I.e., 
nom. event. 

Shrub, 1-1.2 m. Branchlets terete, 5-7 mm 0, 
lepidote at the very tips, early glabrescent; 
internodes 4-10 cm. Leaves in 3-4 merous pseudo- 
whorls at the upper 2 (-3) nodes, large, subsessile, 
oblong or subovate-oblong, apex shortly to 
rather long-acuminate, acute or subacute, broadly 
attenuate towards the base (very base truncate- 
rounded or a little cordate), coriaceous, practi- 
cally glabrous above at full age, rather laxly 
lepidote beneath (scales relatively small, marginal 
zone irregularly and rather shallowly lobed or 
dented, centre but slightly impressed), edge short- 
revolute, 12-20(-25) by (4-)5-8.5(-9) cm, midrib 
a continuation of the petiole, narrow and marked- 
ly impressed above, much prominent on the 
underside below, nerves 8-10(-14) pairs, with 
additional shorter ones, irregular and ascending, 
inarching before the edge, slightly raised above, 
more distinctly so beneath, reticulation lax, but 
slightly raised on both sides; petiole grooved above, 
4-7 (rarely up to 10) by 3-4 mm. Umbels (4-) 
6-9-flowered. Outer perulae ovate-lanceolate, long- 
acute, inner ones broad-ovate-acuminate and 
subacute to oblong-spathulate, all sericeous and 
lepidote dorsally, glabrous internally, up to 5.5 
by 1.5 cm. Bracteoles linear-filiform, lepidote and 
puberulous, up to 3 cm by c. 1 mm. Pedicels 
erect, laxly lepidote, not hairy at all, (6-)7-ll 
cm, c. 1.5 mm 0. Calyx suboblique, patellar, very 
shortly obtusely 5-lobed, c. 5 mm 0, lepidote 
dorsally. Corolla funnel-shaped, orange or red- 
dish to cerise-pink, flushed yellow outside, c. 8 
cm long in all, laxly stellate-lepidote outside, 
laxly hairy at the tube inside, tube 2.5-3 cm long, 
base markedly 5-pouched, 5-6(-8) mm 0, gradu- 
ally extended to 1.5-2 cm at apex, lobes obovate, 
much spreading, 5-6 by c. 3 cm. Stamens 10, 



reaching halfway the corolla lobes; filaments 
linear and densely hairy in the lower, more 
filiform and glabrous in the upper half; anthers 
linear-oblong, slightly curved, 6-7 by 1 mm. 
Disk much prominent, velutinous in the upper 
half, glabrous below. Ovary subcylindric, greyish 
velutinous, with numerous minute rusty scales 
between the hairs, 9-10 by nearly 3 mm, passing 
gradually into the style which ± equals the sta- 
mens and is covered in its lower '/2-% with short 
hairs and well visible scales; stigma thick-capi- 
tate, 5-lobed. Capsule elongate-cylindric, laxly 
hairy and lepidote, 6-10 cm by c. 5 mm; pedicel 
2 mm 0. 

Distr. Malesia: New Guinea. 

Ecol. Epiphytic in Agathis forest, or terrestrial 
in rain-forest, 400-600 m, also reported from 
rocks near river, 1220-2745 m. Fl. March, April, 
July. 

Vern. Tiriz, Movoi: Ueli. 

231. Rhododendron wentianum Koord. Nova 
Guinea 8 (1909) 188; J.J.S. ibid. 12 (1917) 508; 
ibid. 12 (1918) t. 199, b-f, 203, a; Schltr, Bot. 
Jahrb. 55 (1917) 143; Sleum. Reinwardtia 5 
(1960) 200; Blumea 11 (1961) 127. — R. calli- 
chilioides Wernh. Trans. Linn. Soc. II, Bot. 9 
(1916) 95, incl. var. minor Wernh. I.e. 96. — 
R. wollastonii Wernh. I.e. 99. 

Epiphytic shrub, 0.5-1.5 m. Branchlets terete, 
1.5-3 (-4) mm 0, upper internode laxly to densely 
covered with flat scales; internodes 2-10(-20) cm. 
Leaves mostly 3-4 in a pseudowhorl, occasionally 
2 opposite, at the upper 1-2 nodes, ovate to 
lanceolate-ovate, subsessile, apex gradually and 
± elongately acuminate to subcaudate, acute or 
nearly so, base rounded to subcordate, rarely very 
broadly attenuate, subcoriaceous to coriaceous, 
glabrous above at full age, laxly lepidote beneath 
(scales small, thin, marginal zone irregularly dented 
or incised, centre slightly immersed), margin 
slightly or not revolute, (7-)8-13(-16) by (2.5-) 
3.5-6.3 cm, midrib slightly impressed above, very 
prominent beneath, nerves (8-)9-ll pairs, upper 
ones spreading and ± straight below, curved 
towards the edge and inarching, but little promi- 
nent on both faces, reticulation rather dense, 
faintly visible on both sides, or only above so; 
petiole densely lepidote initially, grooved above, 
1.5-6 (rarely up to 10) by 1-1.5 mm. Umbels 
(3-)4— 6-flowered. Perulae oblong-ovate, acutely 
acuminate, sericeous and laxly lepidote dorsally, 
up to 2.5 by c. 1 cm. Bracteoles subulate, up to 
2 cm. Pedicels slender to rather stoutish, densely 
to laxly lepidote, not hairy, (2.5-)3-4.2 cm. 
Calyx irregularly 5-lobed or -dented, wavy, 
densely lepidote, c. 4 mm 0. Corolla funnelform- 
campanulate in outline, tubular below, rose-red, 
tube suff"used with light green or yellowish white, 
(3-)4— 4.8 cm long in all, rather sparsely lepidote 
outside, laxly hairy to glabrous and not rarely 
yellowish inside, tube 1-1.5 cm, bluntly 5-angular, 
(3-)4-6 mm 0, manifestly 5-saccate at the base, 
slightly dilated upwards, lobes ± erecto-patent, 
somewhat recurved at the apex, broadly obovate, 



Jan. 1966] 



Ericaceae (Sleumer) 



629 



(2-)3-3.4 by (1.3-)2.5-3.2 cm. Stamens 10, sub- 
equal, manifestly exserted from the throat; fila- 
ments linear, densely patent-pubescent in the lower 
half, glabrous distally; anthers oblong, 2.5-3 mm. 
Disk glabrous below, pubescent in the upper half. 
Ovary conical, obtusely 5-angular, densely lepidote 
and velutinous. 4-6 by 1.5-2 mm, gradually 
tapering to the style, the latter thick, pubescent 
and lepidote in about the lower half or third, 
glabrous for the rest; stigma thick-capitate, lobes 
manifest. Capsule slenderly fusiform, laxly 
lepidote and hairy, 6.5-9 by 0.4-0.5 cm, on 
slender pedicel 3-4 cm. 

Distr. Malesia: New Guinea (Main Range, 
from Mt Carstensz to the Star Mts, and the 
Upper Sepik R. region). 

Ecol. In forest, 600-1500 m, locally not rare. 
Fl. Jan. -Aug., fr. July. 

232. Rhododendron cuspidellum Slelm. Rein- 
wardtia 5 (1960) 200. 

Terrestrial shrub, c. 1.5 m. Branchlets rather 
robust, terete or sometimes obtusangular, laxly 
to subdensely substellate-lepidote at the upper 
internodes, glabrous lower down; internodes 
8-17 cm. Leaves in 4.5(-7)-merous pseudowhorls 
at the upper 2 nodes, lanceolate to subovate- or 
oblong-lanceolate, apex rather long (subcaudate-) 
acuminate, the acute acumen not rarely somewhat 
curved, — broadly attenuate towards the base 
(very base subtruncate-rounded to subcordate), 
coriaceous, rather stiff, minutely rugose above and 
glabrous at full age, laxly lepidote beneath (scales 
small, irregularly substellate-lobed or -dented in 
the marginal zone, darker centre slightly impressed, 
flat, (9-)10-15 by (2.5-)3-5 cm, midrib dilated 
for the lowest 1-2 cm, becoming very narrow up- 
wards, impressed above, as thick as the petiole, 
blunt and much prominent beneath, nerves 8-10 
pairs, spreading, — straight below, rather indis- 
stinctly curved-anastomosing before the edge, 
faintly raised on both faces, or not rarely sub- 
inconspicuous especially beneath; petiole a little 
flattened, shallowly grooved above, lepidote, 5-10 
by 2-3 mm. Umbels 9-12-flowered. Flower buds 
suborbicular-ovoid, 3.5 by 2.5 cm. Outher perulae 
ovale, apex very shortly subulate, the subse- 
quent inner ones larger, ovate, nearly blunt, 
innermost ones spathulate, all laxly substellate- 
lepidote and very shortly pubescent dorsally and 
along the margin, up to 3.5 by 2.3 cm. Bracteoles 
linear to linear-subspathulate, short-pubescent, 
up to 3 cm by 1-4 mm. Pedicels laxly lepidote, 
epilose, (2.5-)3.5-6 cm by c. I mm. Calyx disk- 
like, wavy and obscurely lobed, densely lepidote 
dorsally, c. 3 mm 0. Corolla tubular below, much 
expanded at the limb, orange or orange-yellow, 
(3.5-)4— 4.5 cm long in all, subdensely or more 
laxly substcllatc-lcpidotc outside, sparsely or 
very sparsely pubescent in the lower part of the 
tube inside, tube 1.7 2.1 cm, straight, 5-angular, 
at base much 5-saccalc, c. 5 mm 0, widened 
upwards to 1.2-1.5 cm at apex, lobes broadly 
obovatc-spathulatc or suborbicular, 2.3-2.7 by 
1.5-2.2 cm. Stamens 10. subcqual, slightly exserted 



from the throat, c. 2.5 cm; filaments linear and 
densely patently hairy in the lower, more filiform 
and glabrous in the upper half; anthers oblong, 
c. 4 by 1 mm. Disk white-sericeous. Ovary sub- 
cylindric-conical, white-velutinous and lepidote, 
5-6 by c. 2.5 mm, rather abrupt with the style, 
the latter robust, nearly equalling the stamens, 
hairy and manifestly lepidote in the lower '73-^3; 
stigma capitate, deeply 5-lobed. 

Distr. Malesia: New Guinea (Wissel Lakes 
region). 

Ecol. Secondary forest, locally common, 1900 
m. Fl. March. 

Vern. Warade, Kapauku. 

233. Rhododendron commutatum Sleum. Reinward- 
tia 5 (1960) 201. — R. gracile {nan Low ex 
LiNDL.) Becc. Malesia 1 (1878) 203. 

Epiphytic or terrestrial shrub growing with 
dense-packed, slender stems, not rarely subarbor- 
escent, 1.5-3 m. Branchlets terete, 2-4 mm 0, 
densely set with substellate rufous scales at the 
upper internodes, early glabrescent in the lower 
ones; internodes (3-)6-l 2 cm. Leaves opposite or in 
3 (-5 )-merous pseudowhorls at the upper 2-3 nodes, 
lanceolate or oblong-lanceolate, apex — elongate, 
sometimes rather abruptly acuminate, subacute, 
base broadly and acutely attenuate to the petiole 
(never rounded), subinequilateral, thin-coriaceous 
lepidote above for a rather long time, finally 
glabrescent, minutely rugose by inflated epidermis 
cells and — persistently laxly to subdensely 
lepidote beneath (scales minute, the marginal zone 
substellate-lobed or -incised, centres mall and 
slightly impressed), edge flat, very sligthly revolute, 
(8-)10-l5(-l7) by (2-)3-5(-5.5) cm, midrib 
very narrow and markedly depressed above, as 
thick as the petiole, blunt and much prominent 
beneath, often more densely lepidote than the 
intervenium, nerves 10-14 pairs, spreading at 
almost 90 \ straight below, curved and obscurely 
anastomosing before the edge, with additional, 
less distinct nerves, :: parallel, faintly sunk above, 
minutely raised or subinconspicuous beneath, 
reticulation rather dense, visible above only; 
petiole flattened and grooved dorsally, densely 
lepidote, 1-1.5 cm by 1.5-2 mm. Umbels (3-)4-5- 
flowered. Outer perulae narrow-ovate-acuminate, 
extended at the apex into a fine sharp point, 
lepidote along the dorsal middle line and in the 
upper part, glabrous at the margin, inner ones 
much narrower, nearly linear, up to 2 by 0.7 cm. 
Bracteoles filiform, laxly lepidote, c. 1.5 cm. 
Pedicels laxly substellate-lepidote, not hairy, 
l-1.5(-2) cm by c. 1 mm. Calyx subobliquely 
disk-shaped, very shortly and bluntly 5-lobcd, 
densely lepidote dorsally, c. 4 mm 0. Corolla funnel- 
shaped, bright rose or red with magenta flushed 
throat, 5-6(-6.5) cm long in all. lobcd to about 
the upper third, laxly minutely subslcllatc-lcpidotc 
outside, laxly short-pubescent in the lower halt 
of the tube inside, lube 3 4 cm. base 5-saccatc, 
4--5 mm o, gradually dilated to r. 1.5 cm at apex, 
lobes ereclo-patcnt, broadly obovatc-spathulate or 
suborbicular. 2-2.5(-2.7) by (1.5-)l.8-2.4 cm. 



630 



Flora Malesiana 



[ser. I, vol. 6* 



Stamens 10, unequal, the longest reaching halfway 
the corolla lobes; filaments linear, subdensely 
papillose-puberulous below, papillose in the 
middle, glabrous in the upper third; anthers 
oblong, 3-3.5 by 1 mm. Disk glabrous below, 
short-pubescent on the upper margin. Ovary sub- 
cylindric, greyish velutinous and densely lepidote 
(scales ± covered by the forward directed hairs), 
rather abrupt with the style, 7-8 by c. 2 mm; 
style thickish, densely subpatent-hairy to nearly 
the top, less densely lepidote, nearly as long as 
the corolla; stigma thick-capitate. Capsula narrow- 
cylindric, curved, laxly hairy and densely lepidote, 
c. 8 cm by 4 mm. 

Distr. Malesia: Borneo (Sarawak). 

Ecol. In swampy jungle or white sand podsols 
at low elevations. Fl. Jan. -Dec. 

Vern. Jemalan, Iban. 

234. Rhododendron subcordatum Becc. Malesia 
1 (1878) 203; Merr. En. Born. (1921) 463; 
Sleum. Reinwardtia 5 (1960) 202. 

Terrestrial shrub. Branchlets terete, 2-3 mm 0, 
the upper internodes laxly set with flattish scales, 
no hairs; internodes 3-7 cm. Leaves opposite or 
in 3(-4)-merous pseudo whorls at the upper 1-2 
nodes, oblong- to lanceolate-ovate, apex mostly 
longish, rarely more shortly and rather abruptly 
acuminate, acute, base rounded to slightly cordate, 
coriaceous, glabrous above at full age, laxly and 
± persistently lepidote beneath (scales very small, 
narrow marginal zone irregularly substellate-lobed 
or -dented, early dissolute, dark centre minute, 
markedly impressed), edge very slightly or not 
revolute, 9-17 by (3.5-)4-6 cm, midrib broadened 
at the very base, otherwise narrowly impressed 
above, as stout as the petiole and obtusely promi- 
nent in the lower part, gradually narrowed towards 
the apex of the lamina beneath, nerves 8-10(-12) 
much spreading pairs, straight below, curved-in- 
arching before the edge, faintly sunk above, 
slightly prominent or sometimes subinconspi- 
cuous beneath, reticulation rather obscure on 
both sides; petiole subterete, not grooved above, 
rugose, lepidote initially, finally glabrous, 1.5-2.5 
cm by 2(-3) mm. Umbels 4-6-flowered. Bracteoles 
subfiliform, laxly lepidote. Pedicels ± densely 
very shortly pubescent and laxly lepidote, 2-2.8 
cm by 0.7 mm. Calyx oblique, disk-shaped, very 
shortly lobed or wavy, c. 3 mm 0, lepidote outside. 
Corolla narrow- funnel-shaped, red, 5-5. 5 (-6) cm 
long in all, laxly lepidote outside, subdensely 
short-pubescent at the tube inside, tube 3-3.5 cm, 
base 5-saccate, 4-5 mm 0, gradually widened 
upwards to 1-1.5 cm at apex, lobes expanded, 
obovate-oblong, 2-2.5 by 1.2-1.5 cm. Stamens 10, 
subequal, attaining ± halfway the corolla lobes; 
filaments linear and densely subpatent-hairy in 
the lower half, filiform and glabrous for the rest; 
anthers oblong, 3-4 by 1 mm, base biunt. Disk 
densely pubescent at the upper margin. Ovary 
subcylindric-conical, velutinous (the short hairs 
covering minute scales), continuous with the 
style, the latter nearly 4 cm long in full anthesis, 
densely covered with short hairs and scales to 



nearly the top; stigma thick-capitate, 2.5 mm 0. 

Distr. Malesia: Borneo (Sarawak: Prov. Batan 

Lupar, Mt Tian Laju), c. 1000 m. Fl. April, June. 

235. Rhododendron vanvuurenii J.J.S. Bull. Jard. 
Bot. Btzg 111, 1 (1920) 399, t. 48; H. J. Lam, 
Blumea 5 (1945) 577; Sleum. Reinwardtia 5 
(1960) 198. — R. kjellhergii J.J.S. in Fedde, 
Rep. 68 (1937) 202. 

Shrub or treelet, terrestrial, often with rather 
long and flexile branches, 1.2-4 m. Branchlets 
reddish, terete, 2-4(-5) mm 0, covered with a dense, 
scurfy, detersile tomentum of deeply stellate and 
subdendroid, rufescent scales at the upper inter- 
nodes, early glabrescent below; internodes (3-) 
5-10(-15) cm. Leaves mostly somewhat spaced, 
5-9 in the uppermost part of the upper l-2(-3) 
internodes, rarely more approximate in a pseudo- 
whorl at the nodes, oblong to lanceolate, rarely 
elliptic- or ovate-lanceolate, apex short-acumi- 
nate, subacute, base ± broadly attenuate, subacute 
or mostly blunt, subinequilateral, thin-coriaceous, 
densely stellate-lepidote on both faces, glabrescent 
above with age, underneath the detersile, loose 
tomentum consisting of rufous longer persistent 
scales (these deeply and narrowly stellate-incised 
in the marginal zone, centre depressed and 
subdendroid-extended downwards, very slightly 
impressed), edge very slightly or not revolute, 
(5-)6.5-12(-14) by (1.5-)2.5-3.7 (rarely up to 5) 
cm, midrib narrowly impressed above, broader 
and very prominent beneath, nerves (6-)7-9 pairs, 
somewhat irregular, spreading, rather straight 
below, curved-anastomosing before the edge, 
mostly rather faintly raised, reticulation 0; petiole 
subterete, rather slender, ± densely lepidote, 
(4-)6-10 by 1-1.5 mm. Umbels (4-)5-10-flowered. 
Outer perulae ovate, more inner ones ovate- 
elliptic to broad-oblong, innermost ones oblong- 
spathulate, all densely very shortly appressed-hairy 
and lepidote outside, laxly hairy inside, up to 3 
by 1.6 cm. Bracteoles filiform to linear-spathulate, 
laxly hairy, up to 2.2 cm. Pedicels rather stoutish, 
mostly densely, rarely more laxly subpatent- 
pubescent and caducously lepidote (especially at 
both ends), occasionally only laxly to subdensely 
lepidote and epilose, (0.9-)l. 3-2.5 cm. Calyx 
oblique, shortly obtusely 5-lobed or -dented, c. 
5 mm 0, densely hairy and lepidote. Corolla 
funnel-shaped, (3.7-)4.5-5.5 (sometimes up to 6) 
cm long in all, orange to yellow, also occasionally 
white or pink, laxly minutely stellate-lepidote 
outside, pubescent at the tube inside, tube obtusely 
5-angular, 2-3 cm, base manifestly 5-saccate, 
5-6 mm 0, gradually dilated to 1-1.3 cm at 
apex, lobes broadly obovate, 1.5-2.5 (rarely up to 
3) by 1-2 cm. Stamens 10, subequal, reaching ± 
halfway the corolla lobes; filaments linear and 
hairy below, filiform and glabrous in the upper 
half; anthers oblong, (3-)4-5 mm, base obtuse. 
Disk densely pubescent at the upper margin only. 
Ovary subcylindric, tapering to the style, densely 
yellowish brownish velutinous and lepidote, 
(5-)7-9 by 2-2.5 mm; style slender, i equalling 
the stamens, densely pubescent and lepidote below, 



Jan. 1966] 



Ericaceae (Sleumer) 



631 



less so upwards, glabrous for the rest; stigma thick- 
capitate. Capsule subfusiform-cylindric,subdensely 
hairy and lepidote, 5.5-7 by 0.6-0.7 cm. 

Distr. Malesia: Central Celebes. 

Ecol. In light dry forest, grass and fern scrub 
vegetation, on rocks or bare slopes, locally 
common and gregarious, (600-) 800- 1600 m. Fl. 
June, July. 

Note. The showy flowers vary in colour and 
are used for decoration by the people. 

236. Rhododendron leptobrachion Sleum. Rein- 
wardtia 5 (1960) 203. 

Shrub or treelet, up to 3 m. Branchlets terete, 
1.5-3 mm o, very tips laxly lepidote, glabrous 
otherwise, pruinose in dry specimens; internodes 
(3.5-)6-12 cm. Leaves in — 5-merous pseudo- 
whorls at the upper 1-2 nodes, l(-2) additional 
on the internode, lanceolote or suboblong- 
lanceolate, apex shortly subabruptly acuminate, 
subacute, base acutely attenuate to the petiole, 
subcoriaceous, — densely lepidote at both faces 
when young, glabrescent and minutely blackish 
punctulate above with age, more persistently and 
laxly lepidote beneath (scales rather small, 
relatively wide marginal zone early dissolute, 
variously substellate-lobed or -dented, minute 
centre slightly impressed, blackish with age), 
margin faintly or not revolute, (5.5-)6.5-9 by 
1.8-2.8 cm, midrib very narrow and impressed 
above, much and obtusely prominent beneath, 
pruinose, nerves 6-8(-10) pairs, irregular, spread- 
ing, obscurely inarching, with additional shorter 
intermediate veins, all minutely sunk or rather 
obscure above, raised beneath, reticulation rather 
dense, subinconspicuous above, more distinct 
beneath; petiole compressed and grooved dorsally, 
(l-)l.2-1.8(-2) cm by 1.5-1.8 mm. Umbels 4-6 
(-lO)-flowered. Pedicels slender, laxly lepidote, 
epilose, 0.8-1.4 cm. Outer perulae ovate to ovate- 
oblong, inner ones spathulate, very densely 
sericeous-hirsutulous outside, glabrous (the laxly 
pubescent base excepted) inside, up to 2 by 1 cm. 
Bracteoles linear, densely hirsutulous, 1.5-2 cm by 
0.6 mm. Calyx obliquely disk-shaped, faintly 
5-lobed, densely lepidote dorsally, 3-4 mm o. 
Corolla tubular, scarlet to fiery red, 3-3.5(-4) cm 
long in all, laxly or very laxly lepidote especially 
towards the base outside, subdensely very shortly 
pubescent at the tube inside, tube subcylindric, 
straight, c. 2.5 cm, the base manifestly 5-saccate, 
c. 4 mm 0, dilated to 5-6 mm at apex, lobes 
broadly obovate or suborbicular, crecto-patent at 
anthesis, 1-1.3 by^).l-0.9{~\)cm. Stamens 10, une- 
qual, (he longest reaching : halfway the corolla lo- 
bes; filaments linear, densely subpatcntly pubescent 
in the lower y^, glabrous above; anthers obovate- 
oblong, 2 by I mm. Disk prominent, glabrous. 
Ovary subobovoid-cylindric, densely subpatcntly 
hairy and lepidote, 4-6 by 2-2.5 mm, tapering to 
the style, the latter rather slender, nearly as long 
as the stamens, subdensely to more laxly hairy 
in the lower half, sparsely or not lepidote; stigma 
capitate. Capsule obliquely sublusiform-cylindric, 
•lightly curved, subdensely hairy and lepidote. 



2.5-3 by 0.5-0.6 cm, on laxly lepidote pedicel 
2.2-3 cm by 1 mm. 

Distr. Malesia: Central Celebes. 

Ecol. In mossy forest. 2000-2800(-3000) m, 
locally common. Fl. May-June. 

237. Rhododendron schlechteri Laut. in K. Sch. & 
Laut. Nachtr. (1905) 338; Schltr, Bot. Jahrb. 
55 (1918) 158, f. 5; Sleum. Reinwardtia 5 (1960) 
120. 

Epiphytic shrub, few-branched, 1-1.5 m. 
Branchlets terete, glabrous; internodes 6-8(-10) 
cm. Leaves spiral or subverticillate, crowded 
apically, ± spreading, oblong-elliptic, apex 
broadly obtusely attenuate, base broadly attenuate 
to rounded, coriaceous, firm, glabrous above at 
maturity, rather densely lepidote beneath (scales 
small, irregularly stellate-dentate or -lacerate in 
the marginal zone, centre not or very slightly 
immersed, finally caducous, leaving blackish 
points beneath), flat, 13-17 by 6-8 cm, midrib 
nearly flat above, robust and strongly raised 
below, nerves 8-9 pairs, rather straight or slightly 
curved, obsoletely anastomosing, slightly raised 
on both faces, reticulation obscure; petiole 
subterete, 1.5-2 cm by 2-3 mm. Umbels ± 
12-flowered. Pedicels densely covered with rusty 
scales, which are deeply incised to or nearly to 
the small centre, otherwise laxly hairy imme- 
diately below the calyx, 1.2-2 cm by 2.5-3 mm. 
Calyx patelliform., ± oblique, shaiiowly 5-lobed 
and wavy, stellate-lepidote outside, glabrous inside, 
c. 6.5 mm 0. Corolla tubular below, funnelform 
distally, snow white, very fragrant, (I5-)I6(-17) 
cm long in all, tube cylindric, 10 — 1 2 cm long, base 
manifestly 5-saccate, c. 1-1.2 cm 0, gradually 
dilated to c. 1.4 cm in the middle, to 2-2.5 cm 
at apex, glabrous outside, rather densely hairy 
and laxly lepidote inside, lobes 5, obliquely and 
broadly obovate, rounded at the apex, ± spread- 
ing, margin wavy, c. 5-6(-7) by 3.5-4(-5) cm. 
Stamens 10, manifestly exserted; filaments linear, 
flat, c. 1.5 mm wide below, gradually attenuate 
and finally filiform distally, densely hairy (the 
hairs denser and m retrorse at the margin) 
glabrescent in the upper third. 11-13.5 cm; 
anthers elongate-oblong, dorsifix, versatile, 1.6- 
1.8 cm by 1.8 mm, cells minutely apiculate at the 
base. Disk lobes set with tufts of short hairs. 
Ovary oblong-cylindric, very densely clad with 
yellowish, i spreading hairs mixed with stellate 
scales, and covering them i completely, c. 2.5 
by 0.7-0.9 cm. gradually tapering to the very 
stout cylindrical style, (c. 12.5 cm by 3.5 mm), 
equally covered with yellowish or fulvous sub- 
patent hairs and stellate scales for the lower c. 
7 cm, becoming almost exclusively lepidote above, 
and entirely glabrous for 2 cm below the stigma, 
the latter broadly and shortly turbinate, coarsely 
5-crcnatc. 

Distr. Malesia: NE. New Guinea (Kani and 
Bismarck Mts). 

Ecol. Rain-forest epiphyte on high trees, 
1000-1700 m. //. Jan., June. 



632 



Flora Malesiana 



[ser. I, vol. 6* 



238. Rhododendron leucogigas Sleum. Blumea 12 
(1963) 102; Jahrb. Rhodod. Ges. Bremen (1963) 
11, //; text, f. 7-8; Schneider, Jahrb. Deutsch. 
Rhodod. Ges. Bremen (1965) f. 2^. 

Epiphytic lanky shrub, l-2(-3) m, erect, few- 
branched. Branchlets densely stellate-lepidote at 
the new shoots, early glabrescent; upper internodes 
(3-)6-12(-20) by (0.5-)0.8-1.2 cm. Leaves in 
(3-)5-6(-7)-merous pseudowhorls at the upper 
3-4 nodes, spreading, oblong or subelliptic- or 
subobovate-obiong, apex gradually attenuate to 
subacuminate, towards the base broadly attenuate 
to cuneate (base rounded-obtuse or often slightly 
cordate), coriaceous, young ones stellate-lepidote- 
tomentellous (scales of two different types, i.e. 
subdendroid much stellate-incised, rufous, early 
caducous ones, and flattish, minor and paler, 
less deeply dentate and more longer persistent ones, 
the impressions of the latter found on the mature 
leaves beneath), later ± glabrescent, flat, (18-) 
20-28 (in cultivation -36) by 6-10.5 cm, midrib 
broadened below, gradually narrower and becom- 
ing a little impressed distally above, as thick as 
the petiole and much prominent at the base, 
nerves (9-)10-l 1(-12) pairs, curved-ascending, 
inarching before the edge, slightly raised (their 
middle-line often faintly impressed) as is the lax 
network of the veins and veinlets above, much 
less prominent beneath; petiole somewhat flatten- 
ed, 7-1 8 (-20) by (3-)4-5 mm. Umbels (5-)6-8- 
flowered. Flower bud ovoid, of a dull carmine; 
outer perulae ovate, apiculate, more inner ones 
obovate to spathulate, covered with a tomentum of 
short, a little scabrid appressed hairs and scales 
outside, and by a tomentum of softer and longer 
hairs inside, 6.5-8 by 2-4 cm. Bracteoles linear, 
dilutely carmine, glabrous, c. 6 cm by 2-3 mm. 
Pedicels densely lepidote, c. 1 by 0.4-0.5 cm. 
Calyx rim-like, sometimes with irregular linear 
lobes up to 8 by 2 mm. Corolla 7-merous, tubular- 
funnelform, white suffused with pale carmine at 
angles of lobes with tube, and partly on the tube 
itself, subdensely stellate-lepidote outside, densely 
hairy at the tube inside, c. 14 cm long in all, 
strongly carnation-scented, tube straight, slightly 
dilated distally, 8-9 by 1.5-2 cm, lobes ± spread- 
ing, obovate, apex emarginate, edge crenulate, 
5-5.5 by 3-4 cm. Stamens 14; filaments hairy, 
exserted for c. 1 cm; anthers linear, base obtuse, 
c. 12 by 1.5 mm. Disk densely hairy. Ovary 
thick-cylindrical, c. 1.5 by 1 cm, tomentose. 
Style thick, c. 7 cm by 2 mm, tomentose in the 
lower, hairy and lepidote in the upper half, 
with a short glabrous zone below the stigma. 
Submature capsule obfusiform, densely covered 
with hairs and scales, 14—16 cm by 1.4-1.8 cm, 
continuous with the style, the latter 10-11 cm by 
2-3 mm, densely hairy and lepidote below, more 
laxly so upwards in the lower %, apparently 
exclusively lepidote for 5-6 mm, glabrous for the 
rest; stigma large, 7-lobed. 

Distr. Malesia: W. New Guinea (Cycloop 
Mts). 

Ecol. On tree trunks in riverine forest or in 
submossy Nothofagus forest at 1200-1350 m, 



not rare locally. Fl. apparently April-May, fr. 
June. 

Note. This species has the largest leaves in 
New Guinea, and, to conclude from the size of 
the developed ovary cq. immature fruit, probably 
bears the largest flowers in the genus, nearest to 
R. schlechteri Laut. 

It is now in cultivation from cuttings and the 
description and size of the floral parts are taken 
from the living plant. The flowers of the latter 
may be smaller than those of the wild plant. 

239. Rhododendron lompohense J.J.S. Bull. Jard. 
Bot. Btzg III, 1 (1920) 402, t. 50; Sleum. Reinward- 
tia 5 (1960) 204. 

Shrub. Branchlets terete, 2.5-4 mm 0, rather 
densely lepidote at the upper internode, elsewhere 
glabrous; internode 2.8-9 cm, with the scars of 
several rudimentary, very early caducous leaflets. 
Leaves in 4(-7)-merous pseudowhorls at the 
upper nodes, oblong-ovate, apex gradually 
acuminate, nearly acute, base ± rounded, 
rather stiffly coriaceous, initially densely lepidote 
on both faces, at maturity glabrescent above, ± 
persistently and subdensely lepidote beneath 
(scales flat, small, marginal zone irregularly 
incised, centre very slightly impressed), the 
margin faintly or not revolute, 7-10.5 by 2.8-4.4 
cm (the upper leaves in the same whorl not rarely 
smaller), midrib broadened below, — flat above, 
strongly raised beneath, nerves (8-)10-12(-14) 
spreading pairs, straight at the base, curved 
upwards and inarching distally, a little raised 
above, more distinctly so beneath, reticulation 
indistinct above, dense and minutely prominent 
beneath; petiole 6-12 by 1.5-2 mm. Umbels ± 
6-flowered. Perulae orbicular to broadly ovate- 
acuminate, lepidote-fimbriate, lepidote and very 
shortly pubescent near the base outside, finally 
glabrescent, remaining faintly verruculose, up to 
3 by 1.5 cm. Pedicels stoutish, subdensely lepidote, 
epilose, 1.2-1.6 cm. Calyx inconspicuous, disk- 
shaped, oblique, c. 3 mm 0. Corolla tubular 
below, more funnel-shaped upwards, white, 
glabrous outside, pubescent at the tube inside, 
tube subangular-cylindric, 2.6-3.1 cm, at base 
manifestly 5-gibbous, 4-5 mm 0, slightly dilated 
towards the apex, lobes obovate, erecto-patent, c. 
1.5 by 1.2-1.3 cm. Stamens 10, somewhat exserted 
from the throat; filaments linear, rather densely 
hairy in the lower 2/3, glabrous for the rest; 
anthers oblong, base obtuse, c. 3 by 1 mm. 
Disk glabrous below, set with short white hairs 
at the upper margin. Ovary cylindric-conical, 
5-angled, c. 9 by 2 mm, appressed-hairy (the 
hairs covering numerous scales), gradually 
developed into the style, the latter c. 2 cm long, 
densely hairy and laxly lepidote in the lower 
2/3, glabrous above; stigma large, capitate, 
5-lobed. 

Distr. Malesia: Central Celebes (Mt Lompoh), 
once found. Fl. Dec. 

240. Rhododendron buruense J.J.S. in Fedde, Rep. 
30 (1932) 168; Sleum. Reinwardtia 5 (1960) 



Jan. 1966] 



Ericaceae (Sleumer) 



633 



204. — R. lompohense J.J.S. var. grandifolium 
J.J.S. in Fedde, Rep. 30 (1932) 169. 

Terrestrial shrub, up to 3.5 m. Branchlets 
terete, 3-4 mm o, laxly appressed-lepidote at the 
upper internode which is — distinctly pruinose; 
internodes (3-)7-16 cm. Leaves in 4-5 (-6)- 
raerous, lax pseudo whorls at the upper 1-2 nodes, 
1-2 additional ones on the internode which has 
also scattered spaced scars of reduced, very 
early caducous leaflets, ovate-oblong to elliptic, 
apex short-acuminate, rather obtuse, base broadly 
attenuate, sometimes nearly rounded, coriaceous, 
glabrous above at full age, laxly ± persistently 
lepidote beneath (scales small, the early disappear- 
ing marginal zone variously lobed or dented, lea- 
ving dark spots, centre small, shallowly impressed), 
edge very slightly or not revolute, (7-) 12-1 6 by 
(4-)5-7.5 cm, midrib grooved above, obtusely 
prominent beneath, nerves 8-12 pairs, mostly 
straight below, curved-anastomosing before the 
edge, minutely impressed above, distinctly raised 
beneath, reticulation rather lax and but faintly 
prominent beneath; petiole subterete or slightly 
flattened, rugose, often pruinose, 1-2 cm by 
1.5-2 mm. Umbels 7-10-flowered. Pedicels ± 
laxly lepidote, laxly to very laxly pubescent, 
(2-)2.2-3.3 cm by c. 1 mm. Calyx obliquely 
disk-shaped, lepidote and with some short hairs 
dorsally, 4-5 mm 0. Corolla broad-funnel-shaped 
in outline, tubular below, white, 4-4.5(-5) cm 
long in all, with some very scattered scales, i.e. 
practically glabrous outside, short-pubescent at the 
tube inside, tube cylindric, straight, 2-2.6 cm, at 
base manifestly 5-saccate, 5-6 mm 0, slightly 
widened towards apex, lobes — expanded, broad- 
obovate to suborbicular, (1.2-)1.5-2.3 by (1.2-) 
1.5-2 cm. Stamens 10, unequal, the longest 
reaching halfway the corolla lobes; filaments 
linear and subdensely patent-hairy in the lower 
Vi to '/2. more filiform and glabrous above; 
anthers oblong, 3-4 by 1 mm, the very base 
slightly thickened, gland-like. Disk prominent, 
glabrous or short-hairy at the upper margin. 
Ovary subcylindric, densely clothed with subpatent 
short hairs which cover small scales, 6-7 by 
2.5-3 mm, rather abrupt with the style, the latter 
2: densely hairy and laxly lepidote to the lower 
2/3, glabrous for the rest; stigma capitate. 

Distr. Malesia: Moluccas (Buru). 

Ecol. On loamy mouldered limestone, 1290- 
1900 m. /-/. Feb., April. 

241. Rhododendron blocmbcrgenii Sleum. Rcin- 
wardlia 5 (I960) 204. 

Treelet, c. 3 m. Branchlets subangular and 
very sparsely apprcsscd-lcpidotc at the tips, 
otherwise terete, glabrous, not rarely pruinose, 
4-9 mm 0; internodes 5-20 cm. Leaves opposite 
or in 3-mcrous pseudo whorls at the upper I 2(-3) 
nodes, with l-2(-3) additional leaves (scattered 
on internode), broad-oblong, apex shortly and 
not rarely rather abruptly acuminate, subacute, 
base broadly attenuate, subinequilatcral, firmly 
coriaceous, sparsely lepidote or entirely glubrcsccnt 
above with age, subpcrsistently laxly to subdensely 



lepidote beneath (scales minute, flat, the rather 
broad, early dissolved marginal zone irregularly 
substellate-lobed or -dented, centre very small, 
hardly impressed), edge very shortly revolute in 
dry state, (9-)ll-19 by (3-)4-8 cm, midrib as 
wide as the petiole in the lower part, flat or shal- 
lowly impressed above, very thick and obtusely 
prominent beneath, nerves 9-12 pairs, high-ascend- 
ing at a rather acute angle, curved and anasto- 
mosing before the edge, with several intermediate 
smaller nerves, all ± parallel, faintly impressed 
or raised above, distinctly prominent beneath, 
reticulation slightly raised beneath only; petiole 
flattened, l-1.5(-2) cm by c. 3 mm, lepidote, 
lengthwise striate as is the midrib. Umbels ± 
10-flowered. Outer perulae ovate, blunt, 1.5-2 
by c. 1 cm, initially (especially at the base) 
sericeous dorsally, early glabrescent, manifestly 
rough to the touch, inner ones oblong to oblong- 
spathulate, very densely subsericeous-hirsutulous 
dorsally, up to 6 by 2.5 cm. Bracteoles linear- 
subspathulate, ± densely hairy, c. 2 cm by 1-1.5 
mm. Pedicels subdensely substellate-lepidote at 
anthesis, 1.2-1.7 cm by c. 1.5 mm. Calyx disk- 
like, very short, c. 3 mm 0. Corolla long-tubular 
below, expanded at the limb, white, fragrant, 
glabrous outside, densely shortly pubescent at the 
tube inside, (6.5-)7-7.5 cm long in all, tube 
4.5-5 cm, cylindric, straight, base ± 5-saccate, 
5-6 mm 0, hardly dilated to apex, lobes broadly 
obovate-spathulate or suborbicular, 2-2.5 by 
1.5-2 cm. Stamens 10, reaching ± halfway the 
corolla lobes; filaments linear and densely patent- 
hairy to the lower 2/3, glabrous and filiform above; 
anthers oblong, 6-7 by c. 1.5 mm, each cell 
protracted in a short basal appendix. Disk rather 
prominent, glabrous, Ovao' subcylindric, yellowish 
tomentose (scales not visible), 9-10 by 3-4 mm, 
continuous with the style, the latter thick, slightly 
exceeding the stamens, densely hairy below, 
more subdensely or laxly so and minutely lepidote 
in the upper part, glabrous for the upper 6-8 mm; 
stigma thick-capitate, c. 3 mm 0. Capsule thick- 
subfusiform-cylindric, laxly shortly hairy, some- 
what rugose, slightly curved, 3.5-4 by 0.7-0.8 cm; 
pedicel 3-3.5 cm by 2 mm. 

Distr. Malesia: W. and Central SW. Celebes 
(Mt Ngilalaki, E of Lake Lindu; Latimodjong 
Mts). 

Ecol. Summit vegetation, 2200-2335 m. Fl. 
July, fr. Aug. 

242. Rhododendron macgregoriac F-. v. M. J. Bot. 

29(1891) 177; J.J.S. Nova Guinea 18 (1936) 97; 
Slbum. Rcinwardtia 5 (1960) 205; Blumca II 
(1961) 127; ihiU. 12 (1963) 112; Jahrb. Rhodod. 
Gcs. Bremen (1963) f. 5, 20. 32. — R. Iuin.\emanni 
Warh. Bot. Jahrb. 16 (1892) 26; K. Sc n. & Laut. 
Fl. .Schut/gcb. (1901) 487; S( hi.tr, Bot. Jahrb. 
55 (1917) 141. f. I; Lanil-Pooli:, For. Res. 
Papua (1925) 130; Wiiiii & Iran( is, Proc. R. 
Soc. Ouccnsl. 39 (1928) 68; Diii.s, Bot. Jahrb. 
62 (1929) 486. R. vonrormcri Kookd. Nova 
Guinea 8 (1912) 879. t. 155; J.J.S. ibid. 12 (1914) 
143; GiHBS, Arfak (1917) MO.—R.lauterbacliianum 



634 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 39. Rhododendron macgregoriae F. v. M. Mt Nettoti, 1750 m, NW. New Guinea (van Royen & 

Sleumer 7415) (Sleumer, 1961). 



FoERSTER in Fedde, Rep. 13 (1914) 224; Stonor, 
Rhodod. Yearbook 6 (1951) 49, f. 49— i?. calceo- 
larioides Wernh. Trans. Linn. Soc. II, Bot. 9(1916) 
95.— i?. gorumense Schltr, Bot. Jahrb. 55 (1917) 
142.— Fig. 39. 

KEY TO THE VARIETIES 

1. Corolla ± densely hairy at the tube inside. 

1. var. macgregoriae 

1. Corolla glabrous inside. 

2. Filaments densely hairy below 2. var. mayrii 

2. Filaments very sparsely hairy below, or quite 

glabrous 3. var. glabrifilum 



1. var. macgregoriae. 

Shrub or treelet, 0.5-5 m, rarely tree up to 15 m, 
terrestrial, rarely epiphytic. Branchlets rather 
slender, terete, upper internodes sparsely lepidote, 
lower parts glabrous; internodes (2-)4-10 cm. 
Leaves mostly in 3-5(-7)-merous pseudowhorls, 
rarely subopposite at the upper 1-3 nodes, 
ovate-lanceolate to subovate- or sometimes 
elliptic-oblong, rarely ovate, apex ± shortly 
acuminate, subacute or more bluntish, base 
broadly and ± acutely attenuate to the petiole, 
rarely rounded (then petiole longer than 4 mm), 
subcoriaceous to rather coriaceous, glabrous above 



Jan. 1966] 



Ericaceae (Sleumer) 



635 



at maturity, subdensely lepidote beneath (scales 
brownish, flat, thin, marginal zone irregularly 
dentate or incised, centre small, shallowly or 
hardly impressed), — flat, (4-)5-8 (rarely up to 
15) by (1.7-)3.5(-6) cm, midrib a little impressed 
above, prominent beneath, nerves (6-)8-10 pairs, 
spreading, — straight and markedly inarched 
before the edge, slightly raised above to faintly 
impressed, somewhat prominent beneath, reticula- 
tion very slightly raised on both sides, rather 
dense and mostly well visible; petiole flattened, 
grooved above, lepidote, (3-)6-12 mm. Flower 
buds subglobose. Umbels (4-)8-15-flowered. 
Outer perulae ovate, inner ones obovate to 
spathulate, all dorsally laxly lepidote and minutely 
sericeous, ciliate, up to 2 by 1 cm. Bracteoles 
linear to filiform, subspathulate distally, very 
laxly hairy or glabrous, up to 1.5 cm. Pedicels 
slender to rather stoutish, sparsely lepidote, 
normally epilose, but sometimes laxly set with 
very fine, short hairs, (l-5.)2.5-3.5(-4.5, rarely up 
to 7) cm. Calyx oblique, patellar, sometimes 
rudimentary, lobes indistinct, margin — ciliate. 
Corolla tubular below, generally much (sub- 
horizontally) expanded at the lobes, light yellow 
to dark orange or yellow at the tube and/or centre, 
reddish orange at the lobes, variable in size, 
mostly scentless, (l-1.8-)2-2.8 cm long in all 
(in natural hybrids with other species of ser. 
Javanica tending towards R. macgregoriae, 
larger, up to 4 cm long with more erect corolla 
lobes), 5-lobed to Vj-'/s' laxly lepidote outside, — 
densely set with spreading hairs at the lube inside, 
tube straight, somewhat 5-angular, with 5 basal 
rounded-saccate pouches, c. 1 cm long, 3-5 mm 
at base, slightly or not dilated upwards, lobes 
broadly obovate-spathulate, laxly lepidote outside, 
glabrous inside, 1.2-1.8(-2) by 0.8-1. 3(-l. 6) cm. 
Stamens 10, unequal, almost as long as the corolla; 
filaments linear and densely clothed with sprea- 
ding hairs or subvillous in the lower, glabrous 
and narrower in the upper half; anthers oblong, 
2.5-3.2 by c. 1 mm. Disk with thick lobes, mostly 
glabrous, rarely shortly hairy in the upper half. 
Ovary elongate-conical, densely covered with 
subappressed hairs, which completely cover the 
scales, c. 4 by 1.8 mm, tapering to the style, the 
latter 1-1.5 cm, densely to laxly hairy and lepidote 
in its lower, glabrous in upper half; stigma thick- 
capitate, somewhat deepened in the middle, 
lobes distinct. Capsule cylindric, slightly curved, 
manifestly 5-ribbed, attenuate at apex and base, 
hairy and lepidote, 3.5-4.5 by 0.3-0.4 cm, on 
slender fruit-pedicel 4-5 cm; valves twisting. 
Seeds c. 4 mm including the long tail at both ends. 

Distr. Malesia: New Guinea. 

Ecol. Distributed over the whole of New 
Guinea, fairly common, locally abundant to such 
degree, that it can be recognized from a far 
distance when in flower, both in primary mossy 
forest, secondary brushes and moderately dry 
sunny situations, among trees on mountain slopes, 
in thin canopy shade (Quercus forest), also along 
creeks, on precipitous banks of streams, scattered 
over open (Imperaia-, maybe rcgrowth) grassland. 



not rarely on landslides and formerly cultivated 
land, invading artificial clearings (air strips), on 
sandy or often on peaty soil, also stony clay and 
limestone ridge, at (120-)50O-30O0(-3350, perhaps 
even more) m. Fl. Jan. -Dec. 

Vern. Gupou, Ueli: Movoi (Kunimaipa Valley), 
nektamuk, Dani, warade, Kapauku, gorogoro, 
Guale: Movoi, liriz, Ueli: Movoi (Upper Zevi 
Valley), nekaloa, Andjai, hongkempa, Sosomepari, 
hakessa, Asaro: Kefamo, kibimume, Chimbu: 
Masul, wumbamb, Wahgi: Minj, wambu, Hagen: 
Togoba, atikro, Kebar: Karoon, ron, Enga, 
xdipndan, Hattam, wamp, Mendi, favell, Wapi. 

Note. Reported to be poisonous to mules 
(Edie Creek). Natural hybrids, apparently in the 
first place with the equally common R. zoelleri, 
not rare locally. Such plants show the general 
habit and foliage of R. macgregoriae, but have 
larger flowers. 

2. var. mayril (J.J.S.) Sleum. Reinwardtia 5 (I960) 
206.— 7?. mavrii J.J.S. Nova Guinea 18 (1936) 
97, t. 20, f. 2. 

Corolla glabrous inside, light red, otherwise 
as in var. macgregoriae. 

Distr. Malesia: W. New Guinea (Arfak Mts: 
Mt Ditschi), at 1100 m, once found. 

Vern. Sangsaia. 

3. var. glabrifilum (J.J.S.) Sleum. Reinwardtia 5 
(1960) 206; Blumea 1 1 (1961) 127; ibid. 12 (1963) 
112.—/?. glabrifilum J.J.S. Nova Guinea 12 (1914) 
134, t. 31. — R. torricellense Schltr, Bot. Jahrb. 
55 (1917) 142. 

Filaments very sparsely hairy below or quite 
glabrous. Corolla glabrous inside (yellow or red), 
otherwise as in var. macgregoriae. 

Distr. Malesia: New Guinea. 

Ecol. Apparently with preference (or exclusi- 
vely?) epiphytic, 200-1675 m. 

Vern. Munako, Kamora. 

243. Rhododendron rosendahlii Sleum. Reinwardtia 

5 (1960) 207. 

Erect shrub, c. 1.5 m. Branchlets erect, laxly 
substellate-lepidote at the upper internodes 
(1.5-7 cm by 1.5-2 mm), glabrescent at the lower 
ones (3-10 cm by 2-3 mm). Leaves in 3-4(-5)- 
merous pseudowhorls at the upper 2-3 nodes, 
ovate to ovate-elliptic, apex longer or shorter 
and gradually acuminate or attenuate, :; bluntish, 
more rarely subacute, sometimes almost rounded 
and minutely emarginate, at base slightly but 
distinctly cordate, coriaceous, laxly punctate by 
the remains of scales above, laxly to subdensely 
and subpersistently lepidote beneath (scales small, 
variously subslcllatc-lobed or -dented at the early 
evanescent marginal zone, centre minute, much 
impressed), margin slightly rcvolute. (3-)4-6 
(-7.5) by (2-)2.5-3.5 cm. midrib narrow and 
immersed above, a continuation of the thick 
petiole and obtusely prominent in the lower 
part beneath, nerves 6-8 pairs, spreadmg, curved- 
anastomosing before the edge, slightly impressed 
above, faintly so beneath or almost inconspicuous. 



636 



Flora Malesiana 



[ser. I, vol. 6* 



reticulation ± obscure; petiole subterete, grooved 
above, 2-4(-5) by 1 .5-2 mm. Umbels 3-5-flowered. 
Outer perulae subcoriaceous, ovate, short-acumi- 
nate, laxly lepidote and minutely sericeous in the 
middle line dorsally, 5-6 by c. 3 mm, inner ones 
much thinner, obtuse, oblong to obovate-spathu- 
late, densely and very shortly sericeous, not 
lepidote, up to 2 by c. 0.7 cm. Bracteoles filiform 
below, subspathulate-dilated distally, pubescent, 
up to 1.5 cm by c. 1 mm. Pedicels densely sub- 
patently pubescent and very laxly or not lepidote, 
(1.5-)2-2.5(-3.5) cm by c. 0.6 mm. Calyx disk- 
shaped, very shortly obtusely 5-dented, shortly 
pubescent dorsally, c. 3 mm 0. Corolla tubular- 
funnel-shaped, slightly oblique, dark red, sparsely 
substellate-lepidote outside, glabrous inside, 3-3.5 
cm long in all, tube (2-)2.5 cm, at base manifestly 
5-saccate, c. 4 mm 0, dilated to 6-8 mm at apex, 
lobes broadly spathulate or suborbicular, 1-1.2 
by 0.8-1 cm. Stamens 10, unequal, the longest 
reaching i halfway the corolla lobes; filaments 
linear, glabrous. Disk tomentellous especially at 
the upper margin. Ovary subcylindric, continuous 
with the style, tomentose, the spreading hairs 
hiding minute scales, 6-7 by 2.5 mm; style thickish, 
tomentose in the lower, less densely hairy in the 
upper part, glabrous for the ultimate 2-3 mm; 
stigma thick-capitate. 

Distr. Malesia: New Guinea (Wissel Lakes 
region). 

Ecol. Terrestrial, in peaty swamp, 1380-1800 
m. Fl. Sept., Jan. 

Vern. Wagadei, Kapauku. 

244. Rhododendron comparabUe Sleum. Reinward- 
tia 5 (1960) 208. 

Erect shrub, up to 2 m. Branchlets (2-)3-4 mm0, 
subterete, laxly and caducously substellate-lepidote 
at the upper internodes, early glabrescent below, 
not rarely pruinose; internodes 2-8(-ll) cm. 
Leaves in 4— 5-merous pseudowhorls at the upper 
2-3 (-5) nodes, elliptic to subobovate-elliptic, apex 
very broadly attenuate or very shortly subabruptly 
acuminate, obtuse, sometimes rounded, base 
broadly attenuate to the petiole, coriaceous, 
glabrous above at full age, laxly and subpersistently 
lepidote beneath (scales minute, the marginal 
zone irregularly substellate-lobed or -dented, 
centre slightly impressed), edge slightly but 
distinctly revolute, (3.5-)4.5-8(-9) by 2.5-4(-5) 
cm, midrib narrowly impressed above, thick and 
obtusely prominent beneath, nerves (5-)6-9 pairs, 
spreading, sometimes irregular, or in part divided 
before the edge, mostly subparallel to each other 
and distinctly curved-anastomosing, slightly im- 
pressed above, somewhat raised beneath, reticula- 
tion faintly impressed above in old leaves, not 
rarely subinconspicuous; petiole subterete, grooved 
above, (8-) 10- 18 (-23) by 1-2 mm. Umbels (8-) 
9-13-flowered. Outer perulae ovate, subacumi- 
nate, or orbicular, lepidote at the median or to the 
apex dorsally, inner ones obovate to spathulate, 
early glabrescent dorsally, or glabrous from the 
beginning, all with marginal scales, up to 2 by 
1.5 cm. Bracteoles linear-spathulate, glabrous, 



c. 1.5 cm by 0.5-1.5 mm .Pedicels rather densely 
substellate-lepidote, 2-2.5(-3) cm by c. 1 mm. 
Calyx obliquely disk-shaped, very shortly and 
bluntly 5-lobed, lepidote dorsally, c. 3 mm 0. 
Corolla funnel-shaped, white, fragrant, (4-)4.5 
(-4.8) cm long in all (5 cm at limb when fresh), 
laxly to subdensely set with minute substellate 
scales outside, short-pubescent at tube inside, 
tube straight, subcylindric, c. 2.5 cm, at base 
markedly 5-saccate, 5-6 mm 0, gradually dilated 
to 8-12 mm at apex, lobes broadly obovate or 
suborbicular, spreading, (1.5-) 2 cm 0. Stamens 
10, subequal, exserted; filaments linear and laxly 
patently hairy in the lower 2/3, more filiform and 
glabrous apically; anthers narrow-oblong, 3-3.5 
by 0.8 mm. Disk hardly prominent, glabrous. 
Ovary subcylindric, densely clad with forward 
directed hairs which i cover the scales, 5-6 by 
c. 1 mm, rather continuous with the style, the latter 
slightly shorter than the stamens, densely hairy 
and lepidote to the lower Y^, lepidote at apex; 
stigma capitate, deeply 5-lobed. Capsule sub- 
cylindric, 3-3.5 by 0.5-0.6 cm, on fruit-pedicel 
2.5-3 cm by 1.5 mm. Seeds very narrow, 4—5 mm 
including the long tails. 

Distr. Malesia: E. New Guinea (Milne Bay 
Distr.: Sudest L, on Mt Riu), 600-800 m, scattered 
in stiff" scrub on summit or on edge of a precipice. 
Fl. Aug. 

Note. Much related to R. lochae F. v. M. from 
NE. Queensland (the only Rhododendron in 
Australia), which, however, has red corollas, 
laxly hairy ovary (the scales thus clearly visible) 
and 2-5 (rarely up to 7) flowers per umbel. 

245. Rhododendron luraluense Sleum. Notizbl. 
Berl.-Dahl. 12 (1935) 485; Reinwardtia 5 (1960) 
209. 

Small, epiphytic shrub or tree, up to 9 m. 
Branchlets subterete, 2-3 (-4) mm 0, laxly lepidote 
at the upper internode, glabrous below; internodes 
3.5-8 cm. Leaves in 4-7-merous pseudowhorls at 
the upper 2-3 nodes, obovate-elliptic to obovate- 
oblong, apex broadly attenuate, subapiculate or 
obtuse, rarely rounded, base broadly cuneate to 
the petiole, coriaceous to subcoriaceous, glabrous 
above at full age, laxly to very laxly and more 
persistently lepidote beneath (scales very small, 
the early dissolved marginal zone irregularly 
dented, centre dark, slightly impressed), edge 
somewhat recurved, (3.5-)4-5.5(-6) by 2-2.8(-3.2) 
cm, midrib broad and impressed in the lower part 
above, broadly and obtusely prominent beneath, 
nerves 4— 5(-6) irregular pairs, spreading, in- 
conspicuously inarching before the edge, faintly 
impressed or hardly visible above, slightly promin- 
ent beneath, reticulation lax, a little raised beneath 
only; petiole flattened, grooved above, 3-4(-5) by 
1-1.5 mm. Umbels (3-)4-5-flowered. Outer 
perulae ovate-acuminate, inner ones obovate to 
spathulate, all glabrous or laxly lepidote apically 
outside, 0.5-1.2 by 0.3-0.6 cm. Bracteoles filiform, 
glabrous, c. 1 cm. Pedicels rather stoutish, laxly 
substellate-lepidote, more densely so at apex, c. 
L3 cm. Calyx oblique, disk-shaped, very shortly 



i 



Jan. 1966] 



Ericaceae (Sleumer) 



637 



5-lobulate, c. 3 mm 0. Corolla funnel-shaped, 
(3-)3. 5-4(4.5) cm long in all (limb c. 5 cm 
when fresh), white, membranous, sparsely lepidote 
outside, very laxly and shortly pubescent at the 
tube inside, tube 1.5-2 cm, at base c. 4 mm 0, 
gradually widened to 7-9 mm at apex, lobes 
obovate-suborbicular, spreading, (1-)1.5 by 1-1.4 
cm. Stamens 10, reaching — halfway the corolla 
lobes; filaments linear and densely hairy below, 
filiform and glabrous above; anthers oblong, c. 
2.5 mm. Disk prominent, glabrous. Ovary sub- 
cylindric, densely white-hairy and lepidote (the 
hairs ± covering the scales), c. 4 by 1.8 mm, 
rather abrupt with the style, the latter slender, not 
quite equalling the stamens, subdensely to laxly 
hairy and lepidote to about the lower Ya; stigma 
capitate. 

Distr. Melanesia: Solomon Is. (Bougainville, 
Crown Prince Range), twice found. 

Ecol. In rain-forest, or stunted vegetation, 
locally common, at 900-1500 m. Fl. Aug., Oct. 

Vern. Kiribui. 

246. Rhododendron flavoviride J.J.S. Med. Rijks- 
herb. n. 25 (1915) 4; Nova Guinea 12 (1917) 505; 
ibid. 12 (1918) t. 200 a, 204 b-e; Sleum. Reinward- 
tia 5 (1960) 207. 

Much-branched, terrestrial shrub, up to 4 m. 
Branchlets subterete, — densely lepidote at the 
upper intemodes, glabrescent, 2-4 mm 0; inter- 
nodes (1.5-)2.5-5(-8) cm. Leaves in (4-)6-8- 
merous pseudowhorls at the upper 1-2 nodes, 
elliptic or ovate-elliptic, apex obtuse or rounded, 
base mostly rounded, more rarely broad-attenuate, 
coriaceous ,r: stiff, initially lepidote on both faces, 
glabrescent leaving dense minute blackish pits 
above, more persistently and subdensely lepidote 
beneath (scales rather large, flat, marginal zone 
wide, shortly substellate-lobed or -dented, minute 
centre very dark, somewhat impressed), edge 
much revolute, 2.5-4.5 by (l.5-)2-2.7 cm, midrib 
narrow and slightly sunk above, as robust as the 
petiole and much prominent beneath, nerves 6-8 
pairs, spreading, rather irregular, anastomosing, 
slightly depressed above and raised beneath in 
old leaves, reticulation rather dense, slightly 
impressed above, faintly to markedly raised 
beneath; petiole lepidote, subterete, 3-5 by 1-1.5 
mm. Umbels 3-5-flowered. Outer perulae sub- 
orbicular, lepidote dorsally, inner ones obovate to 
spathulatc, glabrous, up to 1.6 by 0.8 cm. Bracte- 
olcs linear to linear-spathulatc, glabrous or lepidote 
apically, up to 1.9 cm. Pedicels stoutish, densely 
substcllalc-lepidote, epilosc, I 1.7 cm. Calyx 
obliquely disk-shaped, very shortly and obtusely 
5-lobcd, 4-5 mm 0. Corolla tubular in outline, 
somewhat oblique and curved, ycllow-grccn or 
pale green, fragrant, (3.5 )4--4.5 cm long in all, 
laxly substcllatc-lcpidotc outside, short-pubcsccnl 
at the tube msidc, tube subcylindric, 2.5-3 cm, 
at base 5-saccatc, 4-5 mm 0, slightly dilated 
upwards, lobes suborbicular, subercct or somewhat 
sprcadmg, 1(-1.2) cm 0. Stamens 10, : as long as 
the corolla; filaments linear below, more filiform 
distally, subdcn.scly to laxly pubescent to nearly 



the top; anthers oblong, c. 2.5 by 0.8 mm, base 
obtuse. Disk hardly prominent, glabrous. Ovary 
subcylindric-conical, base attenuate, apex some- 
what constricted to the style, yellowish velu- 
tinous, the straight hairs covering small scales, c. 
5 by 2 mm; style as long as the stamens, finally a 
little exserted, hairy and lepidote to the lower 
14-'/^' lepidote and whether or not very laxly 
hairy up to the middle, glabrous above; stigma 
capitate. 

Distr. Malesia: W. New Guinea (Oranje Mts: 
Kajan Mts and near Lake Habbema). 

Ecol. In open, swampy localities, or edge of 
forest, c. 3200 m. Fl. Feb., Aug. 

247. Rhododendron christianae Sleum. Reinward- 
tia 5 (1960) 211; Blumea 11 (1961) 128; Crutt- 
WELL, J. R. Hort. Soc. 87 (1962) 81, f. 21. 

Rather compact terrestrial shrub, up to 3 m. 
Branches terete, 7-8 mm 0; bark brownish greyish. 
Branchlets obtusangular and sparsely lepidote at 
the tips, subterete and glabrous at the older parts; 
intemodes (2-)3.5-l I(-I3) cm. Leaves in 3(-5)- 
merous pseudowhorls at the upper l-3(-4) nodes, 
± broadly elliptic or subobovate-elliptic, apex 
rather abruptly and very shortly or more gradually 
acuminate, blunt, sometimes rounded or sligthly 
emarginate, terminal gland thick, not emerging, 
base broadly attenuate into the petiole or more 
rarely cuneate, coriaceous, juvenile ones laxly 
lepidote on both faces, mature ones quite glabrous 
above, laxly to very laxly lepidote for a long time 
beneath (scales small, marginal zone variously 
lobed or dented, centre slightly impressed), flat. 
4-7.5 by 3-4.5 cm, midrib slightly or faintly sunk 
above, much and bluntly prominent in the lower 
part beneath, nerves 5-7 pairs, spreading, divided 
and curved-anastomosing before the edge, rather 
obscure above, a little raised beneath, reticulation 
inconspicuous; petiole subterete, grooved dorsally, 
somewhat rugose, glabrous, (5-)6-10 by 1-1.5 
mm. Umbels (2-)3-4(-5)-flowered. Outer perulae 
ovate to obovate, innermost ones oblong-spathu- 
late, hirsutulous on the dorsal middle line and at 
the apex, margin with early caducous scales, up 
to 1.5 by 0.8 cm. Bracteoles linear-subspathulate 
to filiform, laxly pubescent, c. 1.5 cm. Pedicels 
densely and shortly hairy, laxly lepidote at both 
ends, l-1.8(-2.7) cm by c. I mm. Caly.\ oblique, 
disk-shaped, shortly obtusely 5-lobcd, lepidote and 
pubescent dorsally, c. 4 mm o. Corolla widely 
tubular-subcampanulate, normally deep yellow 
shading to glowing orange, or bright orange at the 
lobes, and yellow at the throat, sometimes salmon 
or paling almost to yellow and darkening to 
russet, occasionally red outside and greenish inside, 
sparsely lepidote outside, laxly hairy at the lower 
third of the tube inside, tube straight, (2.5-) 
3-3.5 cm, at base much 5-saccatc, 7-8(-IO) mm 
0, gradually dilated to 1.5-2 cm at apex, lobes 
broadly obovate or suborbicular, (l.5-)2-2.5 by 
r. 2 cm. Stamens 10, very unequal, the longest 
reaching hallway the corolla lobes, the shortest 
hardly cxscrlcd; filaments linear, r. I mm wide at 
the base, densely and very shortly papillose- 



638 



Flora Malesiana 



[ser. I, vol. 6* 



pubescent in the lower third, glabrous at apex; 
anthers narrow-oblong, (3-)4-5 by 1 mm, base 
very shortly biappendiculate. Disk slightly promi- 
nent, white-velutinous at the upper margin. Ovary 
subcylindric, rather abrupt with the style, white- 
tomentose (and lepidote but scales hidden by the 
hairs), c. 6 by 2.5 mm; style 3-4 cm long, ± as 
long as the corolla tube, densely or upwards more 
laxly hairy and lepidote to the lower Y^, glabrous 
for the 5-7 mm below the thick-capitate, manifest- 
ly 5-lobed stigma. 

Distr. Malesia: SE. New Guinea (Milne Bay 
Distr.: Daga country). 

Ecol. On precipitous rock face or cliff edges, 
overhanging deep gorge or gullies, in full sun. 
"Locally abundant from 600-1525 m. When in 
flower (properly about June), it is a mass of orange 
and a conspicuous sight for miles, even from an 
aeroplane" (N. E. G. Cruttwell, in litt.). Fl. 
May-Oct. 

248. Rhododendron laetum J.J.S. Nova Guinea 
12 (1914) 139, t. 35; in Gibbs, Arfak (1917) 169; 
Kaneh. & Hatus. Bot. Mag. Tokyo 56 (1942) 
479; Sleum. Reinwardtia 5 (1960) 212; Blumea 
12 (1963) 112; Jahrb. Rhodod. Ges. Bremen 
(1963) f. 35. — R. uliginosum J.J.S. Nova Guinea 
12 (1914) 136, t. 33; Kaneh. & Hatus. Bot. 
Mag. Tokyo 56 (1942) 479.— Fig. 40. 

Terrestrial shrub, 0.5-1. 5(-3) m. Branchlets 
obtusangular and laxly lepidote at young shoots, 
3-5 (-6) mm 0, terete and practically glabrous 
later; internodes (2.5-)3-16 cm. Leaves in 4-5(-7)- 
merous pseudo whorls at the upper 1-2 nodes, 
broadly elliptic or subovate-elliptic, apex mostly 
very shortly acuminate or subcuspidate, more 
rarely broadly attenuate, subacute or bluntish, 
sometimes rounded, base mostly rounded, rarely 
very broadly attenuate or subcordate, coriaceous, 
dark green and glossy above, paler beneath, sub- 
densely lepidote on both faces initially, glabrous 
above at full age, ± persistently and rather laxly 
lepidote beneath (scales small, marginal zone 
irregularly substellate-lobed or -dented, early dis- 
solved, minute centre shallowly impressed), finally 
glabrescent, but remaining punctate beneath, 
margin ± flat, (4-)5-9.5 by (2.5-)3-5.3 cm, 
midrib narrow and grooved above, as stout as 
the petiole and obtusely prominent at the base 
beneath, nerves 6-8 pairs, spreading, rather 
obscurely anastomosing before the edge, faintly 
impressed, or not rarely inconspicuous above, 
slightly to dinstinctly raised beneath, reticulation 
lax, a little prominent beneath, or often ± 
obsolete; petiole depressed and grooved dorsally, 
2-5 (-7) by 2-3 mm. Umbels (5-)6-8-flowered. 
Flower buds ovoid, c. 4.5 by 2.5 cm. Outer 
perulae rounded to ovate, apiculate or obtuse, 
initially dorsally with very short substrigose 
hairs, glabrescent and ± smooth finally, inner 
ones ovate-oblong to subspathulate, glabrous or 
nearly so dorsally, to 3.5-5 by 1 .5-2 cm. Bracteoles 
fiiliform, subspathulate at apex, very laxly hairy 
or glabrous, c. 4 cm by 0.5-1 mm. Pedicels 
subdensely stellate-lepidote, moreover puberulous 




Fig. 40 Rhododendron laetum J. J. S. Swamp near 

Iray, Anggi Gigi Lake, Arfak Mts, NW. New 

Guinea, 1900 m (Sleumer, 1962). 



to various degree, 2.5-5.5 by c. 1 mm. Calyx 
obliquely disk-shaped, very shortly obtusely 
5-lobed, lepidote and puberulous dorsally, wavy, 
c. 4 mm 0. Corolla rather broadly funnel-shaped, 
fleshy and fragrant, of a deep pure yellow first, 
becoming suffused with red, orange or salmon 
in later stages, or sometimes dilutely rose-coloured 
then, laxly stellate-lepidote and may be with some 
hairs at the base outside, short-pubescent at the 
tube inside, 6.5-7 cm long and c. 5 cm at the 



ji£ 



Jan. 1966] 



Ericaceae (Sleumer) 



639 




Fig. 41. Rhododendron scabridibracteum Sleum. a. Flowering branchlet, X 2/3, b. stamen, x 1 '/s, c. 
ovary and style, x 1 1/3, rf-e. two inner perulae from outside, x 2/3,/. bracteole, x 2/3 (a-f Hoogland 

& PULLEN 5840). 



limb (when fresh), (4.5-)5-5.5(-6) cm long in all 
when dry, tube 3-3.5 cm, straight, very distinctly 
5-anguIar, at base 5-saccate, 0.7-0.8 cm (1.2 
cm when fresh), dilated to 1.2-2 cm at apex, 
lobes broadly obovate, 2-2.5(-3) by 1.5-2.2 cm. 
Stamens 10, subequal, slightly exscrted; filaments 
linear and subdensely short-pubescent in the 
lower, narrower and quite glabrous in the upper 
half; anthers broad-oblong, base obtuse, 4-5 by 
1.5 mm. Disk much prominent, glabrous in the 
lower, tomcntosc in the upper half Ovary sub- 
cylindric, rather abrupt with the style, 6-8 by 2.5-3 
mm, white-tomentosc (the hairs covering minute 
scales); style stoutish, tomentosc at the base, 
more laxly hairy above to the lower '/2 to 2/,, 
mostly without trace of scales; stigma thick-capi- 
tate. C'a/J5M/esubfusiform-cylindric, curved, densely 
hairy and Icpidote, (3.5-)4-5(-5.5) by c. 0.8 cm, 
on fruit-pcdicci 4-5.5 cm by 2 mm. Seeds 4 6 mm 
including the tails. 

Distr. Malesia: NW. New Guinea (Anggi 
Lakes area in Arfak Mis). 



Ecol. On edge of primary and secondary forest, 
and in open marsh, often in swamps near lake 
shore, 1800-2300 m. Fl. Jan. -Dec. 

Note. Much related to R. zoelleri. 

249. Rhododendron scabridibracteum Sleum. 
Rcinwardtia 5 (i960) 215; Biumca 11 (1961) 
128; ibid. 12 (1963) 112.— Fig. 41. 

Terrestrial or epiphytic shrub or tree, (l-)3-5 
(-12) m, sparsely branched. Branchlcts (3-)4-6 
mm 0, laxly substcllatc-lcpidote at the upper inter- 
nodes, early glabrcscent; intcrnodcs (2-)4-18 cm. 
Leaves in 3-4(~6)-merous pscudowhorls at the 
upper 1-2 nodes, elliptic-oblong, apex broadly 
attenuate, obtuse, sometimes rounded or emargi- 
ate (apical gland not projecting), broadly attenu- 
nate towards the base (very base not rarely sub- 
truncate or very shortly or occasionally broadly 
rounded), coriaceous, glabrous above at full age, 
laxly Icpidote beneath (scales small, dark chestnut 
brown, marginal zone variously substellate-lobed 
or -incised, mmutc centre slightly impressed), 



640 



Flora Malesiana 



[ser, I, vol. 6* 



somewhat convex above, edge ± flat, (6.5-) 
8-13 by (3-)3.5-6(-7) cm, midrib narrowly 
impressed above, as thick as the petiole, blunt 
and much prominent in the lower part beneath, 
nerves 10-12 pairs, spreading, rather straight 
below, curved upwards and anastomosing before 
the edge, with some additional intermediate veins, 
faintly impressed above, prominent beneath, 
reticulation rather dense, hardly impressed 
above, raised beneath, veinlets less visible; petiole 
subterete, rugulose, grooved above, 10-15 by 2-3 
mm. Umbels 6-12-flowered. Outer perulae ovate, 
inner ones obovate to subspathulate, obtusely 
apiculate, at both sides with a tomentum of 
scabrid, subappressed yellowish hairs inflated at 
base, up to 3 by 1.8 cm. Bracteoles almost filiform, 
densely hirsute, c. 2 cm by 0.5-1 mm. Pedicels 
with dense rufescent, substellate and somewhat 
scurfy scales, 2-3.2(-5) cm by c. 1 mm. Calyx 
disk-shaped, shortly (c. 1 mm) 5-dented or -lobed 
(one or the other tooth occasionally up to 4 mm), 
densely lepidote dorsally, 4-5 mm 0. Corolla 
tubular, pale to deep red or crimson, distinctly 
oblique, (5-)5.5-6 cm long in all, densely stellate- 
lepidote outside, hirsutulous by retrorse hairs at 
the tube inside, tube subcylindric, curved, 3-4.5 
cm, at base c. 6 mm 0, slightly dilated to 1-1.5 
cm at apex, lobes erecto-patent, obovate-spathu- 
late, 1.5-2 by 1-1.2 cm. Stamens 10, as long as 
the tube or finally exerted; filaments linear and 
densely clad with stout, forward directed hairs 
below, more laxly to very laxly so and filiform 
upwards, glabrous for some mm at apex; anthers 
oblong, c. 5 by 1.3 mm, base ± obtuse. Disk 
thick-prominent, glabrous below, tomentose above. 
Ovary subobconical-cylindric, rather abrupt with 
the style, yellowish hirsute and lepidote (scales 
completely hidden by the hairs), 8-10 by 3-4 mm; 
style thick, densely set with substrigose hairs and 
minute (here well visible) scales below, more 
laxly so above to nearly the top; stigma turbinate- 
capitate, oblique, very thick. 

Distr. Malesia: New Guinea (Mt Antares and 
Western, Southern and Eastern Highlands). 

Ecol. In lower montane rain- forest or in 
Castanopsis- or Nothofagus-iorest, not rarely on 
tree trunks if epiphytic, at 1950-2400 m. Fl. 
April-Sept. 

Vern. Ome, wamp wamp, Mendi. 

250. Rhododendron longiflorum Lindl. J. Hort. Soc. 
Lond. 3 (1848) 88, fig. on p. 89; Planch. Fl. 
Serres 5 (1849) 477^80, fig.; Becc. Malesia 1 
(1878) 205; Henslow, J. R. Hort. Soc. 13 0891) 
263, f. 42 (fol.), 43 b (fl.); K. & G. J. As. Soc. 
Beng. 74, ii (1905) 77; Ridl. J. Linn. Soc. Bot. 
38 (1908) 314, repr. J. Fed. Mai. St. Mus. 2 
(1908) 120; J. Fed. Mai. St. Mus. 4 (1909) 43; 
ibid. 6 (1915) 158; ibid. 1 (1916) 46; Millais, 
Rhodod. (.1917) 203; Merr. En. Born. (1921) 
462; RiDL. J. Fed. Mai. St. Mus. 10 U922) 250; 
Fl .Mai. Pen. 2 (1923) 219; Burk. & Menders. 
Gard. Bull. S.S. 3 (1928) 280; Holttum, M.A.H. 
A. Mag. 9 (1939) iA, phot.; Renders. Mai. Nat. 
J. 6 (1950) 262; Sleum. Reinwardtia 5 (1960) 



209; Blumea 11 (1961) 127; ibid. 12 (1963) 
112. — R. tubiflorum Low ex Lindl. J. Hort. Soc. 
Lond. 3 (1848) 88, in obs.—R. lobbii Veitch, Cat. 
(1870) 22; Anon. Florist & Pomologist (1870) 
233; Anon. Gard. Chron. (1871) 1323; Henslow, 
J. R. Hort. Soc. 13 (1891) 262, fig. on p. 281 
(fol.), on p. 282 (/{.); Millais, Rhodod. (1917) 
203. — R. javanicum (Bl.) Benn. var. tubiflorum 
Hook. Bot. Mag. (1885) t. 6850; Ridl. J. Str. 
Br. R. As. Soc. n. 23 (1891) 144; ibid. n. 33 
(1900) 103. — R. orion Ridl. var. aurantiacum 
Ridl. Fl. Mai. Pen. 2 (1923) 217; Henders. 
Gard. Bull. S.S. 4 (1928) 280. 

var. longiflorum. 

Large shrub, terrestrial or sometimes epiphytic, 
up to 3 m. Branchlets terete, initially laxly fuscous- 
lepidote at the upper internode, early glabrescent 
below; internodes 2-8 (-12) cm. Leaves opposite 
or mostly in 4— 5-merous pseudowhorls at the 
upper 1-3 nodes, narrow-obovate or obovate- 
elliptic, or 4-_ elliptic, rarely more oblong-elliptic, 
apex shortly and bluntly attenuate or subacumin- 
ate, base usually ± cuneate, rarely broadly 
attenuate, i coriaceous, bright green, juvenile 
ones rather densely lepidote on both faces, 
mature ones glabrous above, rather laxly and sub- 
persistently lepidote beneath (scales small, margin- 
al zone irregularly incised or lacerate, centre 
deeply immersed), flat or slightly convex above, 
margin hardly or not revolute, (3.5-)4-9(-l 1.5) 
by (1.7-) 2.5-3. 5(-4, rarely up to 6) cm, midrib 
slightly immersed above, prominent in its lower 
part beneath, nerves 5-6 pairs, faintly impressed 
above, even less so beneath, or hardly visible on 
both sides, reticulation 0; petiole somewhat 
flattened, grooved above, variable in length, 
3_8(-l2, rarely up to 15) by 1.5-2 mm. Umbels 
rather laxly 5-10-flowered. Outer perulae ovate 
to suborbicular, inner ones broadly spathulate, all 
laxly lepidote dorsally, sometimes also minutely 
sericeous there, i shining, ciliolate, up to 1.2 by 
0.8 cm. Bracteoles filiform to linear-spathulate, 
up to 1.5 cm. Pedicels rather slender, mostly laxly, 
rarely densely hairy, (l-)1.2-1.8(-3) cm, laxly and 
± caducously lepidote. Calyx a thin, oblique, 
very short rim. Corolla tubular below, expanded to 
various degree at the limb, or nearly salver-shaped, 
mostly a little oblique, 4-5(-6) cm long in all, 
magenta or coral red to crimson, rarely very 
dainty orange-scarlet, salmon-pink or pink, 
very rarely pure white, laxly lepidote outside 
especially in the lower half, or glabrescent, pubes- 
cent in the lower part inside, tube cylindric, 
mostly markedly curved, (2.5-)3-4 cm, at the 
distinctly 5-pouched (sometimes yellowish) base 
3-5 mm 0, widened to c. 1 mm at apex, lobes 
suberect or ± spreading, obovate to suborbicular, 
1.5-2(-2.5) by l-1.5(-2) cm. Stamens 10, exserted 
for 1-1.5 cm, not rarely exceeding the corolla; 
filaments linear and subdensely to laxly hairy in 
the lower, glabrous and nearly filiform in the 
upper half; anthers oblong, 2.8-3 by 1 mm, 
base obtuse. Disk densely hairy in the upper half, 
glabrous below. Ovary elongate-conical, 5-6 by 



1 



Jan. 1966] 



Ericaceae (Sleumer) 



641 



2 mm, densely set with ascendent, subappressed 
hairs and with scales, the latter ± covered by the 
hairs, rather abrupt with the slender style, which 
is 3.5^.8 cm long and x exserted, densely 
hairy below, more laxly so upwards to c. y^ or 
more of its length, glabrous for the last 5-10 mm 
below the shortly turbinate-capitate, obscurely 
5-lobed stigma. Capsule elongate-obconical, base 
slightly attenuate, apex acuminate, laxly hairy 
and lepidote, 3-3. 5(^) by 0.4-0.5 cm, the 5 ribs 
conspicuous. Seeds very narrow, c. 4 mm includ- 
ing the long filiform tail on both ends. 

Distr. Malesia: Sumatra (incl. Mentawei Is.), 
Malay Peninsula, Karimata Arch., Borneo. 

Ecol. Epiphytic on lofty trees in primary or 
secondary forest, often along rivers or on mang- 
rove, a woody scrambling terrestrial shrub on 
rocks, hard quartzite ridge, sandstone pavement 
or sandy clay, in extreme 'kerangas' heath forest, 
but also in poor forest in stagnant water, from 
sea-level up to c. 1500 m. Fl. Jan. -Dec. 

var. bancanum Sleum. Reinwardtia 5 (I960) 210. 
— Rhododendron sp. KuRZ, Nat. Tijd. N. I. 27 
(1864) 215. 

Differs from var. longiflorum mainly by smaller 
flowers and longer petioles. Leaves elliptic or 
oblong-elliptic; petiole (9-) 12-20 by 1.5-2 mm. 
Corolla 2-2.5 cm long in all, tube ± straight, base 
5-gibbous, 2.5-3 mm 0, widened to 3.5-4.5 mm 
at apex, lobes obovate-spathulate, 1.5-1.6 by 
0.8-1 cm. Stamens c. 3.5 cm; anthers 2.5 by 1 mm. 
Ovary c. 4 by 2 mm; style 2.3 cm. Ferulae manifest- 
ly sericeous dorsally. 

Distr. Malesia: Banka (top of Mts Maras and 
Menumbing, and Sg. Liat), 425-700 m. 

251. Rhododendron polyanthemum Sleum. Blumea 
12 (1963) 111; ibid. 12 (1964) 341. 

Tree or treelet, 6(-15) m, trunk stoutish or 
slender, or low epiphytic shrub. Branchlets at 
tips with a furry tomentum of rufous stellate 
scales, early glabrescent, subterete; internodes c. 
1 1 cm by 5-8 mm. Leaves in 5-7-merous pseudo- 
whorls at the upper 2 nodes, elliptic-oblong or 
elliptic, apex gradually broadly attenuate or sub- 
acuminate (tip bluntish) to base broadly attenuate- 
rounded (very base shortly cordate and compli- 
cate), young ones at both sides and on the petiole 
covered by a similar indumentum (scales sub- 
dendroid, narrowly stellate-divided up to the 
minute centre, fugacious, leaving no pits), mature 
ones early glabrescent except the midrib, finally 
quite glabrous, (7-)8-l5 by (3-)4-8(-9) cm, 
midrib c. 2 mm wide at base, gradually narrower 
and slightly impressed upwards above, as thick 
beneath at base as the petiole, nerves 9 11 pairs, 
subrcgular, slightly curved, partly deeply forked, 
anastomosmg before the edge, faintly though even 
distinctly impressed above, not much raised or 
inconspicuous beneath, no reticulation; petiole a 
little depressed, grooved above, 2 4 cm 4-5 mm. 
Umhelx (l8-)20 24-flowered. Inner pcrulae thin, 
^ npathulate. glabrous, caducously Icpidote- 
flmbnatc, up to 2.5 by 1.2 cm. Bracteules linear 



to filiform, glabrous, c. 2 cm. Pedicels laxly 
stellate-lepidote, 6-8 cm by c. 1.5 mm. Calyx 
rather oblique, patellar or rim-like, obtusely 
shortly 5-lobed or merely wavy, 4-5 mm 0, 
densely lepidote dorsaly. Corolla funnel-shaped, 
thin, bright scarlet, sweet-scented, subdensely 
lepidote at the tube, less so at the lobes outside, 
laxly hairy in the lower part of the tube inside, 
c. 4 cm long in all, limb c. 3.5 cm 0, tube straight, 
c. 2 cm long, at base 5-6 mm 0, 5-pouched, 
dilated to 1.5-2 cm at apex, lobes suberect- 
patent, broadly obovate-spathulate, c. 2 by 
1.5-1.8 cm, slightly retuse. Stamens 10, unequal, 
reaching halfway the lobes; filaments linear and 
for their lower '/j subdensely to laxly hairy, 
narrower and glabrous above; anthers elliptic, base 
obtuse, 2.7-3 by 1.2 mm. Disk thick, glabrous 
at the much prominent ribs, hairy between. Ovary 
subconical, 5-ribbed,densely clad with atomentum 
of white, forward directed hairs which covers the 
scales, 5-6 by 3 mm, rather abrupt with the style, 
the latter thickish, 1.6-2 cm, densely to more 
laxly and rather patently hairy in the lower half, 
glabrous for the rest; stigma thick, depressed- 
capitate, bearing 5 short club-shaped lobes. 

Distr. Malesia: North Borneo (summit area 
of Mt Trus Madi and Mt Kinabalu: Mesilau 
East R. Basin). 

Ecol. In dense mossy or submossy forest on 
crest of ridge, 1720-2285 m. Fl. Jan. 

252. Rhododendron kochii Stein, Gartenflora 34 
(1885) 193, t. 119; Jahresber. Schles. Ges. Vaterl. 
Cult. f. d. Jahr 1885 (1886) 413, 414; Vidal, 
Rev. PI. Vase. Filip. (1886) 172; Merr. Publ. 
Gov. Lab. Philip, n. 29 (1905) 41; Philip. J. Sc. 3 
(1908) Bot. 380; Elm. Leafl. Philip. Bot. 3 (1911) 
1105; MiLLAis, Rhodod. (1917) 199; Merr. En. 
Philip. 3 (1923) 243; Millais, Rhodod. ser. 2 
(1924) 168; Copel./. Philip. J. Sc. 40 (1929) 160, 
t. 7, f. 6-7, t. 8, t. II, f. I; Sleum. Reinwardtia 
5 (I960) 212; Blumea II (1961) \2%.— Rhododen- 
dron sp. (aff. R. javanicum) Vidal, Sinopsis Atl. 
(1883) 30, t. 60, f. P.—/?. javanicum [non (Bl.) 
Benn.] F.-Vill. Nov. App. (1883) 353; Vidal, 
Rev. PI. Vase. Filip. (1886) 170.—/?. schaden- 
bergii {non Warb.) Merr. Publ. Gov. Lab. 
Philip, n. 29 (1905) 41, p.p. 

Shrub, sometimes tree-like; stem 3-6(-10) m, 
7-15 cm 0. Branchlets subterete, (2-)3-4 mm 0, 
fugaciously lepidote; internodes (2.5-)5-9 cm. 
Leaves in (3-)4-5-nierous pseudowhorls at the 
upper 1-2 nodes (several other reduced and early 
caducous ones scattered along the internodes), 
oblong to ovate-oblong, or subovate-clliptic, apex 
rather shortly and ! abruptly acuminate, acute, 
base mostly broadly attenuate to the petiole, 
rarely subrolundatc, subinequilatcral, coriaceous, 
densely lepidote on both sides initially, early 
glabrescent above, laxly subpersistenlly lepidote 
beneath (scales flat, rather small, relatively wide 
marginal /one variously substcllalc-lobcd or -dent- 
ed, minute centre shallowly impressed), finally 
glabrescent beneath too. entire or slightly wavy, 
very slightly or not revolulc at the margin, 8-14 



642 



Flora Malesiana 



[ser. I, vol. 6* 



(-18) by (2.5-)3-5 cm, midrib dilated at the base, 
rather suddenly narrowed and a little impressed 
above, as thick as the petiole and obtusely promin- 
ent in the lower part beneath, nerves 8-10 pairs 
spreading and ascending, inarching near the edge, 
slightly impressed or raised above, distinctly 
prominent beneath, reticulation rather dense, 
raised to various degree on both faces, or at least 
underneath; petiole — flattened, reddish, 1.5-2 
cm by c. 2 mm. Umbels 1 0-20-flowered. Outer 
perulae ovate, obtuse, inner ones obovate to 
spathulate, laxly lepidote apically or glabrous 
dorsally, often rugose by minute tubercules, 
margin set with caducous scales, up to 2 by 1.2 
cm. Bracteoles filiform to linear-subspathulate, up 
to 2 cm. Pedicels — densely clothed with pale 
substellate scales, no hairs, 2-3(-3.5) cm by 1 mm. 
Calyx obliquely disk-shaped, shortly obtusely 5- 
lobed, 3-4 mm 0, densely lepidote dorsally. 
Corolla tubular, white, scentless, 3.5-4(-4.5) cm 
long in all, ± laxly set with substellate scales 
outside, subdensely short-hairy at the tube inside, 
tube (2-)2.5(-3) cm, base much 5-saccate, 5-6 mm 
0, slightly widened to apex, lobes subhorizontal- 
ly expanded, obovate-suborbicular, 1.2-1.6 by 
1-1.5 cm. Stamens 10, subequal, reaching about 
half the corolla lobes, filaments linear and sub- 
densely patent-pubescent to the lower Vi-V^, 
glabrous and filiform for the rest; anthers obovate- 
oblong, base shortly though distinctly biap- 
pendiculate, (3-J3.5 by 1 mm. Disk glabrous. 
Ovary subcylindric, white-velutinous, the hairs 
covering the also present scales, gradually narrow- 
ed to the style, 5-6 by 2.5 mm; style nearly as long 
as the stamens, subdensely to laxly and exclu- 
sively hairy to the lower V2-H. glabrous below the 
capitate stigma. Capsule subcylindric, laxly lepi- 
dote, more densely hairy, (2.5-)3-3.5 by 0.5-0.6 
cm, on fruit-pedicel up to 5 cm by 1 mm. 

Distr. Malesia: Philippines (Luzon, Mindoro, 
Negros, Mindanao). 

Ecol. On ridges in mossy forest, 1000-2300 m. 
Fl. Jan. -Dec. 

Vern. Malagas, Bag., hablang. If. 

253. Rhododendron brevipes Sleum. Reinwardtia 
5 (1960) 213. 

Shrub, up to 3 m. Branchlets subterete or 
obtusangular and subdensely clothed with flat, 
substellate scales in the youngest parts (3-4 mm 
0), terete and glabrous in the older ones (c. 7 mm 
0); internodes 7-10 cm. Leaves in i 5-merous 
pseudowhorls at the upper 1-2 nodes, oblong to 
elliptic-oblong, apex shortly acuminate, subacute, 
broadly attenuate towards the base (very base 
subtruncate-obtuse), subsessile, coriaceous, upper 
surface glabrous but densely minutely pitted, 
undersurface subdensely lepidote (scales flat, thin, 
rufescent, variously lobed or dented at the margin- 
al zone, rather small centre hardly sunk), flat, c. 
15 by 5-7 cm, midrib as wide as the petiole at the 
base, much narrowed upwards and slightly 
impressed above, much prominent beneath, nerves 
8-10 pairs, with additional, less distinct ones 
between them, all erecto-patent, ± straight at 



the base, more curved distally, indistinctly 
anastomosing along the edge, flat or faintly 
raised above, more distinct and raised beneath, 
reticulation visible above, nearly obscure beneath; 
petiole flattened, l-2(-3) by 3-4 mm. Umbels 
6-7-flowered. Outer perulae ovate-acuminate, 
with a distinct sharp point {"i-A mm), very laxly 
lepidote along the dorsal middle-line or glabrous 
outside, densely lepidote at the apex inside, up to 
2 by 1 cm, inner ones oblong-subspathulate, 
glabrous, c. 2.5 by 0.6 cm, all caducously lepidote 
along the margin. Bracteoles filiform, subglabrous, 
c. 2 cm by 1 mm. Pedicels dilated at the very 
apex, laxly lepidote, not hairy, 4-5 (-6) cm by 
c. 1.5 mm. Calyx oblique, rather obsolete, a 
wavy rim, hardly 3 mm 0. Corolla funnel-shaped, 
intensely orange at the limb, yellow at the tube, 
7.5-8 cm long in all, laxly to very laxly substellate- 
lepidote outside, short-pubescent at the lower ^^ 
of the tube inside, tube straight, c. 4 cm, at base 
manifestly 5-saccate, 4—5 mm 0, widened to 
1.5 cm at apex, lobes expanded, obovate, c. 4 by 
2.5 cm. Stamens 10, unequal, the longest reaching 
halfway the corolla lobes; filaments linear and 
subdensely hairy in the lower, filiform and glabrous 
in the upper half; anthers oblong, base obtuse, 
c. 4 by 1.2 mm. Disk very prominent, whitish 
tomentellous at the upper margin, glabrous below. 
Ovary cylindric, continuous with the style, very 
densely lepidote, more laxly and shortly white- 
hairy, 10-12 by 2-2.5 mm; style c. 5.5 cm, ± 
densely lepidote and short-pubescent in the lower 
2/3, more laxly lepidote and epilose to apex; 
stigma thick-capitate, with 5 distinct, almost club- 
shaped lobes. Capsule cylindric, lepidote and 
hairy, c. 8 cm by 3^ mm; pedicel 7.5-8 cm by 
c. 2 mm. 

Distr. Malesia: E. New Guinea (Western 
Highlands: Karap, Jimmy Valley), c. 1725 m. 
Once found. Fl. fr. June. 

254. Rhododendron aurigeranum Sleum. Reinward- 
tia 5 (1960) 214; Blumea 11 (1961) 128; ibid. 
12 (1963) 112; Jahrb. Rhodod. Ges. Bremen 
(1963) f. 16. 

Shrub or treelet, (0.5-)l-2.5(^) m. Youngest 
internodes 2 mm 0, a little flattened and densely 
lepidote, the older ones 3-4(-5) mm 0, terete 
and early glabrescent; internodes 5-15 cm. 
Leaves in ± 5-merous pseudowhorls at the upper 
1-2 nodes, oblong, apex short-acuminate, sub- 
acute, base cuneately narrowed into the petiole, 
thin-coriaceous, glabrous above at full age 
(minutely pitted under the lens), densely lepidote 
beneath in young, less so in mature leaves (scales 
rather small, rufescent, scurfy, — detersile, sessile 
or very shortly stalked, marginal zone irregularly 
and — deeply stellate-incised, centre not rarely 
deepened and extended into a very short stipe 
or foot, not or hardly impressed, flat, (7-)8-10 
(-13) by 3-4(-5) cm, midrib dilated at the base, 
narrowed and slightly impressed, thick and promi- 
nent beneath, nerves 6-8 pairs, rather straight 
below, curved and obsoletely anastomosing before 
the edge, nearly flat above, slightly raised beneath. 



Jan. 1966] 



Ericaceae (Sleumer) 



643 



reticulation subinconspicuous on both faces; 
petiole compressed and grooved dorsally, lepidote, 
6-8 by 1.5-2 (rarely up to 3) mm. Umbels 8-10 
(-14)-flowered. Outer perulae ovate-acuminate, 
apex blunt or very shortly contracted into a 
sharp point, — densely lepidote on both sides at 
the ape.x, glabrous for the rest, inner ones obovate- 
oblong to subspathulate, glabrous dorsally, all 
with caducous scales along margin, 1.5-2 by 
0.4-1 cm. Bracteoles linear-subspathulate, 1-1.5 
cm by 1-2 mm. Pedicels stoutish, 2-4(-5) cm by 
1-1.5 mm, subdensely stellate-lepidote, very 
la.xly though distinctly short-pubescent. Calyx 
disk-shaped, wavy, c. 3 mm o. Corolla funnel- 
shaped, orange or orange-yellow initially, fading 
to yellow, 6-7(-8) cm long in all, laxly stellate- 
lepidote outside, laxly pubescent at the lower half 
of the tube inside, tube straight, rather markedly 
5-angular, (3-)3.5-4 cm, at base very distinctly 
5-saccate, 4-5 mm 0, gradually widened to 1-1.5 
cm at apex, lobes expanded, obovate, 2-3 by 
1.8-2.5 cm. Stamens 10, subequal, reaching zr 
halfway the corolla lobes; filaments linear, sub- 
densely hairy in the lower third, glabrous above; 
anthers broad-oblong, 3.5-4 by c. 1.3 mm, base 
obtuse. Disk very prominent, white-tomentellous 
at the upper margin. Ovary subcortical- cylindric, 
tapering to the style, densely and shortly hairy 
and lepidote, c. 10 by 2.5 mm; style robust, 
densely hairy and lepidote to the lower Y^, 
densely and exclusively lepidote for the sub- 
sequent c. 2 mm, glabrous for the upper c. 10 mm; 
stigma thick-capitate and deeply 5-lobed. Capsule 
elongate-subfusiform-cylindric, shortly attenuate 
at the base, longer so at the apex, hairy and 
lepidote, — curved, (6-)8-10 cm by 5(-6) mm; 
valves much twisted. Seeds (7-)8-10(-12) mm 
including the tails. 

Distr. Malesia: New Guinea (Morobe Distr.: 
mainly the Bulolo-Wau area). 

Ecol. Occasionally in floating immature peat 
in forest clearing, generally in rather dry grassy 
or shrubby hillside together with Grevitlea papuana 
DiELS or on schistes or steep rock, 915-1740 m, 
locally common. Fl. Feb. -Oct., fr. July-Sept. 

Note. Worth to be cultivated for its showy 
flowers. 

255. Rhododendron zoelleri Warb. Bot. Jahrb. 16 
(1892 J 15, 24; K. .SfH. & Laut. Fl. Schutzgcb. 
(1901) 488; Schltr, Bot. Jahrb. 55 (1918) 156; 
Sleum. Rcinwardtia 5 (I960) 216; Blumca II 

(1961) 128; Cruttwell, J. R. Hort. Soc. 87 

(1962) 85, f. 22; Sleum. Blumea 12 (1963) 113; 
Jahrb. Rhodod. Ges. Bremen (1963) f. 33; 
Sf.HNEiDER, Jahrb. Dcutsch. Rhodod. Gcs. 
Bremen (1965) f. I. — R. asparanoides Wernh. 
Trans. Linn. Soc. 11, Bot. 9 (1916) 94.—/?. 
moszkowskii Schltr, Bot. Jahrb. 55 (1918) 161. 
—R. oranum J.J.S. in Fcddc, Rep. 30 (1932) 
167.—/?. dociersii J.J.S. Nova Guinea 18 (1936) 
98, t. 21, I, cf. DorTER.s van Leeuwen, Trop. 
Natuur 25 (1926) 182, f. 9—/?. laelum Uum J.J.S. 
1914) J.J.S. NovaGuinca 18 (1936)98.— Mr, 42-43. 

Weak epiphytic or ilillly erect, also sub- 



scandent terrestrial shrub respectively treelet, up 
to 6(-10) m, stem up to 15 cm 0; bark light grey. 
Branchlets terete, 3-5 mm 0, laxly and i caduc- 
ously lepidote at the upper internodes, glabrescent 
below; internodes (3-)5-10(-14) cm. Leaves in 
4-5(-6)-merous pseudowhorls at the upper 1-2 
nodes, (broadly) elliptic to oblong-elliptic, rarely 
subovate-elliptic or ovate-oblong, apex short- 
acuminate, subacute or blunt, base broadly 
attenuate, sometimes rounded or nearly so, often 
slightly inequilateral, ± thin-coriaceous, glabrous 
above at full age, laxly lepidote for a rather long 
time beneath (scales small, marginal zone various- 
ly substellate-lobed or -dented, minute centre ± 
shallowly impressed), finally glabrescent but re- 
maining punctulate beneath, edge ± flat,7-14(-17) 
by (3-)4-7(-9, very rarely up to 10) cm, midrib 
narrowly grooved above, as stout as the petiole 
and obtusely prominent at the base beneath, 
nerves 10-12(-14) pairs, oblique, irregular, often 
running a little down on the midrib, curved up- 
wards and anastomosing before the edge, mostly 
but slightly raised (or in very old leaves even 
somewhat impressed) above, more distinctly 
prominent beneath, reticulation rather lax, 
mostly ± distinct, but not much raised on both 
faces; petiole flattened and grooved dorsally, 
lepidote initially, (0.5-)l-1.5 cm by 1.5-2 mm. 
Umbels (4-)5-8-flowered. Floral buds oblong- 
ovoid, 3.5-5 by 2-2.5 cm; outer perulae sub- 
orbicular, more inner ones ovate to oblong- 
obovate or spathulate, all obtuse and covered for 
a long time by a tomentum of sericeous to sub- 
strigose. greyish yellowish rather long hairs, in 
which the sparse scales are well visible as dark 
points, ± glabrescent but mostly remaining rough 
to the touch in later stages, margin with caducous 
scales, up to 5 by 2.5 cm. Bracteoles linear- 
subspathulate, short-hairy or hirsutulous, up 
to 3 cm. Pedicels laxly to subdensely stellate- 
lepidote, moreover mostly sparsely pubescent 
in the upper part, sometimes laxly pubescent and 
lepidote all along, (1.5-)2-2.5(-4) cm by 1-1.5 
mm. Calyx obliquely disk-shaped, laxly lepidote 
and hairy outside, obtusely 5-lobed, c. 5 mm 0. 
Corolla funnel-shaped, lobed almost halfway, 
yellow to orange or salmon, or salmon-reddish, or 
the tube yellow and the lobes sufl"uscd with red, 
very rarely white or greenish white t? hybrid with 
R. phaeopeplum, cf. there), often sweet-scented, 
(5-)6-7.5(-8.5, occasionally up to 10) cm long in 
all, sometimes with some hairs at the base (within 
the pouches) and otherwise sparsely lepidote 
outside, laxly hairy at the lower half of the tube 
inside, tube straight or nearly so, 3-4(-4.5) cm, 
base distinctly 5-saccatc, 5-8 mm o, gradually 
widened to 1.5-2 cm at apex, lobes expanded, 
obovate-spathulate. 2.5-3.5(-4, rarely up to 4.5) 
by 2-2.5(-3) cm (up to 7.5 cm when fresh). 
Stamens 10, subequal, reaching I halfway the 
corolla lobes; filaments linear and J densely to 
more laxly set with spreading hairs in the lower 
1/3-I/2, glabrous and fililbrm for the rest; anthers 
oblong, somewhat curved, base obtuse or rarely 
obscurely biappcndiculatc, (4-)6 8 by l.5(-2) mm. 



644 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 42. Rhododendron zoelleri Warb. a. Habit, x 2/3, b. stamen, x 1, c. ovary and style, x \, d. fruit 

X 2/3 {a-d BW 3058 Versteegh). 



Disk prominent, densely hairy to tomentose at 
least in the upper part. Ovary subcylindric, 
whitish tomentose by rather stiff hairs, which 
are directed forward and cover minute scales, 
subabrupt with the style, 9-10(-12) by c. 3 mm; 
style apparently first ± equalling the corolla tube, 
finally ± equalling the stamens, densely to 
laxly hairy in the lower 2/3-34, the scales there 
absent or present, in the latter case more obvious 
above the lower half of the style, where the 
hairs become more lax or even may disappear 
completely, glabrous for the rest; stigma turbinate- 
capitate. Capsule elongate-cylindric, attenuate at 
both ends, slightly curved, ± densely hairy and 
lepidote, 5-7(-8) cm by (5-)6-7(-8) mm, rarely 
shorter; pedicel 3^ cm by c. 2 mm. Seeds 8-10 
mm including the long tail at both ends. 

Distr. Malesia: Moluccas (W. Ceram: Ora 
Mts) and New Guinea. 

Ecol. Epiphyte in high forest, terrestrial in 
lighter mountain forest of Castanopsis; also in 
secondary growths, in open spots (artificial 



grassland), hanging over precipices and rivers, in 
hill scrubs, from sea-level to 1500, rarely up to 
2000 m, not rarely gregarious, on poor sandy or 
clayey soil, but said to occur also on limestone. 
Fl. fr. Jan. -Dec. 

Vern. Krobua, mamba'uk, Kebar, atek, Karoon, 
warade, Kapauku, mompera, Samor, powida, 
Kutubu, fabeh, Wapi, tindili, Orne. 

Note. A form different only by smaller anthers 
(3-4 mm) apparently limited to SE. New Guinea 
(Central and Milne Bay Distr.). For natural 
hybrids with R. phaeopeplum see there. 

DocTERS VAN Leeuwen observed the flowers of 
R. zoelleri visited during daytime by butterflies as 
Ornithopthera and Papilio (Trop. Natuur 15, 
1926, 182, 185). 

256, Rhododendron maxwellii Gibbs, J. Linn. Soc. 
Bot. 42 (1914) 103; Merr. En. Born. (1921)462; 
Sleum. Reinwardtia 5 (1960) 217; Blumea 11 
(1961) 128; ibid. 12 (1963) 113; Jahrb. Deutsch. 
Rhodod. Ges. Bremen (1965) f. 17. 



i 



Jan. 1966] 



Ericaceae (Sleumer) 



645 




Fig. 43. Rhododendron zoelleri Warb. Kebar valley, 700 m, with Gleichenia, NW. New Guinea (van 
RoYEN & Sleumer 6850) (Sleumer, 1961). 



Shrub or treelet, occasionally epiphytic. Branch- 
lets terete, 3-4(-5) mm 0, glabrous or practically 
so; intcrnodes 2-9 cm. Leaves in 3-4(-5)-mcrous 
pseudowhorls at the upper l-2(-3) nodes, 
elliptic to oblong-obovate, sometimes obovate, 
apex broad-attenuate and obtuse, not rarely 
rounded, base broadly attenuate to subrotundate, 
coriaceous, glabrous above at full age, laxly 
lepidotc beneath (scales very small, marginal zone 
variously lobcd or dented, i- early dissolved, 
centre dark, persistent, i- deeply impressed), 
margin t revolute, lamina i convex above in 
herb.. (4-)5-9.5 by (2)2.5-5 cm, midrib narrow 
and deeply impressed above, as thick as the petiole 
at the base beneath, obtusely prominent and 
longitudmally wrinkled, nerves 5-6 pairs, spread- 
ing and curved-anaitomosing. ± deeply impressed 



above and rather sharply prominent beneath 
providing a markedly bullate aspect, reticulation 
lax and subimpressed above, or mostly rather 
inconspicuous; petiole flattened and grooved 
dorsally. lepidotc initially, rugulose, 5-10 by 
2-3(-4) mm. Umbels 5-1 3-flowered. Flower buds 
ovoid-ellipsoid. 2 by 1.5 cm in a not yet fully 
developed stage, outer perulac suborbicular, 
apiculate, inner ones obovate, obtuse, all dorsally 
laxly lepidote, margin with caducous scales. Pedi- 
cels subdensely rufous-stellate-lcpidotc, besides 
whether or not laxly puberulous, 2-3.5 cm by 
0.8 mm. Calyx disk-shaped, obscurely lobcd. lepi- 
dotc dorsally, c. 4 mm o. C«w//rt shortly funnel- 
shaped, i.e. tubular below, much expanded at the 
limb, lobcd from halfway to nearly Vy, yellow, 
sometimes deep cream to pink or pale salmon, 



646 



Flora Malesiana 



[ser. I, vol. 6* 



with a strong delicate scent, c. 2.7 cm long in all, 
± densely set with substellate to subentire scales 
outside, laxly hairy in the lower part of the tube 
inside, tube straight, 1-1.2 cm, base 4-5 mm 0, 
widened to 6-8 mm at apex, lobes spreading, 
oblong-obovate, 1.3-1.8 by 0.9-1.2 cm. Stamens 
10, subequal, somewhat exserted; filaments linear 
and with patent hairs in the lower third, more 
filiform and glabrous apically; anthers oblong, 
2.5-3.3 by 0.8 mm. Disk hairy at the upper 
margin. Ovary subconical, densely clothed with 
subpatent, rufescent hairs covering the scales, ± 
abrupt with the style, c. 4 by 2 mm; style slender, 
patent-hairy at and near the base only; stigma 
subcapitate. Capsule apparently c. 1.5 cm long. 

Distr. Malesia: North Borneo (Mt Kinabalu). 

Ecol. In high forest, mossy jungle or open 
exposed crest of spur among bracken, scrub and 
grasses, rather rare, 1830-3050 m. Fl. Feb. -Aug. 

257. Rhododendron nervulosum Sleum. Bot. 
Jahrb. 71 (1940) 146; Reinwardtia 5 (1960) 217; 
Blumea 11 (1961) 128; /6/</. 12 (1963) 113; Jahrb. 
Deutsch. Rhodod. Ges. Bremen (1965) f. 20. — 
Fig. 44. 

var. nervulosum. 

Small, few-stemmed shrub, generally terrestrial, 
rarely epiphytic. Branchlets 1.5-2(-2.5) mm 0, 
glabrous; internodes 2-10 cm. Leaves in 3-5- 
merous pseudowhorls at the upper 2(-4) nodes 
(other much reduced fugacious ones scattered 
along the internodes), narrow-lanceolate, apex 
(sub)acutely acuminate, cuneate towards the 
the base to a very short petiole, sessile or nearly 
so, coriaceous to subcoriaceous, finally glabrous 
above, laxly lepidote beneath (scales small, 
marginal zone irregularly, often but shallowly 
substellate-lobed, early dissolved and marked by 
a dark halo, centre minute, faintly or not impres- 
ed), edge ± flat, (2.7-)5-7.5(-9) by (0.6-)l-1.4 
(-1.6, very rarely up to 2) cm, midrib narrow and 
and slightly impressed above, i broadened and 
obtusely prominent beneath, nerves c. 5 pairs, at 
a rather acute angle, steeply ascending, ± straight, 
obsoletely or not anastomosing, forming a ± 
dense and slightly prominent network with the 
veins and veinlets on both faces; petiole 1-2 by 
1-2 mm. Umbels (l-)2-3-flowered. Ferulae mem- 
branous, outer ones ovate, inner ones ovate- 
oblong, all ± long-acuminate, the very apex 
tapering to a 2-3 mm long sharp point, glabrous 
dorsally, up to 2.8 by 1 cm. Bracteoles subulate, 
up to 1.5 cm. Pedicels laxly lepidote, 1-1.3 cm by 
1(-1.5) mm. Calyx oblique, thick, disk-shaped or 
more rim-like, obsoletely 5-lobed, subglabrous, c. 
5 mm 0, Corolla subcampanulate, lobed, to i half- 
way, rather fleshy, shining, bright scarlet, 3-3.5 
cm long in all, glabrous in- and outside, tube 
1.7-2.3 cm, straight, base 6-8 mm 0, gradually 
widened to c. 1.5 cm at apex, lobes suberect, 
obovate-subspathulate to suborbicular, c. 1.5 by 
1 cm. Stamens 10, c. 2 cm; filaments linear, 
dilated and rather laxly hairy in the lower Vi, 
glabrous and. narrower for the rest; anthers 




Fig. 44. Rhododendron nervulosum Sleum. var. 

nervulosum. On old log above Mesilau East R., 

1950 m, Mt Kinabalu, N. Borneo (Sleumer 4733) 

(Sleumer, 1963). 

elongate-oblong, (4.5-)5-6 by 1.5 mm, each cell 
shortly appendiculate at base. Disk prominent, 
glabrous. Ovary conical, tomentose, the hairs 
covering the scales, 5-6 by 2.5-3 mm, abrupt 
with the style, the latter slender, glabrous, 8-10 
mm; stigma subcapitate. Immature capsule 
ovoid-cylindric, densely hairy and lepidote to 
judge from the developed ovary (1 by 0.6 cm); 
pedicel 2.8 cm. 

Distr. Malesia: N. Borneo (top of Bt Raja 
(1900 m) and Mt Kinabalu). 

Ecol. In mossy forest or more open mossy 
jungle, on open and sunny boulders or landslides 
with sparse scrub, not common, (1900-)2440- 
3050 m. Fl. March- Aug., Dec. 

var. exuberans Sleum. Blumea 12 (1963) 113. 

Epiphyte. Leaves (7-)9-ll by (2-)2.5-3.7 cm. 
Corolla scarlet, c. 2.5 cm long in all. 

Distr. Malesia: Borneo (Sarawak: Mt Mulu), 
mossy forest, 1430 m. Once found. Fl. June. 

258. Rhododendron salicifolium Becc. Malesia 1 
(1878) 202; For. Born. (1902) 147; Merr. En. 
Born. (1921) 463; Sleum. Reinwardtia 5 (1960) 
217, non Bl. 1863-83. 
Terrestrial shrub, c. 60 cm. Branchlets terete, 



) 



Jan. 1966] 



Ericaceae (Sleumer) 



647 



1.5-2(-3) mm 0, upper 1-3 interaodes laxly 
lepidote; internodes 3-8 cm. Leaves in 4-5(-7)- 
merous pseudowhorls at the upper 2-4 nodes, 
narrow-lanceolate, apex acuminate and slightly 
curved, subacute or acute, base cuneate to the 
petiole, thin-coriaceous, glabrous above at full 
age, sparsely lepidote beneath (scales flat, small, 
marginal zone variously, often obtusely and 
shallowiy substellate-lobed, centre minute, hardly 
or not impressed), edge — flat, 10-15 by (0.6-) 
1-2 cm, midrib narrow and impressed above, 
prominent beneath, nerves c. 10 pairs, high- 
ascending and joined along the edge, slightly 
raised above, faintly so or not visible beneath, 
nerves and veins subdensely reticulate and very 
slightly raised above only; petiole grooved above, 
(3-)4— 6 by 1-1.5 mm, passing imperceptibly into 
the lamina. Umbels (l-)2-3-flowered; flowers 
nodding. Bracteoles subulate, glabrous, up to 1 
cm. Pedicels lepidote at base and apex, i glabrous 
between, 1.2-1.7 cm by 0.8 mm. Ca/v.v disk-shaped, 
obtusely and very shortly 5-lobed, laxly lepidote 
dorsally, 3-4 mm 0, Corolla funnel-shaped, 
tubular below, 5-Iobed from halfway to nearly 
•/s, yellow or buff orange, glabrous outside, laxly 
pubescent at the lower half of the tube inside, 
2.8-3.2 cm long in all, tube straight, 1.2-1.5 cm, 
base 5-saccate, c. 4 mm 0, gradually widened to 
c. 1 cm at apex, lobes much spreading, obovate- 
spathulate, 1.4—1.7 by 1-1.2 cm. Stamens 10, 
unequal, c. 2 cm; filaments linear and densely 
pubescent in the lowest 3-4 mm, glabrous and 
more filiform for the rest; anthers obovate- 
oblong, c. 2.5 by 0.8 mm, base obtuse. Disk 
prominent, pubescent at the upper margin. Ovary 
— densly hairy and lepidote, c. 4 by 2 mm, sub- 
abrupt with the style, the latter equalling the 
stamens, pubescent at the base only; stigma capi- 
tate, becoming disk-shaped and deeply 5-lobed. 
Capsule subfusiform-cylindric, 2.5-3.5 by 0.4 cm. 
Distr. Malesia: Borneo (Sarawak: on summit 
rocks of Mt Mattang). Fl. Jan. 

259. Rhododendron lanceolatum Ridl. J. Str. Br. 
R. As Soc. n. 63 (1912) 60; Merr. En. Born. 
(192!) 462; Sleum. Rcinwardtia 5 (i960) 217. 
— /?. pariitum J.J..S. Bull. Jard. Bot. Btzg ill, 13 
(1935) 453. 

Shrub; bark brown. Branchlets 2-3 mm 0, 
very densely clad with rufcscent substcllate scales 
at the upper internodes, glabrcsccnt; internodes 
3.5-8 cm. Leaves in 3-4-mcrous pseudowhorls at 
the upper 1-2 nodes, lanceolate or subovatc- or 
oblong-lanceolate, rarely oblong-ovatc, sessile or 
nearly so, the sometimes slightly curved apex 
acuminate, acute, base obtuse to rounded, very 
densely lepidote on both faces when juvenile, with 
sparse flat scales or finally quite glabrous but 
pitted above, subdensely lepidote beneath for a 
long time, especially at the base of the blade 
(scales rather small, thin and flattish, marginal 
zone irregularly substcllatc-lobed or -dented, 
mmutc centre shallowiy impressed, leaving dark 
pits), flat. 6.5-11.5 by 2.8-4 cm, midrib stout 
and obtusely prominent in the lower half, similarly 



dilated but flat or even slightly impressed beneath, 
nerves (8-) 10-1 2 pairs, at a nearly right angle, 
curved and anastomosing before the edge, 
slightly raised above, obsolete beneath, reticula- 
tion, if any, subinconspicuous above only; 
petiole 0-2 by 2-3 mm. Umbels (3-)4-6-flowered. 
Outer perulae subulate to ovate-acuminate (these 
then often shortly subulate at the apex), inner 
ones oblong-obovate to -spathulate, apiculate, 
all subdensely ± caducously lepidote outside, up 
to 2 by 1 cm. Bracteoles subsubulate-linear, 
lepidote, up to 1.5 cm. Pedicels densely lepidote 
and short-pubescent, 1.5-2.7 cm by 0.7 mm. 
Calyx very oblique, disk-shaped, with distinct, 
very short blunt lobes, lepidote and pubescent 
dorsally, c. 4 mm 0. Corolla subcampanulate, 
5-lobed to ± %, c. 2 cm long in all, orange- 
yellow, membranous, -± densely lepidote at the 
tube and the lower part of the lobes outside, 
glabrous or slightly puberulous inside, tube up to 
7 mm, base 4—5 mm 0, gradually widened to c. 
1 cm at apex, lobes much expanded, oblong to 
subobovate, 1.2-1.7 by 0.8-1.2 cm. Stamens 10, 
unequal, 1 .6-1 .9 cm; filaments linear and woolly in 
the lower 2/3, narrower and glabrous for the 
rest; anthers oblong, curved, 3.5-4 by 1 mm, base 
obtuse or very shortly bilobed. Disk thick- 
prominent, glabrous or somewhat hairy at the 
upper margin. Ovary conical, woolly, obtuse, i.e. 
very abrupt with the style, c. 4 by 3 mm; style 
slenderly columnar, thickened towards the subcapi- 
tate stigma, c. 7 mm, with some hairs at the base. 
Distr. Malesia: Borneo (Sarawak: Mt Batu 
Lawi at Upper Limbang R., and Mt Batu Tiban), 
1125-1700 m. Fl. Oct.-Dec. 

260. Rhododendron sumatranum Merr. Pap. 
Mich. Ac. Sc. 19 (1933) 182; J.J.S. Bull. Jard. 
Bot. Btzg III, 13 (1935) 450; Sleum. Reinwardtia 
5 (1960) 217. 

Much branched terrestrial shrub, 1.5-3 m. 
Branchlets terete, (1.5-)2-3 mm 0, subdensely set 
with flat, substellate scales at the upper internodes, 
glabrescent; internodes (2-)4-8.5 cm. Leaves in 
5-7-merous pseudowhorls at the upper 1-2 nodes, 
unequal, obovate or obovate-elliptic, also oblong- 
obovate, apex obtuse or 4. rounded, not rarely 
somewhat notched, base cuncalc and a little 
decurrent into petiole, coriaceous, glabrous above 
at maturity, laxly and I: persistently lepidote 
beneath (scales small, marginal zone irregularly 
lobed or dented, dark centre impressed), margin 
:i: flat. (2.5-)3.5-7.5( 8.5) by (1.5-)2-3(-4) cm, 
midrib narrowly impressed above, nearly as 
robust as the petiole and obtusely prominent at 
the base beneath, nerves 4-6 pairs, spreading, ± 
distinctly impressed above and rai.sed beneath, 
not rarely subinconspicuous on both laces as is 
the reticulation; petiole somewhat flattened, 
grooved above, lepidote initially, 4-7(10) by c. 
1.5 mm. Umbels (I )2 3( 5)-flowered. Flower 
buds ovoid; outer perulae suborbicular. inner ones 
ovate to oblong-obovate. obtuse, grey-sericeous 
or appresscdly shorl-tomcntose in ihe upper (free) 
part, cilialc, up to 0.8 by 0.5 cm. Pedicels rather 



648 



Flora Malesiana 



[ser. I, vol. 6* 



slender, densely substellate-lepidote, epilose, 1-1.8 
cm. Calyx obliquely disk-shaped, very shortly 
obtusely lobed or dented, 4-5 mm 0. Corolla 
broadly funnel-shaped, ±_ tubular below, scarlet 
to purple or orange red, sometimes salmon, 
2.3-3 cm long in ail, laxly substellate-lepidote 
outside, glabrous inside, tube straight, 1-1.4 cm, 
base 3-5 mm 0, gradually widened to c. 1 cm at 
apex, lobes spreading, obovate, (1. 3-) 1.6-1. 8 by 
1-1.3 cm. Stamens 10, subequal, 1.5-1.7 cm; 
filaments linear, dilated and patently pubescent 
for 2-4 mm at base, glabrous and filiform for the 
rest; anthers obovate-oblong, base subobtuse, 
2-2.5 by 1 mm. Disk glabrous, not prominent. 
Ovary ovoid-conical, i patently villous, 4.5 by 
3-3.5 mm; style very abrupt, terete, glabrous, 
0.8-1 cm; stigma thick-obconical. Capsule sub- 
cylindric, shortly attenuate at both ends, sub- 
densely set with minute scales and short, rather 
robust and subpatent hairs, c. 2 cm by 4-5 mm; 
pedicel up to 3 cm. 

Distr. Malesia: N. Sumatra (Atjeh, Tapanuli, 
East Coast). 

Ecol. Mostly terrestrial in light forests, summit 
vegetation, cricoid ridge scrub, alpine growth 
with ferns or grassland, occasionally epiphytic, 
rather common locally, 1800-2700 m. Fl. Jan. -Dec. 

Vern. Kaju hor hor dolok, Asahan-Toba 
boundary. 

261. Rhododendron arfakianum Becc. Malesia 1 
(1878) 201; J.J.S. Nova Guinea 12 (1914) 134, 
in text Qarfakense'); Sleum. Reinwardtia 5 
(1960) 218; Blumea 12 (1963) 113.—/?. undulati- 
calyx J.J.S. in Gibbs, Arfak (1917) 168. 

Epiphytic, more rarely terrestrial shrub, (0.5-) 
l-2(-2.5) m. Branchlets 2-3 mm 0, sparsely 
lepidote and papillose-pubescent at the upper 
l(-2) internodes, glabrescent; internodes 2-10 cm. 
Leaves in (3-)4-5-merous pseudowhorls at the 
upper 1-3 nodes, oblong-obovate to obovate- 
elliptic, apex obtusely attenuate or nearly rounded, 
base ± cuneate to the petiole, coriaceous, glabrous 
above at full age, subpersistently and subdensely 
to rather laxly lepidote beneath (scales small, 
marginal zone irregularly dented, dark centre 
minute, slightly impressed, the scale surrounded by 
a dark halo), moreover very finely punctulate by 
minute emergences of the epidermis beneath, 
slightly revolute at the margin in herb., (4-)5-8 
(-10) by (1.5-)2-3.5 cm, midrib and petiole 
slightly depressed and with minute, papillose, 
spreading hairs above, as robust and wrinkled as 
the petiole and obtusely prominent in the lower 
part beneath, nerves 6-8 (-10) pairs, spreading, 
± obscurely inarched at margin, slightly promi- 
nent on both faces, reticulation inconspicuous; 
petiole 6-10(-12) by 1-1.5 mm. Umbels (4-)5-7 
(-lO)-flowered. Outer perulae ovate, obtuse, 
inner ones oblong-obovate, laxly lepidote and 
minutely papillose dorsally, finally practically 
glabrous, fimbriate, up to 1 .5 by 0.8 cm. Bracteoles 
filiform below, spathulate-dilated distally, up to 
2.3 cm by 0.5-4 mm, papillate. Pedicels slender, 
laxly to subdensely lepidote and finely patent- 



puberulous, 0.8-1.2 cm. Calyx disk shaped, 
membranous, subobtusely and very shortly 
unequally dented, wavy, spreading to reflexed, 
glabrous dorsally, manifestly ciliate, c. 3 mm 0. 
Corolla tubular below, expanded towards the 
limb, oblique, (2.5-)3(-3.5) cm long in all, ±. 
deeply pink, glabrous outside, laxly pubescent at 
the tube inside, tube subcylindric, straight or 
slightly curved, 1.5-2 cm, base 3-4 mm 0, widened 
to c. 5 mm at apex, lobes suberect, or somewhat 
spreading, obovate-spathulate, 0.7-1 by 0.5-0.8 
cm. Stamens 10, unequal, longest ± as long as 
corolla; filaments linear and laxly pubescent below, 
filiform and glabrous upwards; anthers obovate, 
2 mm. Disk puberulous. Ovary subcylindric, 
densely short-pubescent and minutely lepidote 
(scales covered by the hairs), c. 4 by 2 mm, 
rather abrupt with the style, the latter slender, 
sometimes with some hairs at the very base, i 
equalling the stamens; stigma subclavate-obconi- 
cal, oblique. Submature capsule fusiform, densely 
hairy, 2 by 0.4 cm, crowned by the glabrous 2 
cm long style. 

Distr. Malesia: NW. New Guinea (Arfak and 
Nettoti Mts). 

Ecol. In edge of forest, 1200-2135 m, locally 
common, Fl. Jan. -Dec. 

Vern. Krobiia, Kebar. 

262. Rhododendron fuchsii Sleum. Blumea 12 
(1964) 341. 

Shrub or small tree, 2-4 m. Branchlets robust, 
reddish and shining when dry, laxly lepidote at 
apex; internodes 4-8 cm by 3-4 mm. Leaves in 
(5-)6-8-merous pseudowhorls at the 1-2 upper 
nodes, elliptic-oblong or more rarely oblong, 
apex shortly attenuate and subobtuse, base 
almost rounded to truncate, coriaceous, stiff, 
glabrous above at age, subdensely lepidote 
beneath (scales brownish, flatfish, narrow marginal 
part variously angled or substellately dented, 
centre sunk, leaving dark pits), ± flat, 5-8.5 by 
(2-)2.5-4(-4.5) cm, midrib as thick as the petiole 
below, narrowly impressed above, obtusely 
prominent beneath, nerves 6-8 (-10) irregular 
pairs, rather straight in the lower, more curved 
in the upper part and but obscurely inarching, 
slightly though markedly impressed above, often 
rather indistinct on both sides, petiole thick, 
flattened, (2-)3-5 by 3-4 mm. Umbels (6-)8-10 
(-ll)-flowered.. Outer perulae ovate, apiculate, c. 

2 by 1.5 cm, inner ones spathulate and up to 

3 by 1-1.5 cm, glabrous, rather shining dorsally. 
Bracteoles subspathulate-linear, glabrous, 3-3.5 
cm. Pedicels stoutish, (2-)2.5-3.5 cm, laxly 
patently hairy and laxly lepidote, or more densely 
lepidote only. Calyx oblique, shortly 5-lobed and 
wavy, c. 4 mm 0. Corolla funnel-shaped, salmon- 
pink in bud, pinkish red at anthesis, throat 
yellowish, (very) laxly lepidote outside, hairy at 
the base only inside, (4.5-)5-5.5(-6) cm long in 
all, limb 4(-5) cm 0, tube 2.5-3.5 cm, base 5- 
saccate and 5-6 mm 0, widened to 1 cm at apex, 
lobes spathulate-obovate, slightly refuse, 2-2.5 
by 1.5-2 cm. Stamens 10, ± as long as the corolla 



Jan. 1966] 



Ericaceae (Sleumer) 



649 



tube; filaments linear, hairy but for the lower 
Vi or Yi', anthers oblong, 4 by 1 mm, each cell 
with a basal thickened apiculus. Disk thickly 
10-lobed, glabrous. Ovary conical-cylindric, 5- 
angular or -ribbed, abrupt with the style, densely 
patently hairy and lepidote, 4-5 by 2.5-3 mm; 
style thickish, glabrous, c. 1.4 cm; stigma sub- 
capitate. Capsule thick-cylindrical, 1.2 by 0.6 cm; 
pedicel thick, glabrous, up to 3.5 cm. 

Distr. Malesia: Borneo (N. slope of Mt 
Kinabalu). 

Ecol. Apparently rare, in sclerophyllous 
vegetation or forest border at c. 3350 m. Fl. fr. 
Aug. 

263. Rhododendron impositum J.J.S. in Fedde, 
Rep. 68 (1937) 201; Sleum. Reinwardtia 5 
(1960) 218. 

Shrub or treelet, 3-4 m. Branchlets subterete, 
(2-)3-4 mm 0, young parts laxly set with flattish 
scales, glabrescent. Leaves several (up to 12) 
scattered along the upper internodes (7-18 cm), 
elliptic or mostly obovate-elliptic to obovate, 
sometimes subovate-elliptic, apex generally shortly 
and rather abruptly acutely acuminate, rarely 
obtuse, base broad-cuneate to rounded, sometimes 
subcordate, coriaceous, initially very densely 
lepidote on both faces (scales touching each 
other), glabrous and remaining minutely pitted 
above, densely lepidote beneath (scales only 
partly or not touching each other, dilutely rufes- 
cent, small, marginal zone variously angled or 
dented, minute centre slightly impressed), edge 
finely crenulate by impressed scales (lens!), 
margin revolute to various degree, (3.7-)4-8(-10) 
by (1.8-)2.5-4(-5) cm, midrib nearly as stout as 
the petiole and much convex-prominent in its 
lower part, broadened and rather flat, if not 
impressed in its lower part beneath, nerves 6-9 
pairs, at an angle of c. 45' and rather high-ascend- 
ing, curved-anastomosing before the edge, slightly 
impressed above, a little raised beneath, reticula- 
tion mostly obsolete; petiole robust, rugulose, 
lepidote initially, compressed dorsally or distinctly 
flattened in the lower leaves, (4-)6-10 by (1. 5-) 
2-4 mm. Umbels (6-)IO-15-flowered. Bracteoles 
filiform to linear, almost hirsute, up to 2 cm by 
0.5-1.5 mm. Pedicels densely (in part caducously) 
stellate-lepidote, 2-3.8 cm by c. 0.8 mm. Calyx 
obliquely disk-shaped, normally shortly (r. 1 mm) 
dented (or not rarely one to two teeth elongate 
up to 5 by I mm), teeth rcflexed, lepidote dorsally, 
glabrous inside, margin with caducous scales, c. 
4 mm 0. Corolla funnel-shaped, white, pink or 
red, 5-lobcd : halfway, (3.5)4-5 cm long in all, 
sparsely lepidote outside, pubcrulous in the lower 
part of the tube inside, tube straight, fl.5-)2 cm, 
base 4-5 mm 0, - pouched, gradually dilated to c. 
1 cm at apex, lobes broadly obovate-spalhulate, 
slightly rctusc. 2-2.5(-3) by 1.5 2 cm. Stamens 10, 
a little cx.sertcd; filaments linear and laxly hairy 
below, narrower to filiform and glabrous upwards; 
anthers oblong. 3-3.5 by I mm, base obtuse or 
minutely apiculatc. Disk glabrous, or pubescent 
at upper margin. Ovary ovoid-conical or somewhat 



cylindric-obovoid, abrupt with the style, brownish 
tomentose (hairs covering the scales), 5-6(-7) by 
2.5-3 mm; style rather slender, with some sparse 
hairs at the very base or glabrous, nearly equalling 
the stamens; stigma capitate, deeply lobed. 

Distr. Malesia: SW. Central Celebes (Latimod- 
jong Range). 

Ecol. In low forest, 2400-3000 m. Fl. June. 

264. Rhododendron leytense Merr. Philip. J. Sc. 
10 (1915) Bot. 55; En. Philip. 3 (1923) 244; 
COPEL. /. Philip. J. Sc. 40 (1929) 165, t. 11, f. 
7-9; Sleum. Reinwardtia 5 (1960) 218.—/?. 
flaviflorum Elm. ex Merr. En. Philip. 3 (1923) 
244 /;; obs. pr. R. leytense. 

var. leytense. 

Epiphytic shrub. Branchlets terete, reddish 
brown or greyish, 2-3 (-4) mm 0, laxly lepidote 
at the upper internodes, glabrescent lower; 
internodes 1.5-5 cm. Leaves in 3-5-merous 
pseudo whorls at the upper 1-3 nodes, oblong or 
elliptic-oblong, apex shortly and ± bluntly 
acuminate, base attenuate to somewhat decurrent 
into petiole, thin coriaceous, laxly substellate- 
lepidote on both faces when young, finally glabrous 
above, laxly and subpersistently lepidote beneath 
(scales small, the marginal zone irregularly lobed 
or dented, centre shallowly impressed), finally ± 
glabrescent beneath too, leaving dark points, 
flat, 4-7 (-8) by (1.5-)2-2.7 cm, midrib slightly 
prominent in the lower, ± impressed in the upper 
half above, broadened and obtusely prominent 
below, nerves c. 8 pairs, rather straight-ascending, 
subinconspicuous on both sides, reticulation 0; 
petiole 0.6-1.8 cm by 1.5 mm, flattened. Umbels 
4-5-flowered. Perulae oblong, acuminate, glabrous, 
up to 2.5 by 1 cm. Bracteoles linear. Pedicels 
puberulous, practically elepidote, stoutish, c. 1 
cm, twice as long in young fruit. Ca/v.v obsolete, 
rim-like, c. 3 mm 0. Corolla broadly tubular- 
funnel-shaped or -subcampanulate, yellow, 3.7-4 
cm long in all, 5-lobed about halfway, glabrous 
outside, very laxly hairy at the lower half of the 
tube inside, tube c. 2 cm, at base 5-6 mm 0, 
gradually widened to c. 1.5 cm at apex, lobes 
obovate, 1.5-1.8 by 1.2-1.4 cm. Stamens 10, 
unequal, 2.7-3 cm; filaments dilated and patently 
hairy in the lower third, more linear to filiform 
and glabrous above, anthers oblong. 2-2.5 mm, 
base minutely or not biappendiculatc. Disk 
pubescent at the upper margin. Ovary subcyUndTic, 
tapering to style, rather densely short-pubescent 
and with minute scales : hidden by the hairs, c. 
7 by 2 mm; style hairy and lepidote in the lower 
%, glabrous for the rest; stigma obconical, 
deeply 5-lobcd. 

Distr. Malesia: Philippines (Luzon, Leyte). 

Ecol. Mossy forest, probably above 1000 m. 

Vcrn. Maldgos, Leyte. 

var. loheri (Copel. /.) Slium. Reinwardtia 5 
(I960) 218.—/?. loheri Cofti.. f. Philip. J. Sc. 40 
(1929) 166. t. II. f. 10-12. 

Differs from var. leytense by the corolla only c. 



650 



Flora Malesiana 



[ser. I, vol. 6* 



3 cm long in all and anthers biappendiculate at 
the base. 

Distr. Ma/e.s/a: Philippines (Luzon), once found. 

Vern. Salangon. 

265. Rhododendron brachygynum Copel. /. Philip. 
J. Sc. 40 (1929) 137, 165, t. 11, f. 4-6; Sleum. 
Reinwardtia 5 (1960) 219. 

"Shrub 1 m. Branchlets red brown. Leaves 
either in pseudowhorls at the end of the branchlets, 
or scattered and alternate, oval, apex short- 
acuminate, base obtuse, decurrent into the petiole, 
shining and minutely blackish punctulate above, 
sparsely set with pale scales beneath, c. 10 by 4.5 
cm; petiole 1.5 cm. Umbels rr 5-flowered. Pedicels 
robust, pubescent, c. 2 cm. Corolla narrowly 
obconicaj, yellow, 4 cm long in all, lobes rounded, 
c. 1.5 cm. Stamens 10; filaments minutely pubes- 
cent in the lower part, c. 3 cm; anthers oblong, 
base minutely biappendiculate, 3 mm. Disk hairy, 
in the upper margin. Ovary subcyiindric, densely 
hairy, c. 8 by 3 mm; style subabrupt, glabrous, 
c. 9 mm; stigma subcapitate, 2 mm 0." 

Distr. Malesia: Philippines (Cebu), once 
found. 

Vern. Mogdng-piidpiid. 

Note. No material seen; description after 

COPELAND /. 

266. Rhododendron mindanaense Merr. Publ. Gov. 
Lab. Philip, n. 29 (1905) 41; Philip. J. Sc. 3 (1908) 
Bot. 381; Elm. Leafl. Philip. Bot. 3 (1911) 1106; 
MiLLAis, Rhodod. (1917) 210; Merr. En. Philip. 
3 (1923) 244; Millais, Rhodod. ser. 2 (1924) 188; 
Copel./. Philip. J. Sc. 40(1929) 164, t. 7, f. 10-11, 
t. 11, f. 3; Sleum. Reinwardtia 5 (1960) 219. 

Erect shrub, about 1 m; stems few to several, 
sparingly branched. Branchlets terete, reddish and 
sparingly lepidote at the upper internodes, 
glabrescent and often greyish or pruinose below, 
4—5 mm 0; internodes (2-)5-7 cm. Leaves in 
5-7(-9)-merous pseudowhorls at the upper 1-2 
nodes, oblong-obovate, apex obtuse or mostly 
rounded, not rarely slightly emarginate, narrowed 
to the cuneate base, coriaceous, i rigid, dark 
green above, paler beneath, glabrous above at 
full age, more persistently and laxly lepidote 
beneath (scales rather small, marginal zone 
irregularly lobed or dented, minute dark centre 
slightly impressed), finally glabrescent beneath with 
persistent dark points, margin ~ flat, 5-7(-8.5) 
by (2-)2.5-3.5(-4.5) cm, midrib broadened below, 
slightly grooved above at least in the upper ^4, 
rather flat and but obtusely raised beneath, nerves 
8-10 pairs, at an acute angle, rather high-ascending, 
anastomosing before the edge, slightly but distinct- 
ly impressed above and raised beneath as is the 
± marked lax reticulation; petiole flattened, 
lepidote for a long time, 6-10 by 2-3 mm. Umbels 
(7-)9-12-flowered. Outer perulae ovate, inner ones 
ovate-oblong, obtuse, subchartaceous, glabrous 
and shining dorsally, up to 3.5 by 1.5 cm. 
Bracteoles linear below, subspathulate at the 
apex, glabrous, up to 4 cm by 1-3 mm. Pedicels 
laxly lepidote, 2-2.5 cm by 1 mm. Calyx obliquely 



patellar, shortly obtusely 5-lobed, 3.5 mm 0. 
Corolla tubular below, funnelform-expanded 
above, pure white, very laxly lepidote outside, 
laxly pubescent in the lower 24 of the tube inside, 
4—4.5 cm long in all, tube cylindric, c. 2.5 cm, 
base pouched, c. 5 mm 0, somewhat expanded 
distally, lobes obovate, 2-2.2 by 1.5-1.8 cm. 
Stamens 10, subequal, 2.5-3 cm; filaments 
linear, laxly hairy in the lower third, glabrous 
above; anthers subobovate-oblong, 3.5-4 by 1 mm, 
base minutely biappendaged. Disk slightly promi- 
nent, hairy at the upper margin. Ovary subovoid- 
cylindric, densely yellowish-brown-pubescent, the 
hairs covering numerous, minute scales, 6-7 by 
2.5 mm, tapering to the style, the latter sparsely 
hairy in its lower ^U-^la, ± as long as the stamens; 
stigma subcapitate. Capsule fusiform, slightly 
curved, warty, subdensely hairy and lepidote, 
3-3.5 by c. 0.7 cm, with 5 deep longitudinal 
grooves; pedicel 5 cm. 

Distr. Malesia: Philippines (Mindanao: Mt 
Apo). 

Ecol.Mossy forest near summit, 2700-2895 m. 
Fl. July, Aug. 

267. Rhododendron renschianum Sleum. Bot. 
Jahrb. 71 (1940) 146; Reinwardtia 5 (1960) 220. 
— R.javanicum [non (Bl.) Benn.] Malm in Fedde, 
Rep. 34 (1934) 284.— Fig. 45. 

Shrub, c. 0.5 m. Branchlets 1.5-2(-3) mm 0, 
tips caducously lepidote, older parts glabrous; 
internodes 3-4 cm. Leaves 4-6 spirally arranged 
in the upper third of the upper 2-4 internodes, 
2 or 3 of these leaves more close together in a 
kind of pseudowhorl, elliptic or mostly elliptic- 
or obovate-oblong, apex shortly acuminate, sub- 
acute or obtuse, sometimes rounded, base ± 
cuneate to the petiole, coriaceous, very early 
glabrescent above, subpersistently and subdensely 
lepidote beneath (scales small, marginal zone 
irregularly lobed or dented, centre distinctly 
impressed, blackish with age), slightly revolute, 
(3.5-)4.2-5 by 1.5-2.3 cm, midrib impressed above, 
strongly raised beneath, nerves 4-5 pairs, sprea- 
ding, obscurely anastomosing, slightly impressed 
above, mostly obsolete beneath, reticulation 0; 
petiole lepidote, semiterete, grooved above, 4-6 
by 1-1.5 mm. Umbels (1-3) 5-8-flowered. Perulae 
membranous, outer ones ovate-apiculate or 
-acuminate, inner ones ovate-oblong or obovate 
and obtuse, glabrous dorsally, margin with 
caducous scales, up to 1 by 0.6 cm. Bracteoles 
linear, subspathulate apically, up to 1.2 cm. 
Pedicels rather slender, laxly to subdensely sub- 
stellate-lepidote, c. 2 cm. Calyx obliquely disk- 
shaped, very shortly dented or lobed, lepidote 
dorsally, c. 3 mm 0. Corolla tubular-funnelform, 
3-3.5 cm long in all, orange to red, laxly lepidote 
outside, glabrous or practically so inside, tube 
cylindric, straight, (1.7-)2(-2.5) cm, base some- 
what 5-pouched, 4-5 mm 0, slightly dilated to 
6-7 mm at apex, lobes ± spreading, oblong- 
obovate to obovate, 1-1.5 by 0.8-1.3 cm. Stamens 
10, i as long as the corolla tube; filaments 
linear, hairy for 2 or 3 mm above the base, 



Jan. 1966] 



Ericaceae (Sleumer) 



651 




Fig. 45. Rhododendron renschianum Sleum. Mt 

Geli Mutu, E. Flores, 1500-1700 m (Rensch, 

1927). 

glabrous for the rest; anthers oblong, c. 1 mm. 
Disk prominent, glabrous. Ovary subcylindric, 
densely covered with very short, subpatent, 
greyish hairs and some (early caducous) scales, 
5-7(-9) by c. 1 mm, subabrupt or more continuous 
with the style, the latter with some hairs at the 
base, — as long as the stamens; stigma subcapitate, 
manifestly dilated. Capsule oblong, (l.2-)l.5-2 
cm by (4— )5-6 mm, subdensely short-pubescent, 
elepidotc; pedicel glabrescent, 2-3.5 cm by I mm. 

Distr. Malesia: Lesser Sunda Is. (Flores: Mts 
Geli Mutu and Mandaswai). 

Ecol. In Casuarina forest or in summit vegeta- 
tion, on slopes or edge of crater, on stony ground, 
1500-2300 m. Fl. May-Nov. 

268. Rhododendron culminicolum F. v. M. Trans. 
R. .Soc. Vict. n.s. 1, 2 (1889) 23; .Sleum. Rcinward- 
tia 5 (1960) 220; Blumea II (1961) \2ii; ihid. 12 
(1963) 113.—/?. nodosumC. H. Wrk.ht, Kcw Bull. 
(1899) 103.—/?. kevsseri Foerster in Fcddc, Rep. 
13 (1914) 223; .Sf mltr. Bot. Jahrb. 55 (1918) 155; 
DiEU, ihid. 62 (1929) 487; .Sleum. Rcinwardtia 5 
(1960) 220.—/?. convexum Sleum. I.e. 220.—/?. 
gregarium Sleum. I.e. 221. 

KEY TO THE VARIETIES 

1. Style hairy in the lower Vi-V* part. Filaments 
laxly pubescent to practically glabrous. 

2. var. nublcoia 



1. Style mostly with some sparse hairs at base, 
rarely so up to the lower third. Filaments 
laxly to subdensely hairy in about the lower 
half. 
2. Corolla 2.5-3.5 cm. Fruit 4(-5) mm o. 

3. var. angiense 

2. Corolla 3.5-4.5 (rarely -6) cm. Fruit (5-) 

6-8(-9) mm o 1. var. culminicolum 

1. var. culminicolum. 

Shrub or small tree, up to 5 m. Branchlets 2-3 
(rarely up to 6) mm 0, striate lengthwise, tips 
lepidote, below early glabrescent; internodes 2-5 
(-8, rarely up to 14) cm. Leaves in 3-5(rareIy up 
to 7)-merous pseudowhorls at the upper l-2(-3) 
nodes, elliptic or broadly oblong or obovate- 
oblong, more rarely ovate- or obovate-elliptic or 
obovate, apex shortly, sometimes rather abruptly 
subacuminate or attenuate, subacute or mostly 
obtuse, rarely almost rounded, base ± broadly 
attenuate to the petiole, or rarely rounded or 
even subcordate, i coriaceous, when very young 
densely covered on both faces with flattish, pale 
scales, completely glabrescent above with age, 
still laxly lepidote for a while beneath (scales 
rather small, early whitish or dissolved marginal 
zone irregularly lobed or dented, dark centre ± 
impressed), finally laxly brownish-biackish-punc- 
tate beneath, edge ± revolute, (2.5-)3.5-6(-8, 
rarely up to 9) by (l-)2-3(-4.5, rarely up to 7) 
cm, midrib narrowly sunk above, broadened, 
wrinkled, and much prominent in the lower part, 
nerves (4-)6-8(-10) pairs, curved-spreading and 
rather indistinctly anastomosing before the edge, 
slightly impressed above, somewhat raised beneath 
at full age, reticulation subdense, — obscure above, 
slightly raised beneath; petiole flattened, grooved 
above, (2-)4-7(-9, rarely up to 16) by (1-) 
1.5-2(-3) mm. Umbels (l-2-)4-6(rarely up to 9) 
-flowered. Outer peruiae suborbiculate to ovate, 
sometimes with a short narrow acumen, inner 
ones ovate to obovate, laxly lepidote and some- 
times subsericeous at the base outside, glabrous 
or mostly laxly to subdensely sericeous inside, up 
to 2 (rarely up to 2.5) by 1(-1.5) cm. Bracteoles 
filiform to linear, glabrous, up to 2 by 0.3 cm. 
Pedicels laxly to subdensely lepidote, moreover 
whether or (mostly) not hairy, (l-)1.5-2(-2.4) cm 
by c. 0.7 mm. Calyx obliquely disk-shaped, wavy, 
obtusely and shortly lobed, c. 4 mm 0, lepidote 
outside, generally not ciliate. Corolla ! obliquely 
tubular, red to purple, sometimes reddish pink, 
3.5-4.5 (rarely up to 6) cm long in all, laxly to 
subdensely lepidote outside, sparsely to subdensely 
hairy in the lower part of the tube inside, tube 
} curved, 2.5-3.5 (rarely up to 4.5) cm, base 
4-5(-6) mm 0, gradually widened to 8-IO(-15) 
mm at apex, lobes broadly obovate to suborbicul- 
ar, subcrect or • spreading, (0.8-)l.5(-1.8) by 
0.8-1. 3(-l. 5) cm. Stamens 10, subcqual, ± 
equalling the corolla; filaments Imcar, laxly to 
subdensely hairy in about the lower half, glabrous 
above; anthers obovate to broad-oblong, base 
often minutely biappcndiculate. (2-)2.5-3(-3.5) 
by 1-1.5 mm. Disk _\_ prominent, short-pubescent 



652 



Flora Malesiana 



[ser. I, vol. 6* 



but at the upper margin. Ovary subcylindric, 
yellowish tomentose, and i densely lepidote 
(scales mostly covered by the hairs), subcontin- 
uous with the style, 5-7 by 2-3 mm; style slender, 
finally exserted, mostly with some sparse hairs and 
scales at the base, rarely so up to the lower 
third; stigma subcapitate. Capsule subcylindric, 
attenuate at both ends, laxly to very laxly hairy, 
more densely lepidote (2.5-)3-4 by (0.5-)0.6-0.8 
(-0.9) cm. 

Distr. Malesia: New Guinea (Main Range, 
from Mt Carstensz to the Southern and Western 
Highlands and in the Owen Stanley Range, also 
in the Mt Saruwaged-Rawlinson Range area). 

Ecol. Low mossy forest or subalpine forest- 
grassland edge with shrubberies, on more open 
slopes or summit vegetation, also found in alpine 
pasture, both in primary and secondary communi- 
ties, not rarely as part of the forest undergrowth, 
locally plentiful, (2400-)2800-3600(-4000) m. Fl. 
Jan. -Dec. 

Vern. Kibimune, Chimbu: Masul, kutengaigai, 
Chimbu: Goromagua, gagum, wumbemp, Wahgi: 
Minj, wambu, Hagen: Togoba, bit-bit, Dani. 

Note. R. culminicolum is conceived here in a 
broad sense comprising forms with coriaceous, 
though flexible, distinctly petioled leaves with ± 
attenuate base, and corollas 3.5-4.5 cm, found 
in the upper montane forest from on c. 2400 m, 
and forms with smaller, stiff-coriaceous, short- 
petioled or subsessile leaves with rounded or even 
subcordate bases, and corollas 4-6 cm, found in 
the uppermost patches of the mossy forest or 
forest fringe shrubberies or even in open alpine 
grassland between 3200 and 4000 m. 

It belongs to the small number of rather 
variable species which are distributed all over the 
mountainous part of New Guinea, such as are 
R. zoelleri, R. macgregoriae, and R. konori. 

2. var. nubicola (Wernh.) Sleum. Reinwardtia 5 

(1960) 221; Blumea 12 (1963) 113.—/?. coenenii 
J.J.S. Nova Guinea 12 (1914) 132, t. 30 A; ibid. 
12 (1917) 506; Sleum. Reinwardtia 5 (1960) 153, 
pro syn. sub R. hatamense. — R. nubicola Wernh. 
Trans. Linn. Soc. II, Bot. 9 (1916) 98.— /?. acro- 
cline Sleum. Reinwardtia 5 (1960) \5%.—R.hata- 
mense {non Becc.) Sleum. I.e. 153, p.p. 

Differs from var. culminicolum by the laxly 
pubescent to practically glabrous filaments and 
the style, which is hairy in its lower half to % 
part. Leaves 2-3.5(-4) by (l-)1.5-2(-2.5) cm. 

Distr. Malesia: W. New Guinea (Wissel Lakes, 
Mt Carstensz and Oranje Mts). 

Ecol. Open places in the subalpine forest or 
in open alpine vegetation, (2500-)3200-3600 m. 

3. var. angiense (J.J.S.) Sleum. Blumea 12 (1963) 
\\A.—R. angiense J.J.S. Nova Guinea 12 (1914) 
133, t. 30 B; in Gibbs, Arfak (1917) 169; Kaneh. 
& Hatus. Bot. Mag. Tokyo 56 (1942) 478; 
Sleum. Reinwardtia 5 (1960) 220; Blumea 11 

(1961) 128, pro syn. sub R. culminicolum F. v. M. 
var. culminicolum. — R. gibbsiae J.J.S. in Gibbs, 
Arfak (1917) 169; Sleum. Reinwardtia 5 (1960) 



153, pro syn. sub R. hatamense. 

Diff"ers from var. culminicolum by corollas 
2.5-3.5 cm long in all and smaller fruits, 4(-5) 
mm 0. 

Distr. Malesia: NW. Guinea (Vogelkop 
Peninsula: Arfak, Nettoti, and Tohkiri Mts). 

Ecol. In devastated Nothofagus forest or forest 
border and more open summit areas, (900-)1200- 
2300 m, locally rather common, but often sterile, 
generally terrestrial. Fl. fr. Jan. -Dec. 

269. Rhododendron mollianum Koord. Nova 
Guinea 8 (1909) 187; Sleum. Reinwardtia 5 
(1960) 222. 

Terrestrial shrub, 1 m. Branchlets terete, 2-4(-5) 
mm 0; internodes laxly minutely lepidote, lower 
ones glabrescent and corticate, 6-9(-15) cm. 
Leaves in 4— 5-merous pseudowhorls at the upper 
nodes, lanceolate or oblong-lanceolate, apex 
acutely caudate-acuminate, base broadly acutely 
attenuate, coriaceous, completely glabrous above 
with age, rather laxly lepidote for some time 
beneath (scales minute, marginal zone irregularly 
substellate-lobed, early dissolute, centre shallowly 
impressed), finally ± glabrescent beneath, margin 
strongly revolute, 11-15 by 1.5-3 cm, midrib 
narrow and impressed above the broadened base 
excepted, much and obtusely prominent beneath, 
nerves 10-12(-14) pairs, irregular, spreading, 
obsoletely anastomosing near the edge, slightly 
impressed above and raised beneath, reticulation 
obscure above, rather dense and finely prominent 
beneath; petiole rugose, grooved above, 5-7 by 
1-1.5 mm. Umbels ± 3-flowered. Pedicels laxly 
lepidote, epilose, 2.5-3 cm by 0.7 mm. Calyx 
disk-shaped, obsoletely dented, oblique, c. 3 mm 
0. Corolla funnel-shaped, orange, suff"used at the 
limb with red, 6.5-7 cm long in all, sparsely 
minutely lepidote outside, laxly hairy at the tube 
inside, tube c. 2.5 cm, base markedly pouched, c. 
5 mm 0, dilated to c. 1 cm at apex, lobes ± 
spreading, obovate-oblong, 4—4.5 by c. 2 cm. 
Stamens 10, subequal, reaching halfway the 
corolla lobes; filaments linear and subdensely to 
laxly hairy in the lower third, becoming nearly 
filiform and glabrous above; anthers elongate- 
oblong, slightly curved, 5 by 1 mm. Disk prominent, 
shortly greyish pubescent. Ovary subcylindric, 
tapering to the style, densely minutely lepidote and 
short-pubescent (not tomentose), scales well 
visible between the hairs, c. 8 by 2 mm; style in the 
lower y^-Vi short-pubescent and lepidote, or 
exclusively lepidote above, slender, i equalling 
the stamens; stigma dilated-capitate. 

Distr. Malesia: New Guinea (Noord R.), once 
found. Fl. June. 

270. Rhododendron poremense J.J.S. Bot. Jahrb. 
68 (1937) 203; Sleum. Reinwardtia 5 (1960) 222. 

Epiphytic shrub, c. 1 m. Branchlets 1.5-2.5 mm 
0, very laxly clothed with appressed scales, early 
glabrescent; internodes 4-9 cm. Leaves opposite 
or in 3(-6)-merous pseudowhorls at the upper 1-2 
nodes, obovate- to oblong-elliptic, apex very 
shortly acuminate or obtuse, base acutely attenu- 



Jan. 1966] 



Ericaceae (Sleumer) 



653 



ate, coriaceous, glabrous, densely blackish punc- 
tulate above at full age, laxly appressedly lepidote 
beneath (scales minute, narrow marginal zone 
irregularly lobed or obtusely dented, centre hard- 
ly impressed), finally — glabrescent beneath, 
margin very narrowly revolute, (6.5-)8-l 1 by 
2.9-4.6 cm, midrib flattened and slightly or not 
raised above, obtuse and much prominent at 
least in the lower half beneath, nerves 7 — 9 
pairs, rather narrow and subparallel, curved- 
ascending and obscurely anastomosing along the 
margin, slightly raised on both faces, reticulation 
— obsolete; petiole subterete, lepidote, grooved 
above, 7-9 by c. 2 mm. Umbels 5-6-flowered. 
Pedicels stoutish, rather densely substellate- 
iepidote, epilose, c. 1.5 cm. Calyx obliquely disk- 
shaped, crenulate or obtusely lobed, lepidote 
dorsally, c. 4 mm 0. Corolla funnel-shaped, 
carmine, c. 4 cm long in all, sparsely substellate- 
lepidote outside, finely hairy in the lower half of 
the tube inside, tube c. 2 cm, base 5-saccate, c. 
4 mm 0, gradually dilated to 6-8 mm at apex, 
lobes obovate-spathulate, often retuse, — ex- 
panded, 1.8-2 by 1.4-1.6 cm. Stamens 10, some- 
what exserted; filaments linear, rather laxly 
pubescent in the lower half, glabrous above; 
anthers oblong, base obtuse, c. 4 by 1 mm. 
Disk low, glabrous. Ovary subcylindric, attenuate 
towards the slender style, finely short-pubescent 
and subdensely set with minute, subentire scales, 
c. 5 by 2 mm; style glabrous, 1.7-2 cm; stigma 
thick-capitate. 

Distr. Malesia: SE. Celebes (Mengkoka Mts: 
B. Porema), at 1400 m. Once found. Fl. Oct. 

271. Rhododendron stresemannii J.J.S. in Fedde, 
Rep. 30 (1932) 166; Sleum. Reinwardtia 5 (1960) 
222. 

Shrub. Branchlets subterete, 2-3 mm 0, brown, 
subdensely clad with appressed, small scales at 
upper internodes (c. 10 cm), laxly so in the lower 
ones. Leaves in 3-5-merous pseudowhorls, ovate- 
or oblong-cordate, apex rather shortly acutely 
acuminate, base cordate with rounded lobes, 
glabrescent above with age, but still rather densely 
set with minute blackish pits for some time, laxly 
to subdensely lepidote beneath (scales small, 
marginal zone substellate-lobed or -dented to 
various degree, centre impressed), finally 4: 
glabrescent beneath too, leaving darker i pitted 
spots, flat. 8.5-10 by 4.2-5.3 cm, midrib flattish, 
dilated al the very base, narrower and slightly 
impressed for the upper third above, as robust 
as the petiole in its lower part, obtusely prominent, 
nerves 7-8 pairs, spreading, inarching before 
the edge, slightly impressed above and raised 
beneath, reticulation subinconspicuous; petiole 
compressed dorsally, lepidote. 5-6 by 2 mm. 
Umbels : 7-flowcrcd. Pedicels with lax substcllate. 
flat scales and minute white hairs, 2.4-2.7 cm by 
I mm. Calyx oblique, disk-shaped, obscurely 
lobed. rather fleshy, dorsally with some scales 
and fine short hairs, c. 4 mm 0. Corolla tubular- 
funncl-shapcd, slightly oblique, dark ro.sc-rcd, 
4-4.5 cm lung in all, quite glabrous outside, 



practically so inside, tube 2-2.3 cm, base dilated 
and 5-pouched, 5-6 mm 0, then ± narrowed, 
but upwards again dilated to c. 12-15 mm at 
apex, lobes obovate-spathulate, 2-2.3 by 1.5-1.8 
cm. Stamens 10, subequal, 2.4-2.8 cm; filaments 
linear, sparsely set with longish, ± spreading 
hairs in the lower third, glabrous above; anthers 
oblong, c. 3 by 1 mm. Disk low, hairy except 
the very base. Ovary subovoid-cylindric, densely 
clothed with white, ± forward directed hairs, 
which cover small scales, 6-7 by 3 mm, rather 
abrupt with the style, the latter stoutish, c. 2 cm, 
hairy at the base; stigma thick-capitate. 

Distr. Malesia: Moluccas (W. Ceram). 

Ecol. Steep limestone ridge of the Ora Mts, c. 
1000 m, in Rhododendron copse with a few high 
Casuarina. Once found. Fl. May. 

272. Rhododendron retivenium Sleum. Reinwardtia 
5 (1960) 222; Blumea 12 (1963) 114.—^. brook- 
eanum {nan Low ex Lindl.) Staff, Trans. Linn. 
Soc. II, Bot. 4 (1894) 196. 

Straggling shrub or small tree, up to 6 m, 
terrestrial or epiphytic. Branchlets terete, (3-) 
4-5 mm 0, rather laxly set with flat scales at the 
upper 1-2 internodes, the latter (2-)3-9(-13) cm. 
Leaves partly in 3-4-merous pseudowhorls at the 
upper 1-2 nodes, partly spiral and spaced in the 
upper half of the internode, lanceolate to lanceo- 
late-oblong, apex shortly subacutely acuminate, 
base cuneate to the petiole, not rarely subtruncate 
at the very base, coriaceous, glabrous and indis- 
tinctly impressedly blackish punctulate above at 
full age, rather laxly and subpersistently lepidote 
beneath (scales minute, marginal zone lobed or 
dented, early dissolute, blackish centre faintly 
impressed), margin ± flat, 14-18^-23) by (2-) 
2.5-4(-4.5) cm, midrib dilated in the basal part, 
very narrow and deeply sunk in the upper part 
above, as thick as the petiole and obtusely promin- 
ent in the lower part beneath, nerves 12-18 pairs, 
spreading, anastomosing along the edge, not 
much raised on both faces, reticulation dense, 
fine, but remarkably prominent on both sides; 
petiole somewhat flattened, wrinkled lengthwise, 
lepidote, early glabrescent, v6-)8-20 by 2-3(-4) 
mm. t/m6e/5 4-6(-8)-flowcred. Outer perulac ovate, 
more inner ones broadly ovate-oblong, innermost 
ones spalhulate, obtuse, membranous, all glabrous, 
up to 3 by 1.2 cm. Bractcoles linear, glabrous, c. 
2 cm by 1 mm. Pedicels robust, laxly lepidote, 
more densely so al both ends, not hairy at all, 
2-2.8 cm by 1-1.5 mm. Calyx obliquely disk- 
shaped, the margin wavy and indistinctly 5- 
lobulatc. glabrous. 4-5 mm 0. Corolla funnel- 
shaped in outline, tubular below, expanded up 
to 7.5 cm al the Imib. of a rich pure yellow or 
orange, sweet-scented. 6-7(-7.5) cm long in all, 
glabrous outside, laxly pubescent at the lower half 
of the tube inside, tube rather straight. (3-)3.5-4 
cm, base much saccate or inflated and 6-7 mm 
at base (which includes the calyx), slightly dilated 
upwards, l.5( 2) cm across the mouth, lobes 
broadly obovatc to suborbicular, 2.5-3 cm 0. 
Stamens 10. unequal, slightly exserted from the 



654 



Flora Malesiana 



[ser, I, vol. 6* 



corolla tube; filaments linear (1 mm) and densely 
patently hairy in the lower third, more filiform and 
glabrous upwards; anthers oblong, curved, base 
minutely apiculate, (3.5-)4.5-5 by c. 1 mm. 
Disk very thick and prominent, glabrous, or 
occasionally puberulous at the upper margin. 
Ovary cylindric, continuous with the style, 
seemingly glabrous, but laxly finely pubescent 
(lens!), otherwise set with some sparse scales, 
8-10 by c. 2 mm; style rather slender, nearly as 
long as the stamens, glabrous; stigma capitate, 
manifestly 5-lobed. Capsule subfusiform, very 
shortly attenuate at the base, more longish so at 
the apex, sparsely lepidote, puberulous in the 
lower third, or quite glabrous all over, (4.5-) 
5-5.5 by c. 0.8 cm; pedicel 2.5-3 cm by 1.5 mm. 

Distr. Malesla: Borneo (Mt Kinabalu). 

Ecol. In jungle ridge or crest, or on wet rocks 
in light shade, near waterfall, 1220-2985 m. Fl. 
Jan.-Dec. 

273. Rhododendron lowii Hook. /. in Hook. Ic. 

PI. (1852) t. 883 i! lower); Walp. Ann. 5 (1858) 
447; Staff, Trans. Linn. Soc. II, Bot. 4 (1894) 
195; GiBBS, J. Linn. Soc. Bot. 42 (1914) 102; 
MiLLAis, Rhodod. (1917) 204; Merr. En. Born. 
(1921) 462; Sleum. Reinwardtia 5 (1960) 223; 
Blumea 12 (1963) 114, non Courtois, Mag. 
d'Horticult. 1 (1833) 105, n. 402, pro var. R. 
ponticum L.; Loudon, Gard. Mag. 11 (1835) 79, 
190. 

Spreading shrub or small tree, 3-7 (rarely up 
to 10) m, terrestrial or epiphytic. Branchlets (5-) 
6-10 mm 0, terete, practically glabrous; upper 
internodes 4-10 cm (or longer?). Leaves 4-6 
scattered along the upper 1-2 internodes, 2 or 3 
in a ± pronounced pseudowhorl, large, elliptic 
to subobovate- or oblong-elliptic, rarely obovate, 
apex broadly attenuate or subacuminate, obtuse, 
rarely really rounded, broadly attenuate towards 
the base (very base often i abruptly truncate), or 
even rounded or subcordate, stiffly coriaceous, 
initially very densely lepidote on both sides (the 
scales ± touching), finally quite glabrous above, 
but still blackish-impressed-punctulate, subdense- 
ly lepidote for a while beneath (scales small, 
marginal zone variously lobed or shortly dented, 
minute centre shallowly impressed), gradually 
glabrescent beneath too, the scales leaving a 
pale halo in the epidermis, flat, ( 1 2-) 1 4-23 (-26) 
by (5-)6-13(-15) cm, midrib wrinkled lengthwise, 
as stout as the petiole in the basal part, much 
convex-prominent above, much less so to nearly 
flat beneath, nerves (10-) 16-24 pairs, spreading 
below, often markedly subparallel, in part 
deeply forked, indistinctly anastomosing, ± 
prominent on both faces, reticulation rather dense, 
mostly finely raised on both sides; petiole flattened, 
(3-)5-10 (rarely up to 15) by 3-5(-7) mm, 
rugose, inserted on a thick leaf-cushion. Umbels 
(6-)7-12-flowered. Outer perulae ovate, obtuse, 
inner ones ovate-oblong to oblong, short-acumin- 
ate, innermost ones obovate, obtuse, all stiff- 
membranous, glabrous and lustrous dorsally, up 
to 3.5 by 1 cm. Bracteoles linear below, sub- 



spathulate-dilated distally, up to 3 cm by 2 mm. 
Pedicels laxly substellate-lepidote, epilose, 2-3 
(rarely up to 5) cm by 1.5 mm. Calyx very oblique, 
rim-like or shortly disk-shaped, obscurely 5-lobed 
or -angled, fleshy, subglabrous dorsally, 5-6 mm 0. 
Corolla funnel-shaped (wax-like when fresh), 
canary or golden yellow, orange, salmon red or 
flushed flame-pink, (5.5-)6-7 cm long in all, 
limb as wide as the length of the corolla, scentless, 
glabrous outside, short-pubescent at the tube inside, 
tube straight, ± 3 cm, base markedly 5-saccate, 5- 
7 mm 0, gradually widened to 1.5-2 cm at apex, 
lobes much expanded, broad-obovate to -spathu- 
late, sometimes retuse, (2.5-)3-3.5(-4) by 2-3 cm. 
Stamens 10, unequal, the longest reaching halfway 
the corolla lobes; filaments linear (2 mm) and 
villous in the lower third, narrower to subfiliform 
and glabrous above; anthers obovate-oblong, 
curved, with minute basal appendages, c. 5 by 1.5 
mm. Disk thick, very prominent, glabrous or 
puberulous in the upper margin. Ova/->' columnar, 
± abrupt with the style, greyish velutinous and ± 
densely lepidote (most scales hidden by the rather 
short dense hairs), 7-10 by c. 4 mm; style stoutish, 
glabrous, or with some hairs at the base, (2-)3 
cm by 1 mm; stigma large, capitate, c. 4 mm 0, 
obtusely lobed. Capsule thick-fusiform, (2.5-) 
3.5-4(-4.5) by 0.8-1 cm, short-hairy, practically 
elepidote, finally ± glabrescent, with scattered, 
minute, elliptic-oblong, lenticel-like warts, valves 
woody; pedicel 4-5(-7) by 2-2.5 mm. Seeds c. 
6 mm, long-tailed on both ends, each tail c. 
2.5 mm. 

Distr. Malesia: Borneo (Mt Kinabalu and 
Mt Trus Madi). 

Ecol. Undershrub in high, small tree in low 
mossy forest, on hillside or rocky ridge, (1980-) 
2440-3350 m. Fl. Jan.-Dec. 

Note. This beautiful species is very conspicuous 
on Mt Kinabalu; the flower clusters are reported 
to measure up to 35 cm when fresh. 

274. Rhododendron intranervatum Sleum. Blumea 
11 (1961) 129; ibid. 12 (1963) 114. 

Epiphytic shrub. Branchlets c. 4 mm at the 
glabrous tips; internodes 1.5-3 cm. Leaves 
solitary and subopposite at the ultimate 1-2 nodes 
(leaving subreniform scars 5-6 by 7-9 mm), 
elliptic-obovate, apex rounded-obtuse, or some- 
times very shortly apiculate, broadly subcuneate- 
rounded towards the base (very base subtruncate- 
obtuse), or faintly cordate, coriaceous, firm, ± 
glabrous at full age above, subpersistently 
lepidote beneath (scales minute, flat, marginal 
zone variously substellate-dented, centre small, 
hardly immersed), edge a little revolute, 13-16 
(-18) by 8-10 cm, midrib as wide as the petiole 
and slightly raised below on the upper surface, 
thick-prominent at the base beneath, nerves 12-14 
(-16) pairs, ~ at right angles, rather straight 
and ih parallel, with numerous similar less distinct 
nerves (or primary veins) between, all + mani- 
festly prominent on both sides, anastomosing and 
united into an inner intramarginal nerve, which 
is connected by numerous transverse veinlets 



Jan. 1966] 



Ericaceae (Sleumer) 



655 



with an outer, less distinct one, reticulation dense, 
slightly prominent on both faces, especially 
beneath; petiole compressed dorsally or sub- 
semiterete, glabrous, c. 10 by 5-6 by c. 4 mm. 
Flowers in twos. Inner perulae oblong, glabrous, 
4-5 by c. 1 cm. Bracteoles (or innermost perulae?) 
elongate-subspathulate, c. 4 by 0.5 cm. Pedicels 
very densely patently short-hairy, elepidote, c. 4 
cm by 1.5 mm. Calyx disk-shaped. wa\-y, obtusely 
5-lobed, c. 4 mm o. glabrous lobes sometimes in 
form of 5 elongate obtuse laciniae, appressed to 
the corolla, 4—4.5 by 0.2-0.4 cm. Corolla funnel- 
shaped, light yellow, very shortly hairy for the 
lowest cm inside, quite glabrous otherwise, 5- 
lobed halfway, 5-6 cm long in all, base slightly 
5-saccate, 0.6-0.8 cm o, widened to 1.5-2.5 cm 
at ape.x, lobes obovate-spathulate, erecto-patent, 
2.5-3 by 2-3 cm. Stamens 10, c. 3 cm; filaments 
linear and densely hairy or almost villous in the 
lower half, filiform and glabrous above; anthers 
narrow-oblong, slightly curved, 7-8 by 1.5 mm, 
each cell bearing a basal obtuse swelling or gland. 
Disk prominent, thickly 10-lobed, glabrous below, 
laxly hairy above. Ovary subcylindric, c. 6 by 
2 mm, densely white-hairy or subvillous, appa- 
rently elepidote, tapering to the style, the latter 
rather slender, quite glabrous, 1-1.5 cm at 
anthesis; stigma thick-turbinate with 5 distinct 
lobes. 

Distr. Malesia: Borneo (Mt Palimasan, near 
Tabang on Belajan R. in W. Kutei; Mt Penrissen 
and Mt Berumput in Sarawak). 

Ecol. In Agathis forest on waterlogged, white, 
acid sands or on granite rock faces, 600-1050 m. 

275. Rhododendron toxopei. J.J.S. in Fedde, Rep. 
30 (1932) 168; Sleum. Reinwardtia 5 (1960) 224. 
Shrub. Branchlets subterete, 2-3 mm 0, initi- 
ally lepidote, early glabrescent; internodes 2-4 
cm. Leaves in — 4-merous pseudowhorls, oblong- 
ovate, apex gradually and r shortly acuminate, 
subacute, base obliquely rounded, slightly ine- 
quilateral, coriaceous, glabrous above (scale 
vestiges remaining recognizable by numerous 
minute, blackish pits), laxly subpersistently 
lepidote beneath (scales small, marginal zone 
variously and shortly dented or lobed, but later 
roundish when marginal zone is partly dissolved, 
centre slightly impressed), edge very narrowly 
or not revolute, 6.2-10 by 2-4.6 cm, midrib 
broadened below, wrinkled lengthwise, rather 
flat or a little impressed above, : distinctly 
prominent beneath, nerves 10-12 pairs, curved- 
spreading, inarching near the edge, well visible 
impressed above, slightly raised beneath, reticula- 
tion dense, finely prominent beneath only; petiole 
rugose, flattened, 7-1 1 by 2-2.5 mm. Umbels ■; 8- 
fluwcrcd. Bracteoles said to be linear, glabrous. 
Pedicels sparsely lepidote at both ends, 1.7-2.1 
cm by 1 mm. Calyx oblique, disk-shaped, obtusely 
angled or very shortly lobed, practically glabrous, 
3-4 mm 0. Corolla tubular-funnel-shaped, pale 
rose, 4.5-5 cm long in all, glabrous outside, laxly 
pubescent in the lower Vj of the tube inside, 
lube 2-2.4 cm, straight, base slightly dilated or 



pouched, 5-6 mm 0, widened to c. 1 cm at apex, 
lobes oblong-spathulate, ± spreading, 2.3-2.7 by 
1-1.6 cm. Stamens 10, shortly e.xserted; filaments 
linear below (1 mm) and subdensely patent- 
hairy in the lower, narrower and glabrous in 
the upper half; anthers oblong, curved, base 
obtuse, c. 4 by 1 mm. Disk prominent, short- 
pubescent. Ovary subcylindric. densely pubescent 
or subvelutinous. with numerous minute scales 
well visible between the hairs, rather continuous 
with the style, 6-8 by 2.5 mm; style rather slender, 
pubescent in the lower '/4-'/3, c. 2 cm; stigma 
capitate, c. 2.5 mm 0. 

Distr. Malesia: Moluccas (Buru, near Kuntu- 
run), in peatmoor with Sphagnum, 1075 m. Fl. 
Feb. 

276. Rhododendron brookeanum Low ex Lindl. 
J. Hort. Soc. Lond. 3 (1848) 82, c. icon p. 83; 
Planch. Fl. Serres 5 (1849) 477-480, c. icon, xyl.; 
Anon. Card. Chron. (1855) 404, c. icon.; Hook. 
Bot. Mag. (1856) t. 4935; Houtte, Fl. Serres II, 
2 (1857) 111, t. 1238-39; MiQ. Fl. Ind. Bat. 2 
(1859) 1057; Becc. Malesia 1 (1878) 207; 
Staff, Trans. Linn. Soc. II, Bot. 4 (1894) 196; 
GiBBS, J. Linn. Soc. Bot. 42 (1914) 103; Millais, 
Rhodod. (1917) 130; Merr. En. Born. (1921) 
460; HoLTT. M.A.H.A. Mag. 9 (1939) 35; Sleum. 
Reinwardtia 5 (1960) 224; Blumea 11 (1961) 130; 
ibid. 12 (1963) 114. — Azalea brookeana (Low ex 
Lindl. ) O. Ktze, Rev. Gen. PI. 2 (1891) 387. 

KEY TO THE VARIETIES 

1. Leaves oblong to lanceolate-oblong, ± shortly 

acuminate, (3-)3.5-8(-9) cm wide. 
2. Leaves without hairs . .1. var. brookeanum 

2. Leaves densely short-hairy on both sides 
along the midrib; branchlets hairy. 

2. var. cladotrichum 

1. Leaves lanceolate or oblong-lanceolate, longer 

and/or more acutish acuminate, mostly more 

slenderly. 6-15 by 1.5-2, rarely up to 3 mm 

petioled. 

3. Leaves lanceolate. Corolla (5-6) cm. 

3. var. gracile 
3. Leaves oblong-lanceolate. Corolla 4(-5) cm. 

4. var. cxtraneum 

1. var. brookeanum. 

Epiphytic or terrestrial shrub, up to 2(-4) m. 
Roots large and fleshy. Upper internodes ob- 
tusangular or subterete. laxly set with appressed 
substellatc scales, 2-15 cm by 3-4(-6) mm, lower 
ones terete, glabrescent. Leaves more or less clearly 
in 3 4( -5)-merous pseudowhorls at the upper 1-2 
nodes, oblong or lanceolate-oblong, often subses- 
sile, but sometimes also manifestly petioled, apex 
gradually and mostly rather shortly acuminate, 
subacute, base cuncate or attenuate to the petiole, 
or sometimes subtruncate. or even rounded at 
the very base, 1 coriaceous, glabrous, the former 
scales leaving dense, minute, dark pits, laxly to 
very laxly lepidote for some time beneath (scales 
small, marginal zone variously substcllatc-lobcd 



656 



Flora Malesiana 



[ser. I, vol. 6* 



or -dented, centre minute, shallowly impressed), 
finally ± completely glabrescent beneath, margin 
± flat, 12-25(-28) by (3-)3.5-8(-9) cm, midrib 
a prolongation of the robust petiole below, nerves 
in 14-24 pairs, ± spreading, inarching along the 
edge, slightly raised on both faces, reticulation 
rather coarse and not much conspicuous, some- 
times dense and more visible raised especially 
beneath (possibly due to intercrossing with distinct- 
ly reticulate-leafed species as R. retiveniuml). 
Umbels 5-14-flowered. Ferulae thin, outer ones 
ovate-subacuminate, inner ones oblong to spathu- 
late-oblong, glabrous, up to 4(-4.5) by 1.5(-2) cm. 
Pedicels sparsely lepidote or not, rarely almost 
glabrous, rarely pubescent, (2-)2.5-4(-5) cm by 
(l-)1.5-2 mm in anthesis. Calyx shortly disk- 
shaped or merely rim-like, obscurely angled or 
lobed, glabrous, 4—5 mm 0. Corolla funnel-shaped, 
beautiful orange pink with white or cream centre, 
or golden (pale) yellow, more rarely of a brick-red 
colour, throat then mostly yellow, said to become 
suffused with red or rosy orange outside with age, 
± fleshy, often with a delicate lemon-like frag- 
rance, glabrous outside, ± laxly pubescent in the 
lower part of the tube inside, 5-7.5(-8) cm long 
in all (length and width of corolla 9 cm when 
fresh), tube straight, 3-3.5 cm, markedly obtusan- 
gular, base 5-pouched, 6-8 mm 0, gradually dilated 
to 2-2.5 cm at apex, lobes much expanded, 
broadly obovate or suborbicular, slightly retuse 
or delicately crisped at the edge, (2.5-)3.5(-4.5) 
by 2.5-3(-3.5) cm. Stamens 10, subequal, reaching 
halfway the corolla lobes; filaments linear, 
densely subpatently hairy in the lower third, 
glabrous above; anthers oblong, curved, base 
obtuse, (4-)4.5-5(-6, rarely up to 7) by 1-1.3 mm. 
Disk prominent, puberulous at the upper margin. 
Ovary subcylindric, densely greyish hairy or 
velutinous and lepidote (scales often at least 
partly hidden by the hairs), ± abrupt with the 
style, 8-10 by 2-3 mm; style first shorter than, 
finally ± equalling the stamens, glabrous; stigma 
turbinate-dilated, deeply lobed. Capsule sub- 
fusiform-cylindric, lepidote and short-pubescent 
to various degree, with some elliptic minute 
warts, (3.5-)4-5 by 0.7-0.8(-1.2) cm; pedicel 
3-6 cm by c. 2 mm. Seeds 7-9 mm including the 
tails. 

Distr. Malesia: Borneo. 

Ecol. Epiphytic on mangrove or rocks at 
sea-level, ascending to 1525 m in the mossy 



forest, with preference for trees along rivers. Fl. 
Jan.-Dec. 

Note. Introduced to the lowlands of the Malay 
Peninsula and flowering freely. Formerly used for 
crossings, practically out of cultivation by now. 
Very near to R. triumphans Yersin & Chev. from 
S. Annam, which is mainly distinguished by longer 
and more slender petioles; its fruit is still unknown. 

2. var. cladotrichum Sleum. Reinwardtia 5 (1960) 
224. 

Branchlets and midrib of the leaves ± densely 
clad with fuscescent short and it spreading hairs. 
Pedicels ± glabrous, 6-10 by (l-)2(-3) mm. 

Distr. Malesia: Central E. Borneo (W. Kutei: 
Long Suh), at 300 m. Once found. 

3. var. gracUe (Low ex Lindl.) Henslow, J. R. 
Hort. Soc. 13 (1891) 261, f. 42 C/b/.), 43 a {fl.)', 
MiLLAis, Rhodod. (1917) 130; Sleum. Reinwardtia 
5 (1960) 224; Blumea 11 (1961) 130.—/?. gracile 
Low & ex Lindl. J. Hort. Soc. Lond. 3 (1848) 
84, fig.; MiLLAis, Rhodod. (1917) 174; Merr. 
En. Born. (1921) 461, non Becc. Malesia 1 (1878) 
203 (which is R. commutatum Sleum.). — R. 
obscurinervium Merr. J. Str. Br. R. As. Soc. n. 
76 (1917) 109; En. Born. (1921) 462. 

Leaves lanceolate, ± long-acuminate, rather 
slendery petioled, (l-)2-3(-3.5) cm wide only; 
nerves generally more obscure than in var. brooke- 
anum. Corolla 5 (-6) cm long in all. Branchlets 
and pedicels besides the scales whether or not 
patent-puberulous. 

Distr. Malesia: Borneo (Sarawak; W. and N. 
Borneo; Brunei). 

Ecol. Mostly epiphytic in lowland forest, 
sometimes terrestrial on vertical rocks, up to 
600 m. 

Note. Apparently transient to var. brookeanum. 

4. var. extraneum Sleum. Reinwardtia 5 (1960) 225. 
Branchlets laxly lepidote, epilose. Leaves 

oblong-lanceolate, 9-18 by (2-)2.5-5 cm; petiole 
7-12 by 2-3 mm. Pedicels densely short-pubescent 
and laxly lepidote, c. 1 5 by 1 mm. Corolla funnel- 
shaped, pale orange, 4(-5) cm long in all; other- 
wise as in var. brookeanum. 

Distr. Malesia: Sumatra (West Coast: Harau 
canyon near Pajakumbuh), on sandstone rocks, 
c. 500 m. Fl. Nov.-Dec. 



2. Subgenus Hymenanthes 

(Bl.) K. Koch, Dendrol. 2 (1872) 170 CHymenanthus'); Sleum. Blumea Suppl. 4 (1958) 51; 
Reinwardtia 5 (1960) 225. — Hymenanthes Bl. Bijdr. (1826) 862; G. Don, Gen. Syst. 3 (1834) 
849; D. DiETR. Synops. 3 (1843) 2; Copel./. Am. Midi. Nat. 30 (1943) eU. — Rhododendron 
subg. Eurhododendron (Endl. pr. sect.) Koehne, Deutsch. Dendrol. (1893) 449, 5. 5/r. ; Sleum. 
Bot. Jahrb. 74 (1949) 526, 542. — Fig. 47-^9. 

Plants entirely elepidote. Leaves evergreen. Flowers from terminal buds (several to many); 
leafy branchlets (new foliate shoots) from separate lateral buds, from the axils of the leaves of 
the past season. 

Distr. Europe, Asia, in Malesia: N. Sumatra, Malay Peninsula. 



A 



Jan. 1966] 



Ericaceae (Sleumer) 



657 



1. Section Hymenanthes 

(Bl.) Endl. Gen. (1839) 759; DC. Prod. 7 (1839) 721; Sleum. Blumea Suppl. 4 (1958) 51; 
Reinwardtia 5 (1960) 225. — Rhododendron sect. Eurhododendron Endl. Gen. (1839) 759, 
p.p. et aut. plurim.; DC. Prod. 7 (1839) 721, p.p. — Fig. 47-^9. 

Same as above. 

1. Subsection Irrorata 

Sleum. Dot. Jahrb. 74 (1949) 548; Blumea Suppl. 4 (1958) 52; Reinwardtia 5 (1960) 225. 

Leaves densely tomentose or villous by simple hairs initially (new shoots) or in young stages 
at least beneath, mostly glabrescent at maturity, margin ± cartilaginous characteristically 
undulated when dry; nerves 12-20, ± parallel. 

Distr. SE. Asia, in Malesia: N. Sumatra, Malay Peninsula. Fig. 46. 







Fig. 46. Distribution of Rhododendron subg. Hymenanthes subsect. Irrorata ( approximate 

southern limit of area on the Asiatic continent, • localities in Malesia) and of subg. Tsutsutsi sect. 
Tsutsutsi ( approximate southern limit of area in SE. Asia and Malesia). 

KEY TO THE SPECIES 

1 . Ovary densely set with stalked glands, or glandular and hairy (strigose). Leaves not wax-coated above, 

i.e. unaltered when heated. 
2. Style densely or subdensely glandular to the top. Leaves glabrous at maturity, laxly set with red 

glandular points underneath 277. E. atjchcnsc 

2. Style glabrous or practically so. Leaves clad underneath with a thin, whitish or fawn indumentum 

of interlaced cobweb hairs for a long time 278. R. wrayi 

1. Ovary not glandular at all, otherwise laxly to densely hairy. Leaves wax-coated above, i.e. becoming 
lustrous when heated 279. R. korthalsH 



277. Rhododendron atjchease Sleum. Blumea Suppl. 
4 (I95K) 53, //) clavi (angl.); Reinwardtia 5 (1960) 
226— Fig. 47-48. 

Large shrub or tree, 5-7 m; trunk c. 8 cm o. 
Branches and branchlcts robust, the latter terete, 
upper intcrnodcs densely and 1- persistently set 
with stalked glands; intcrnodcs 1.6-7 cm. Leaves 
in 5-9-mcrous pscudowhorls at or near the upper 
2 nodes, oblong-lanccolatc, apex short-acuminate, 
subacute, base attenuate to the petiole, thin- 
conaccous, though firm, brownish lanatc when 
very young, finally quite glabrous, but with lax. 



reddish glandular points beneath, edge slightly 
revolutc, 7.5-IO(-l3.5) by (2-)2.5-3.5(-5) cm. 
midrib impressed above, very prominent beneath, 
nerves c. 15 pairs, curved, not anastomosing, 
faintly impressed in fully developed leaves above, 
prominent beneath, reticulation dense, not much 
raised; petiole glandular initially, glabrescent, 
grooved above, 1-1.5 cm by 1.5-2 mm. Racemes 
umbel-like. 6-IO(-12)-flowcred. Flower buds 
oblong-ovoid, somewhat glutinous, c. 3 by 1-1.5 
cm. Outer pcrulae broadly ovate, apiculutc, 
ycllowish-or greyish-subsericcous, 0.5-1 by 0.6- 



658 



Flora Malesiana 



[ser. I, vol. 6* 




Fig. 47. Rhododendron atjehense Sleum. Mt Goh Lembuh, 3000 m, N. Sumatra (van Steenis 8991) 

(van Steenis, 1937). 



0.8 cm, inner (middle) ones ± obovate, dorsally 
with stalked, i appressed glands and ± tomentel- 
lous by rufous and yellowish hairs, 1-1.5 by c. 
1 cm, innermost ones obovate-spathulate, yellow- 
ish-tomentose or lanate dorsally, hairy fimbriate, 
up to 2 by 0.8 cm. Bracteoles linear, hairy, up 
to 2 cm by c. 1 mm. Rachis 0.5-2.5 cm, rather 
abruptly decrescent in 0, laxly glandular, with a 
tuft of pale hairs immediately below the insertion 
of each pedicel, these densely clad with spreading 
stalked glands, furthermore with rufous fine hairs 
at the base and immediately below the calyx, 
rarely all over the whole pedicel, 1-1.5 cm by c. 
1 mm. Calyx thick-membranous, wavy, deeply 
5-lobed, lobes ± spreading or erect, short- 
glandular dorsally, laxly to subdensely glandular- 
fimbriate and/or with longish substrigose hairs 
along the margin, 1.5-2.5 mm. Coro//a funnelform- 
campanulate, the attenuate base with 5 pouches, 
thin, glabrous, rose-lilac, (3-)3.5(^) cmlong in all, 
limb 3-3.5 cm 0, with 5 purplish nectar markings 



at base inside and darker spots at throat, lobes 
erecto-patent, broadly oblong to subobovate, 
slightly retuse, 1.5-2 by 1-1.5(-1.8) cm. Stamens 
10, very unequal, reaching to Vi-Vi of the corolla; 
filaments linear and laxly hairy below, filiform 
and glabrous above; anthers obovate, truncate, 
1.5-1.8 mm. Disk glabrous. Ova/-^ conical, obtuse, 
all over densely clad with stalked reddish glands, 
furthermore laxly to subdensely clad with sub- 
strigose rufescent stiff" hairs in the lower half, 
4-5 by 2-3 mm; style abrupt with the ovary, 
subdensely or densely clothed with stalked glands 
to the top; stigma subcapitate. Capsule cylindric, 
obtuse, nigrescent, glandular or strigose below, 
1.5-1.8 by 0.6-0.7 cm, valves 5, woody, with 
persistent calyx; pedicel 1.5-2 cm by 1 mm. 

Distr. Malesia: N. Sumatra (Gajolands: Mt 
Losir: Putjuk Angasan; Mt Kemiri; Mt Goh 
Lembuh). 

Ecol. In mossy forest or subalpine brushwood, 
1800-3000 m, locally common. Fl. Jan.-Feb. 



Jan. 1966] 



Ericaceae (Sleumer) 



659 




Fig. 48. Rhododendron atjehense Sleum., with 

white to light pink Exohasidium galls, on Mt Goh 

Lembuh, N. Sumatra (van Steenis, 1937). 



Note. Most related to R. ningyuenense Hand.- 
Mazz. {R. langbianense Chev. ex Dop) known 
from SW. China (SW. Szechuan-E. Yunnan) and 
Indochina (Annam), distinguished chiefly by much 
larger calyx lobes, a richer inflorescence, and rfc 
persistently glandular ultimate internodes. Not 
yet in cultivation, possibly one of the most 
hardy species of the genus in Malesia. 

278. Rhododendron wrayi K. & G. J. As. Soc. 
Beng. 74, ii (1905) 75; Ridl. J. Linn. Soc. Bot. 
38 (1908) 314, repr. J. Fed. Mai. St. Mus. 2 
(1908) 120; J. Fed. Mai. St. Mus. 4 (1909) 42. 
incl. var. elliptica Ridl. I.e.; ibid. 6 (1915) 49, 
incl. var. minor Ridl. I.e. 157; Fl. Mai. Pen. 2 
(1923) 217; Burk. & MoLrr. Card. Bull. S.S. 3 
(1923) 57; HfNDER.s. J. Mai. Br. R. As. Soc. 5 
(1927) 256; Symington, ibid. 14 (1936) 355; 
Hender.s. Mai. Nat. J. 6 (1950) 261; Sleum. 
Blumca Suppl. 4 (1958) 53, /// clavi; Rcinwardtia 
5 (1960) 221. -R. dubium K. & G. J. As. Soc. 
Bcng. 74. ii (1905) 78; Ridl. Fl. Mai. Pen. 2 
(1923) 221. -/f. corruscum Ridl. J. Fed. Mai. 
St. Mus. 5 (1914) 37; II. Mai. Pen. 2 (1923) 
217.- Fig, 49. 



Compact shrub or small, lanky tree, 3-6(-12) m. 
Branches and branchlets thick, terete, only tips 
pubescent; internodes 2-4(-8) cm. Leaves in 4-5- 
merous pseudowhorls at the end of the upper 
1-2 internodes; besides, much reduced, mostly 
early caducous leaves scattered along these inter- 
nodes; oblong to elliptic-oblong or elliptic, rarely 
somewhat obovate-elliptic, apex short-acuminate 
(very apex mostly subacute or rather obtuse), 
base broadly attenuate to rounded, very coriaceous, 
rusty-floccose-tomentose on both surfaces on 
young shoots, very early glabrous above, indumen- 
tum of undersurface persisting for a long time, 
thin, greyish, whitish or fawn at flowering time, 
± evanescent with age, leaving a very fine detersile 
greyish tomentum of cobweb hairs in the pits of 
the intervenium, finally not rarely quite glabrous, 
becoming silvery-greyish or rather glaucous under- 
neath, edge recurved in dry specimens, (4.5-) 
6-15(-18) by 2-5(-6) cm, midrib grooved above, 
bold beneath, nerves 10-14(-18) pairs, rather 
straight or slightly curved, at an angle of 70-90°, 
indistinctly anastomosing, ± not impressed above, 
prominent beneath, reticulation dense, obsolete 
above, more raised beneath and very distinct with 
age; petiole glabrous, (1.5-)2-2.5 cm by 2-3 mm, 
somewhat flattened, grooved above. Umbel-like 
racemes (3-)8-12-flowered. Outer perulae broad- 
ovate, dorsally clothed with very short hairs and 
minute subsessile glands, especially the lowest 
ones ± glutinous, up to 1.2 by 0.9 cm, inner ones 
larger, ± obovate-spathulate, ± apiculate, silky- 
tomentose on both faces, up to 2.5 by 1.5 cm. 
Bracteoles linear-spathulate, ± tomentose, up to 
10 by 1-1.5 mm. Rachis 0.5-1 cm, laxly floccose 
and glandular. Pedicels rather slender, ± densely 
rusty-floccose and with ± numerous small sub- 
stipitate glands, the floccae finally ± evanescent, 
the glands becoming more visible, 1.5-2 cm. 
Calyx mostly very oblique, small, wavy, rusty- 
floccose-tomentose outside, lobes obtuse, c. 1 mm. 
Coro//a rather wide-campanulate, (2. 5-)3-3.5(-4.5) 
cm long in all, somewhat translucent, flushed 
pink in bud, white in full bloom, whether or not 
tinged with pale pink and red-spotted inside, 
glabrous, lobes broad oblong to obovate-oblong, 
4: spreading, 1.2-1.7 by 0.7-1 cm. Stamens 10, 
4 unequal, mostly a little shorter than the corolla; 
filaments linear and densely patent-hairy in the 
lower third, filiform and glabrous distally; anthers 
oblong, orange-red, 3-3.5 by 1 mm. Disk glabrous 
below, densely whitish ciliate at the upper margin. 
Ovary conical. 3-4 by c. 2.5 mm. dense rusty 
tomcntulose, with some substipitate glands; style 
rather abrupt with (he ovary, showingsomc stalked 
club-shaped glands on its very base, otherwise 
completely glabrous, rather slender, (1.5-)2-2.5 
cm; stigma subcapitatc. Capsule woody, cylindric, 
obtuse, slightly ridged, initially rusty-lomcntose, 
finally glabrous, blackish when dry. (1)1.3-1.6 
(-2) by 0.5-0.6 (rarely up to 0.8) cm, on stoutish 
_h glabrous pedicel (1.5-)2-2.5(-3) cm; valves 5. 
Seeds angular, with very short irregular, often 
lacerate appendices, no tails, c. 2 mm. 

Distr. Malesia: Malay Peninsula (Main Range 



660 



Flora Malesiana 



[ser, I, vol. 6* 




Fig. 49. Rhododendron wrayi K. & G. a. Branchlet with flowers, x 2/3, b. flower, x 1, c. ovary and style, 
X 10/3, d. stamen, X 10/3, e-f. seed, x 10 {a-d Seimund 515, e-/HoLTTUM s.n.). 



from Mt Kerbau on the Perak/Pahang/Selangor 
border to Mt Ulu Kali in Selangor; further on 
Mts Tahan and Tapis in Pahang). 

Ecol. In mossy forest associated with Dacry- 
dium datum, also in dwarf mossy forest or summit 
vegetation, in Leptospermum forest, also in expos- 
ed situations on the actual summits, open ridges, 
quartzite cliffs, (8 5 5-) 1200-2 165 m, locally 
plentiful (so in the Cameron Highlands sometimes 
dominant just below the comb of the ridges). 
Fl. Jan. -Dec. 

Note./?, wrayi is most related to R. pennivenium 
Balf. /. & FoRR. from Yunnan, with which it 
shares the glandular-tomentose ovary and the 
leaves clad underneath with a thin whitish or 
fawn indumentum of interlaced cobweb-like 
hairs, which persists for a long time. R. penni- 
venium diff"ers in the rose- to deep-crimson corolla 
and the leaves which are initially floccose and 
glandular. 

279. Rhododendron korthalsii MiQ. Ann. Mus. 
Hot. Lugd.-Bat. 1 (1863) 43; Millais, Rhodod. 



(1917) 199; Sleum. Blumea Suppl. 4 (1958) 54, 
in clavi; Reinwardtia 5 (1960) 228. 

Shrub? Branchlets thick, terete, glabrous. Leaves 
both in 4-5-merous pseudowhorJs at the nodes 
(c. 6 cm) and laxly scattered on the internodes, 
lanceolate, apex short-acuminate, subacute, with a 
small, not protruding terminal gland, base attenu- 
ate to the petiole, coriaceous, ifc dull on both faces, 
wax-coated above (becoming lustrous there when 
heated), glabrous, rather densely punctulate 
beneath, 8-10 by (1. 5-) 1.8-2.6 cm, midrib im- 
pressed above, prominent beneath, nerves 6-8 
pairs, ± straight below, curved and ± irregular 
distally, faintly anastomosing, slightly impressed 
above, somewhat raised beneath, reticulation dense, 
regular, slightly but well visibly impressed above, 
rather faint beneath; petiole stoutish, grooved 
above, glabrous, 0.7-1.2 cm. Racemes sub- 
umbellate, ± 6-flowered. Rachis elongate-conical, 
c. 2.5 cm, rather densely clad with short-stipitate, 
reddish glands especially in the upper part. 
Perulae oblong, glandular and hispid-hairy, c. 
1 by 0.5 cm. Bracteoles linear-spathulate, hairy. 



Jan. 1966] 



Ericaceae (Sleumer) 



661 



up to 1 cm. Pedicels stoutish, 1.5-2 cm, densely 
set with very shortly stalked glands, but no 
proper hairs. Calyx deeply 5-lobed, lobes ovate, 
rounded-obtuse, c. 2 (rarely up to 3) by c. 2 mm, 
membranous, veined, ciliolate, short-glandular 
(glutinous) at the very base only, glabrous for the 
rest. Corolla wide-campanulate, (white?), c. 2.5 
cm long in all, thin, glabrous except a pubescent 
spot at the base of the tube inside, lobes oblong- 
ovate, 1.5-1.8 by c. 1 cm. Stamens 10, unequal, 
1.2-1.6 cm; filaments linear, glabrous at the very 
base (1 mm), then densely set with papillose 
(inflated) spreading whitish blunt hairs for 4-6 
mm, glabrous in the upper half or to 2/3; anthers 
broad-elliptic, — truncate, 1.5-2 by 1 mm. Disk 
glabrous below, densely whitish ciliate on the 



upper margin. Ovary conical, r. 4 by 2 mm, all 
over with scattered red-brown substrigose hairs 
but apparently no proper glands; style abrupt with 
the obtuse ovary, glabrous, 1.5-1.8 cm; stigma 
short-capitellate. 

Distr. Malesia: Central Sumatra (possibly 
Padang Highlands). One collection. 

Note. R. korthalsii shares the wax-coated sur- 
face of the leaves with R. lepiopeplum BM.r.f. & 
FORR. (Yunnan) and R. lukiangense Franch. 
(SE. Tibet, SW. Szechuan, NW. Yunnan), but 
differs from them in many respects. The most 
related species are probably R. ombrochares 
Balf. /. & Ward (NE. Upper Burma), R. cero- 
chitum Balf. /. & Fork. (W. Yunnan), and R. 
araiophyllum Balf. /. & W. W. Sm. (W. Yunnan) 



3. Subgenus Pentanthera 

(G. Don, pro sect.) Pojarkova, Flora U.R.S.S. 18 (1952) 57; Sleum. Blumea 12 (1964) 345. — 
Azalea (non L. sensu Salisb.) Desv. J. Bot. Appl. 1 (1813) 35, p.p. — Anthodendron Reichb. 
in Moessl. Handb. ed. sec. I (1827) XL, 308. — Azalea subg. Pentanthera (G. Don) K. Koch, 
Dendrol. 2 (1872) 186. — Rhododendron subg. Anthodendron (Reichb.) Endl. exWiLS. &Rehd. 
Monogr. Azaleas (1921) 115. 

Plants entirely elepidote. Leaves deciduous. Flowers (several to many) from a strictly floral 
terminal bud; leafy shoots from lateral buds below, i.e. from the axils of the leaves of the past 
season. 

Distr. N. America, Europe, Asia. 

1. Section Pentanthera 

G. Don, Gen. Syst. 3 (1834) 846; Wils. & Rehd. Monogr. Azaleas (1921) 95, 124; Sleum. 
Blumea 12 (1964) 346. — Rhododendron sect. Anthodendron (Reichb.) Endl. Gen. (1839) 759. 

Stamens 5. Corolla funnel-shaped. 

Distr. As above; in Malesia but one cultivated species. 



280. Rhododendron moUe (Bl.) G. Don, Gen. 
Syst. 3 (1834) 846, non S. & Z. 1846; Hassk. 
Cat. Hort. Bog. (1844) 161; Wilson in Wils. & 
Rehd. Monogr. Azaleas (1921) 95; Rehd. Spec. 
Rhodod. ed. 2 (1947) (A.— Azalea mollis Bl. Cat. 
(1823) 44; Bijdr. (1826) 853; Zoll. Syst. Verz. 
2 (1854) 137. — Azalea sinensis Lodd. Bot. Cab. 
9 (1824) t. 885.—/?. sinense (Lodd.) Sweet, Br. 
Fl. Gard. I. 3 (1829) sub t. 290. 

Sparingly branched shrub, 0.3-1.2 m, with 
rather stout erect branches, villous when young 
and often setose. Leaves deciduous, oblong to 
oblong-lanccolatc, apex obtuse and mucronulate, 
base cuncatc, ciliate and often revolutc at margin, 
soft-pubescent above at least when young, beneath 
with a greyish white dense matted pubescence, 
which becomes glaucous or sometimes confined 
to the veins in later stages, 6-15 by 1.8-5.5 cm; 
petiole pubescent, 2-6 mm. Floral buds ovoid, 
with 9-12 broad-ovate mucronatc or acuminate 



perulae, densely grey-pubescent or velutinous, 
sometimes glabrescent. Inflorescence many-flower- 
ed, flowers expanding before the leaves. Pedicels 
puberulous, with only few or without bristles, 
1.2-2.5 cm. Calyx lobes small, obtuse, pubescent, 
ciliate, with few bristles, 1-2 mm. Corolla wide- 
funnel-shaped, 4-5 cm long, 4.5-6 cm across, 
golden-yellow with a large blotch of greenish 
dots, tomcntulose outside, lobes oval or ovate to 
broad-oblong, longer than the rather wide tube. 
Stamens 5, as long as the corolla; filaments 
hairy below the middle; anthers 2 mm. Disk 
hairy. Ovary densely white-hairy; style glabrous, 
exceeding the stamens. Capsule cylindric-oblong, 
1.8-2.5 cm, finely pubescent and sparingly setose. 

Distr. Native in Eastern and Central China, 
cultivated occasionally in Malesia (Java, intro- 
duced before 1823). 

Note. Azalea mollis Bl. is based on a plant 
cultivated in Java. 



i 



4. Subgenus Tsut.sutsi 

(G. Don) Pojarkova, Flora U.R.S.S. 18 (1952) 55; Sleum. Blumea 12 (1964) 347. — Azalea 
subg. Tsutsutsi (G. Don, pro .'sect.) K. Koch, Dendrol. 2 (1872) 171. — Rhododendron subg. 



662 



Flora Malesiana 



[ser. I, vol. 6* 



Anthodendron (non (Reichb.) Endl. ex WiLS. & Rehd.) Sleum. Blumea Suppl. 4 (1958) 49; 
Reinwardtia 5 (1960) 228. — Fig. 50—51. 

Plants elepidote. Leaves persistent or deciduous in the wild state in Asia, persistent in Malesia 
including the cultivated species introduced from Asia. Flowers (1-3, rarely more), from the 
terminal bud; leafy shoots from the axil of lower perulae of the same bud. 

Distr. E. Asia, in Malesia: Philippines (Luzon). Fig. 46. 

1. Section Tsutsutsi 

Sweet, Br. Fl. Card. I, 5 (1831) sub t. 117 ('Tsutsusr); G. Don, Gen. Syst. 3 (1834) 845, p.p.; 
Planch. Fl. Serres 9 (1854) 76 {'Tsutsia); Miq. Fl. Ind. Bat. 2 (1859) 1059 CTsutsugr); Wils. 
& Rehd. Monogr. Azaleas (1921) 22; Sleum. Blumea Suppl. 4 (1958) 49; ibid. 12 (1964)347.— 
Rhododendron sect. Anthodendron Sleum. Reinwardtia 5 (1960) 228. — Fig. 50 — 51. 

Branchlets with appressed, flattened, mostly brown hairs, or with flattened and normal (not- 
flattened) hairs. Leaves ± scattered on the branchlets, ± strigose. 

Distr. E. Asia, in Malesia: Philippines (Luzon, 1 sp.). Fig. 46. 

KEY TO THE SPECIES 

1. Perulae viscid, especially on the inner surface. 
2. Shoots clothed with appressed, flattened, grey-brown strigose hairs which gradually disappear with 

age. Stamens 10 281. R. scabrum 

2. Shoots densely clothed with spreading, grey to grey-brown soft (whether or not glandular) hairs, 
mixed with few to numerous, loosely appressed or spreading flattened bristle-like hairs. 
3. Stamens normally 10 (sometimes 8). Ovary densely setose, glandless .... 282. R. mucronatum 
3. Stamens normally 5 (6-10 in some garden forms). Ovary glandular-setose. 

283. R. linearifolium var. macrosepalum 
1. Perulae not viscid. 

4. Stamens 5. Leaves crenulate-serrate 284. R. indicum 

4. Stamens (6-)8-10. Leaves entire or nearly so. 

5. Style hairy in its lower 1/4 to 1/3 285. R. subsessQe 

5. Style entirely glabrous 286. R. simsii 



281. Rhododendron scabrum G. Don, Gen. Syst. 
3 (1834) 846; WiLS. in Wils. & Rehd. Monogr. 
Azaleas (1921) 59; Rehd. Spec. Rhodod. ed. 2 
(1947) 102. 

Loosely branched shrub, 1-2 m tall, with rather 
stout branches; shoots clothed in their first year 
with appressed flattened grey-brown strigose hairs, 
glabrescent. Leaves dimorphous, persistent, coria- 
ceous; spring leaves elliptic-lanceolate to lanceo- 
late, apex acute to obtuse and mucronate, base 
broad-cuneate, ciliate and subcrenulate at the 
± revolute margin, 3-9.5 by 1.8-3.5 cm, at 
first with scattered, appressed, straight grey-brown 
hairs, at maturity glabrescent nerves excepted, 
nerves impressed above, raised beneath; summer 
leaves similar, but oblanceolate, smaller and 
more coriaceous; petiole stout, flattened, clothed 
whh appressed grey-brown hairs, 4-8 mm. Floral 
winter buds ovoid, acute, outer perulae clothed 
with strigose rufous hairs, inner ones cinnamon- 
brown, ciliate, viscid especially on the inner 
surface, persisting at the base of the pedicels. 
Inflorescence 2-6-flowered. Pedicels densely ful- 
vous-strigose, 8-12 mm. Ca/yx lobes ample, green, 
normally oval-rounded, c. 6 by 5 mm, occasionally 
all or some lanceolate up to 12 mm long and 
acuminate, always glandular, ciliate and covered 
with appressed straight grey hairs. Corolla broad- 
funnel-shaped, c. 6 cm across and as long, rose- 



red to vivid scarlet or rich magenta, with dark 
dots on the upper 3 lobes, lobed to nearly the 
middle. Stamens 10, shorter than the corolla; 
filaments short-hairy below the middle. Ovary 
glandular and strigose; style glabrous, longer than 
the stamens. Capsule erect, ovoid, 1-1.2 cm long, 
sparsely hairy, subtended by the persistent calyx 
lobes. 

Distr. Native in the Riu Kiu Is., in Malesia 
cultivated occasionally in Java. 

282. Rhododendron mucronatum (Bl.) G. Don, 
Gen. Syst. 3 (1834) 846; Wils. in Wils. & Rehd. 
Monogr. Azaleas (1921) 68; Rehd. Spec. Rhodod. 
ed. 2 (1947) 92; Sleum. Blumea Suppl. 4 (1958) 
49. — Azalea rosmarinifolia Burm./. Fl. Ind. (1768) 
43, t. 3, f. 3; Bl. Bijdr. (1826) 853, non R. rosma- 
rinifolium Vidal, 1886. — Azalea mucronata Bl. 
Cat. (1823) 44; Bijdr. (1826) iST,.— Azalea ledi- 
folia Hook. Bot. Mag. (1829) t. 2901.— /?. 
burmanni G. Don, Gen. Syst. 3 (1834) 846; 
Hassk. Cat. Hort. Bog. (1844) 161; Moritzi, 
Syst. Verz. (1846) 42; Zoll. Syst. Verz. 2 (1854) 
137.—/?. ledifolium (HooK.) G. Don, Gen. Syst. 3 
(1834) 846; Hassk. Cat. Hort. Bog. (1844) 162; 
Miq. Ann. Mus. Bot. Lugd.-Bat. 1 (1863) 34; 
Merr. Philip. J. Sc. 19 (1921) 369; Amsh. in 
Back. Bekn. Fl. Java (em. ed.) 7 (1948) fam. 
162, p. 2; Steen. Fl. Sch. Indon. (1949) 309. 



Jan. 1966] 



Ericaceae (Sleumer) 



663 



i 



Much branched, wide-spreading shrub, 0.9-1.8 
(-2.5-5) m; shoots in their first year, as well as 
petioles and pedicels densely clothed with spread- 
ing grey to grey-brown soft hairs, mixed with few 
to many loosely appressed or spreading, flattened, 
bristle-like hairs, and occasionally with glandular 
hairs. Leaves partly persistent, dimorphous, spring 
leaves deciduous, membranous, lanceolate to 
ovate-lanceolate, apex acute or obtuse and mucro- 
nulate, base cuneate, on both surfaces with appress- 
ed straight and soft grey to rufous hairs, 3-4.5 
by 1-2.5 cm; summer leaves persistent, chartaceous, 
oblong-lanceolate to oblong-obianceolate, apex 
obtuse to rounded and mucronate, base cuneate, 
recurved at margin, pubescent like the spring 
leaves and often glandular, 1-3.5 by 0.6-1.2 cm; 
petiole 2-6 mm. Floral winter buds ovoid, 
outer perulae with strigose hairs along the middle 
line, inner ones viscid. Inflorescence 1-3-flowered. 
Pedicels 5-20 mm. Calyx lobes lanceolate, often 
slightly erose or serrate, glandular-pubescent, 
c. 12 mm. Corolla wide-funnel-shaped, 3.5-5 cm 
long and wide, fragrant, pure white or white with 
a yellowish blotch on the upper corolla lobes in 
the cultivated forms (red or rose coloured in the 
native form), occasionally a few flowers rose or 
striped with rose on an odd branch also in cultivat- 
ed plants, with oval or narrow-oval lobes. Stamens 
10 (sometimes 8), ± as long as the corolla; 
filaments glabrous or subglabrous. Ovary densely 
setose, glandless; style glabrous, slightly exceed- 
ing the corolla. Capsule conic-ovoid, with appress- 
ed setose hairs, c. 1 cm long, shorter than the 
persistent calyx lobes. 

Distr. A native of Japan (Shikoku) in its form 
with rose to purple flowers (R. ripense Making), 
the white flowered form much cultivated in Japan 
and China, early introduced to Java and still 
cultivated there in various forms, up to 1400 m. 
Fl. Jan. -Dec. 

283. Rhododendron linearifolium S. & Z. var. 
macrosepalum (Maxi.m.) Making, Bot. Mag. 
Tokyo 27 (1913) 108; Wils. in Wils. & Rehd. 
Monogr. Azaleas (1921 ) 74; Rehd. Spec. Rhodod. 
ed. 2 (1947) 87. — R. macrosepalum Maxim. 
Gartenflora 19 (1870) 258, t. 662; M6m Ac. Imp. 
Sc. St. P^tersb. VII, 16, n. 9 (1870) 31. 

Loosely branched and usually low shrub, 
0.3-1 (-3) m; shoots and pedicels densely clothed 
with greyish, partly gland-tipped hairs, and a few 
spreading flattened caducous hairs. Leaves mostly 
deciduous, dimorphous, spring leaves membra- 
nous, partly scattered and partly clustered, 
ovate-elliptic or ovate to lanceolate, apex acute 
or short-acuminate, mucronate, base cuneate, 
sparingly glandular-hairy above, slightly rugulose 
and hispid, beneath with soft spreading grey to 
grey-brown glandular hairs, 2.5-6 by 0.6-2.5 cm; 
summer leaves oblanccolatc to oblong-oblancco- 
latc, obtuse and mucronulatc, of firmer texture; 
petiole densely hairy ,2 6 mm. Floral winter buds 
ovoid; outer perulae r. 10, densely covered with 
strigose yellow-brown hairs, inner ones viscid. 
Inflorescence 2-IO-flowcrcd. Pedicels l.2-3(-4) 



cm. Calyx lobes lanceolate, 1.2-3 cm, sometimes 
longer than the corolla, glandular-pubescent. 
Corolla broad-funnel-shaped, 3.6-5 cm across and 
as long, rose-pink to red-purple, fragrant, with 
dark purple dots on the upper lobe, lobes oval to 
broad-oblong. Stamens normally 5 (6-10 in some 
garden forms), slightly shorter than the corolla; 
filaments pubescent below the middle. Ovary 
glandular-setose; style glabrous, longer or as long 
as the corolla. Capsule narrow-ovoid, 8-12 mm, 
glandular-pubescent, subtended by the persistent 
reflexed calyx lobes. 

Distr. Native of Japan, occasionally cultivated 
in Java at c. 1000 m. 

284. Rhododendron indicum (L.) Sweet, Br. Fl. 
Gard. II, 2 (1833) sub t. 128; Hassk. Cat. Hort. 
Bog. (1844) 162, incl. var. A-E, I.e.; Zoll. Syst. 
Verz. 2 (1854) 137; MiQ. Fl. Ind. Bat. 2 (1859) 
1059; Wils. in Wils. & Rehd. Monogr. Azaleas 
(1921) 22; Kgord. Exk. Fl. Java 3 (1923) 7; 
Rehd. Spec. Rhodod. ed. 2 (1947) M.— Azalea 
indica Linne, Sp. PI. (1753) 150; Bl. Cat. (1823) 
44; Bijdr. (1826) 852, incl. var. spathulata Bl. 
I.e., var. angustifolia Bl. I.e. 853 et var. floribunda 
Bl. I.e. 853. — R. breynii Planch. Rev. Hort. 
sdr. 4, 3 (1854) 46; Fl. Serres 9 (1854) 77 (based 
on a plant cultivated in Java). 

Densely branched shrub, up to 2 m, usually low, 
sometimes prostrate. Branches slender, but rigid, 
with flattened, appressed, chestnut-brown hairs. 
Leaves crowded, persistent, narrow-lanceolate to 
oblanceolate, acute at the ends, with scattered, 
closely appressed red-brown hairs on both faces, 
usually remotely crenulate-serrulate and ciliolate, 
1.8-3.6 by 0.3-1 cm; petiole densely clothed with 
appressed chestnut-brown hairs, 1-4 mm. Floral 
winter buds ovoid, pointed, not viscid; outer 
perulae ciliate but otherwise glabrous or with 
a tuft of brown hairs at the apex. Inflorescence 
1-2-flowered. Pedicels with strigose brown hairs, 
hidden by the perulae, 3-5 mm. Calyx lobes oval 
or suborbicular, obtuse, densely strigose and 
ciliate, 2-3 mm. Corolla broad-funnel-shaped, 
5-6 cm long and wide, bright red to scarlet, 
sometimes rose-red or salmon, with oval rounded 
lobes. Stamens 5, 4 as long as the corolla; 
filaments glabrous. Ovary densely clothed with 
appressed shining brown hairs; style glabrous, 
exceeding the stamens. Capsule oblong-ovoid, c. 
6 mm, densely strigose. 

Distr. Native of S. Japan, cultivated occasion- 
ally in Java, mostly in hybrids. The Azalea indica 
of the trade is not this species, but R. simsii 
(and related forms). 

Note. R. indicum (L.) Sweet/, variegatum (Bl.) 
DC. Prod. 7 (1839) 726; Wils. in Wils. & Rehd. 
Monogr. Azaleas (1921) 26. — Azalea indica L. 
var. variegala Hi . Bijdr. (1826) 852, is a form with 
whitc-and-rcd-stripcd and spotted flowers, culti- 
vated in Java in Blume's time. 

285. Rhododendron subst-sslle Rfndi.f, J. Bot. 
34 (1896) 357; Mirk. Publ. Gov. Lab. Philip, n. 
29 (1905) 40; Philip. J. Sc. 3 (1908) Bot. 379; 



664 



Flora Malesiana 



[ser. I, vol. 6* 




d 



Fig. 50. Rhododendron subsessile Rendle. a. Habit, X 2/3, b. flower, X 1, c. stamen, X 8/3, d. ovary 
and style, x 8/3, e. fruit, x 2,/. seed, x 10 (a-t/ Barness 922, e-/ Williams 2001). 



ibid. 5 (1910) Bot. 371; Millais, Rhodod. (1917) 
249; Wils. in Wils. & Rehd. Monogr. Azaleas 
(1921) 51; Merr. En. Philip. 3 (1923) 245; 
Millais, Rhodod. ser. 2 (1924) 244; Copel. /. 
Philip. J. Sc. 40 (1929) 174, t. 16, f. 7-10, incl. 
var. baucoense Copel. /. I.e. 175; Rehd. Spec. 
Rhodod. ed. 2 (1947) 106; Sleum. Reinwardtia 5 
(1960) 228; Blumea 12 (1963) 1 14; Jahrb. Rhodod. 
Ges. Bremen (1963) f. 2. — .'' R. rubropilosum 
Hayata, J. Coll. Sc. Imp. Un. Tokyo 30, art. 1 
(1911) 173; Wils. in Wils. & Rehd. Monogr. 
Azaleas (1921) 52; J. Arn. Arb. 6 (1925) 182; 
Kaneh. Formos. Trees (1936) 547, f. 507; Rehd. 
Spec. Rhodod. ed. 2 (1947) 99.— ?R. caryophyllum 
Hayata, Ic. PI. Formos. 3 (1913) 130, t. 23.— 
Azalea subsessilis (Rendle) Copel. /. Am. Midi. 
Nat. 30 (1943) 597, in text.— Fig. 50-51. 

Much-branched shrub, l-3(-6) m; shoots dense- 
ly clothed with appressed, upwardly directed, 
flattened, bristle-like or strigose chestnut-brown 
hairs, which in the third season are wearing ofi". 
Leaves ascending, persistent, scattered along the 
upper shoots and ± crowded towards their ends, 
elliptic to lanceolate, short-acuminate, acute and 
prominently mucronate, base ± broad-attenuate, 
darker above, firmly chartaceous to subcoriaceous, 
at first hairy on both faces, at maturity laxly 



covered with whitish hairs above, underneath 
densely clad with rufous, later greyish silky long 
bristle-like or strigose, appressed, ± reddish hairs 
and thickened at the base, margin slightly to 
strongly recurved, (l-)2^(-5) by (0.3-)0.5-1.2 
(-1.4-1.8) cm, midrib impressed above, prominent 
beneath, nerves rather indistinct; petiole strigose- 
hairy, 3-6(-8) mm. Floral buds ovoid; perulae 
broad-ovate, abruptly attenuate or mucronate at 
the apex, up to 9 by 6 mm, outer ones with appress- 
ed brown hairs along the midline, ciliate, other- 
wise glabrous, inner ones appressed strigose toward 
the apex, not viscid, long persistent. Inflorescence 
(l-)2-4(6-)-flowered. Pedicels stoutish, with 
appressed rusty hairs, 2-4 mm. Calyx lobes 
broadly oblong or ovate to suborbicular, obtuse, 
2-3 mm, densely clad with upright flattened 
rufescent hairs outside, glabrous inside. Corolla 
mostly light red or lilac to purple-red (magenta), 
sometimes pink, rarely white, apparently never 
spotted in the Philippine specimens, campanulate- 
funnel-shaped, gradually dilated from the base, 
glabrous, 1.8-2.2 cm long, c. 2.5 mm 0, tube 
8-9 mm, lobes 1.2-1.4 by 8-9 mm, ovate-oblong, 
shortly spreading. Stamens (6-)8(-10), slightly 
longer than the corolla; filaments linear, ± 
sparingly whitish papillose hairy towards the base. 



Jan. 1966] 



Ericaceae (Sleumer) 



665 




Fig. 51. Rhododendron subsessile Rendle. Mt 

Data, 2000 m, Luzon, Philippines (van Royen & 

Sleumer 5613) (Sleumer, 1961). 

Ovary subovoid, c. 3 mm, densely covered with 
flattened rufous or rusty, shiny, upwards directed 
long hairs; style c. 2 cm, exserted, clad with the 
same hairs as occur on the ovary in its lower |4 
to Yi, glabrous otherwise; stigma small, capitate. 
Capsule conic-ovoid, densely clothed with appress- 
ed, rufous hairs, 6-8(-10) by 4-5 mm; pedicel to 
9 mm. Seeds very shortly or not winged, 1 mm. 

Distr. Formosa, in Malesia: Philippines (N. 
Luzon: Mountain and Abra Prov.). 

Ecol. In Luzon rather common in the pine 



region and the pine-oak association below the 
mossy forest, rare in the mossy forest (Mt Pulog), 
again on exposed grass-covered slopes above the 
moss forest, often in clumps or masses, 1 300-2400 
m (in Formosa only on open slopes, 2135-3300 
m). Fl. fr. Jan. -Dec. 

Vern. Ausip, ayusip, buga-buga, bugo-gugua, 
salangon, Ig. 

Note. This is the most southern species of the 
"Azalea group. Its var. baucoense Copel. /. is a 
microform, connected by intergrades with the 
larger typical form. As to Formosan R. rubropilo- 
sum Hayata see Sleum. Reinwardtia 5 (1960) 229. 

286. Rhododendron simsii Planch. Fl. Serres 9 
(1854) 78; Wils. in Wils. & Rehd. Monogr. 
Azaleas (1921) 45; Rehd. Spec. Rhodod. ed. 2 
(1947) 105; Sleum. Blumea Suppl. 4 (1958) 51.— 
Azalea indica {non L.) Sims, Bot. Mag. (1812) t. 
1480. 

A much-branched shrub, usually up to 1.5 m, 
rarely attaining 2.5 m; shoots densely clothed with 
appressed, flattened, shining brown hairs. Leaves 
persistent or partly so, and dimorphous in the 
wild plant, always persistent in specimens cultivat- 
ed in the tropics and less distinctly dimorphous 
there; spring-leaves (or early leaves) chartaceous, 
elliptic, sometimes ovate or oblong-elliptic, 
1.8-5 by 0.8-1.8 cm, apex acute or somewhat 
obtuse, base ± cuneate, sparingly strigose above, 
more densely so beneath at least on midrib and 
veins; summer-leaves (or later leaves) obovate to 
oblanceolate, 0.8-3.6 by 0.4-1.2 cm, more 
coriaceous; petiole densely strigose, 2-6 mm. 
Floral winter buds ovoid, not viscid, c. 4 cm 
long or less, with several perulae, these densely 
strigose along the median and ciliate. Inflores- 
cences 2-6-flowered. Pedicels densely strigose, 
5-10 mm. Ca/vAT lobes ovate to lanceolate, strigose 
and ciliate, 2-6 mm persistent. Corolla broad- 
funnel-shaped, 3.5-5 cm long and wide, rose-red 
to bright or dark red, spotted, lobes broad-ovate. 
Stamens (8-9-) 10, i as long as the corolla; 
filaments pubescent below the middle; anthers 
purple. Ovary strigose; style exceeding the corolla, 
glabrous. Capsule ovoid, c. 8 mm long, setose. 

Distr. Native in the temperate parts of Thai- 
land, Indochina, China, and S. Formosa, in 
Malesia occasionally cultivated under the commer- 
cial name 'Indian Azalea'. 



5. Subgenus Azaleastrum 

Planch. Fl. Serres 9 (1854) 75 Csous-type); Rev. Hort. s6r. 4, 3 (1854) 43 {'so us- type); ex 
K. Koch, Dendrol. 2 (1872) 159. descr. subf;.; Drude in E. & P. Pn. Fam. 4, 1 (1889) 37; 
Sleum. Bot. Jahrb. 74 (1949) 528, 552; Blumea Suppl. 4 (1958) 56; Reinwardtia 5 (1960) 230. — 
Azaleastrum (Planch.) Rydh. Mem. N.Y. Bot. Gard. 1 (1900) 297. 

Floral buds, respectively inflorescences, though seemingly terminal, in reality lateral, mcstly 
several together, starting from the axil(s) of the uppermost leaf (or leaves). 

Distr. N. America, E. Asia, Malesia (Malay Peninsula). 

I. Section (honia-strum 
Franch. Bull. Soc. Bot. I r. 33 (1886) 229; Sllum. Bot. Jahrb. 74 (1949) 528, 552; Blumea 
Suppl. 4 (1958) 56; Reinwardtia 5 (1960) 230. — Fig. 53. 



\ 



666 



Flora Malesiana 



[ser. I, vol. 6* 



Calyx small or obscure. Stamens 10. Seeds shortly and irregularly winged at both ends. 
Distr. E. Asia, Malesia (Malay Peninsula). Fig. 52. 




Fig. 52. Distribution of Rhododendron sect. Choniastrum: southern limit of the section; 

southern limit and • localities of Rhododendron moulmainense Hook, respectively; + localities of 

Rhododendron klossii Ridl. 



KEU TO THE SPECIES 

1. Ovary glabrous 287. R. moulmainense 

1. Ovary laxly to subdensely hairy 288. R. klossii 



287. Rhododendron moulmainense Hook. Bot. 
Mag. (1856) t. 4904; Kurz, J. As. Soc. Beng. 
46, ii (1877) 216; Fl. Burma 2 (1877) 94; Clarke 
in Hook./. Fl. Br. Ind. 3 (1882) 463; Brandis, 
Ind. Tr. (1906) 412; Craib, Kew Bull. (1911) 405; 
Ridl. Fl. Mai. Pen. 2 (1923) 218; Dop, Fl. I.-C. 3 
(1930) 734; Hutch. Spec. Rhodod. (1930) 620, 
excl. syn. R. klossii; in Craib, Fl. Siam. En. 2 
(1938) 318; Sleum. Blumea Suppl. 4 (1958) 58; 
Reinwardtia 5 (1960) 230.—^. westlandii Hemsl. 
J. Linn. Soc. Bot. 26 (1889) 31; Millais, Rhodod. 
(1917) 261; Hutch. Spec. Rhodod. (1930) 626.— 
R. oxyphyllum Franch. J. de Bot. 12 (1898) 264; 
Millais, Rhodod. (1917) 222; Hutch. Spec. 
Rhodod. (1930) 621. — Azalea moulmainensis 
(Hook.) O. Ktze, Rev. Gen. PI. 2 (1891) 387.— 
R. siamense Diels in Fedde, Rep. 4 (1907) 289. — 
R. leucobotrys Ridl. J. Fed. Mai. St. Mus. 4 
(1909) 43; ibid. 7 (1916) 46; Fl. Mai. Pen. 2 
(1923) 218; Hutch. Spec. Rhodod. (1930) 619.— 
R. stenaulum Balf. /. & Forr. Not. R. Bot. 
Card. Edinb. 10 (1917) 157; Hutch. Spec. 
Rhodod. (1930) 624, excl. syn. R. mackenzianum 
Forr. — Fig. 53. 

Terrestrial shrub, 3-5 m, or small tree, with 
straight, erect branches. Branchlets glabrous, 
terete; internodes 2-8 cm. Leaves in ± terminal, 
2-4-merous pseudowhorls, elliptic or mostly el- 
liptic-lanceolate, rarely obovate-elliptic or oblong- 
oblanceolate, apex acute, base narrowed, ± 
coriaceous, glabrous, 6-12(-15) by (2.5-)3-5(-6) 



cm, midrib impressed above, much raised beneath, 
nerves 9-12 pairs, straight, spreading and ± 
parallel, initially slightly prominent or somewhat 
impressed, more conspicuously inarched in mature 
leaves above, prominent beneath, veins slightly 
or not visible; petiole often red in fresh specimens, 
somewhat winged, (6-)8-12(-15) mm. Inflores- 
cences 2-4, each in the axil of one of the upper 
leaves; inflorescences 2-4-flowered. Floral buds 
ovoid-oblong, subacute, c. 2.5 by 0.8 cm. Outer 
perulae small and ovate, obtuse, inner ones 
gradually larger and ovate to ovate-oblong or 
obovate-oblong, all glabrous or rather faintly 
sericeous outside especially towards the tip, 
densely white-ciliate. Bracteoles linear, glabrous, 
up to 2.5 by 0.1 cm. Pedicels rather slender, 
glabrous, (l-)1.5-2.5 cm. Calyx very small, 
margin undulately lobed, glabrous. Corolla 
funnel-shaped, deeply 5-lobed, 3.5-4 cm long in 
all, tube 1-1.5 cm, base 5-saccate, 3-4 mm 0, 
lobes oblong, subacute, ± spreading, 2-2.5 by 
0.5-0.9 cm, glabrous, mostly white with a yellow 
spot inside on the upper lobe, also rose or of a clear 
orchid-purple with a ± orange spot, rarely 
reddish, fragrant. Stamens normally 10 (some- 
times a few more), ± as long as the corolla; 
filaments linear, hairy except the very base and 
upper half, occasionally entirely glabrous; anthers 
short-oblong, 2.5 by 1 mm. Disk glabrous. 
Ovary glabrous, cylindric, slightly 5-ribbed, mostly 
continuous (rarely abrupt) with the glabrous 



Jan. 1966] 



Ericaceae (Sleumer) 



667 




Fig. 53. Rhododendron moulmainense Hook. a. Habit, X ^3, b. ovary, X 2, c. stamen, x 2 {a-c Poilane 

28769). 



style, c. 3 cm long; stigma capitate. Capsule 
ovoid-fusiform, rather deeply 5-grooved, 2-2.5 
by c. 0.6 cm, beaked by the persistent rest of the 
style even after the capsule is split into 5 rather 
thin, not twisted valves. Seeds flat, broad-oblong, 
very shortly and irregularly winged (not tailed) 
on both ends, c. 2 mm. 

Distr. Burma, S. China, Indochina, Thailand, 
in Malesia: Malay Peninsula (Kedah Peak: Mt 
Jerai), 855-1220 m. /•/. May-Dec, fr. May. 



2S8. Rhododendron klossii Ri[>i.. J. Fed. Mai. St. 
Mus. 4 (\W)) 43; M. Mai. Pen. 2 (1923) 218; 



\ 



Sleum. Blumca Suppl. 4 (1958) 59; Rcinwardtia 
5 (1960) 231.—/?. laoticum Dop, Fl. I.-C. 3 
(1930) 735. 

Shrub or trcclet, occasionally a tall tree up to 
15 m high. Branchlets glabrous, terete; inlcrnodes 
2.5-6 cm. Leaves mostly lanceolate to oblong- 
lanceolate, rarely lanceolate-clliplic, (6-)8-13 by 
(2-)2.5-6 cm, very similar to those of R. moul- 
mainense. Injiorescences 3 6, seemingly terminal, 
each with 2 flowers, making an umbel of 6-12 
flowers. Flower buds ovoid-oblong, subacute, 
2-2.5 by f. 0.6 cm. Perulac and bracteolcs as in R. 
moulmainense. Pedicels rather slender, glabrous 



668 



Flora Malesiana 



[ser. I, vol. 6^ 



or initially very finely pubescent, c. 2.5 cm. 
Calyx small, obscurely 5-dentate, an occasional 
lobe accrescent to 3 mm. Corolla funnel-shaped, 
while, fragrant, c. 3J cm long in all, tube cylin- 
dric, c. 1.2-1.5 cm by 3-4 mm, lobes oblong, 
obtuse, 2-2.5 by 0.5-0.8 cm. Stamens ± as long 
as the corolla; filaments slender, pubescent in the 
middle, glabrous at the base and in the upper 
half; anthers oblong, 2.5 by I mm. Disk glabrous. 
Ovary cylindric, 5-ribbed, laxly or mostly sub- 
densely clothed with simple subapprcsscd hairs; 
style continuous with the ovary, slender, glabrous, 



somewhat exserted. Capsule apparently similar to 
that of R. moulmainense. 

Distr. Indochina (Central Laos and SW. 
Cambodge), in Malesia: Malay Peninsula (Main 
Range on the Perak-Pahang border, from the 
Cameron Highlands to the Frazer Hill region and 
in SW. Pahang). Fig. 52. 

Ecol. Rocky ridges, 1600-1800 m. Fl. April, 
June, Nov., fr. June. 

Note. Very close to R. moulmainense, different 
from it by the hairy ovary. 



i!: Ill III III 

3 5185 00202 4964 




FLORA MALESIANA 

SCIENTIFIC COMMUNICATIONS 

concerning Flora Malesiana should be addressed to the General Editor, 
Dr C. G. G. J. VAN Steenis, Rijksherbarium, Leyden, Holland 



SUBSCRIPTIONS 

should exclusively be directed to N.V. de Erven P. Noordhoff, 
P.O. Box 39, Groningen, Holland 

Volume 1. Cyclopaedia of collectors & collections. 1950. pp. clii. + 639. Dfl. 55.50, £ 5,10.-, US$ 14.65. 

Volume 2. Malaysian vegetation In preparation. 

Volume 3. Malaysian plant geography In preparation. 

Volume 4. General chapters and revisions. 1948-1954. pp. ccix + 631. Dfl. 69.-, ;^6.16.-, US$ 18.15. 
Volume 5. Bibliography, specific delimitation, & revisions. 1955-1958. pp. cccxlii -|- 596. Dfl. 92.-, 

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For sale only , Not for exchange 
INDEX TO REVISED FAMILIES 



Aceraceae . . 


4: 3, 


592 


Droseraceae 


. . 4: 


377 


Pentaphylacaceae* 


. 5 


121 


Actinidiaceae s.str 


. 4: 


37 


Elatinaceae 


. . 4: 


203 


Philydraceae . 


. 4 


5 


Aizoaceae . 


. . 4: 


267 


Epacridaceae 


. . 6: 


422 


Phytolaccaceae 


. 4 


229 


Alismataceae . 


. . 5: 


317 


Ericaceae . 


. . 6: 


469 


,Pittosporaceae. 


. 5 


345 


Amaranthaceae 


. 4: 69, 


593 


Erythroxylacea 


e . . 5: 


543-' 


Plumbaginaceae 


. 4 


107 


Ancistrocladaceae 


. 4: 


8 


Ficoidaceae 


. . 4: 


267 


Podostemaceae 


. 4 


65 


Aponogetonaceae 


. . 4: 


11 


Flacourtiaceae 


. . 5: 


1 


Polemoniaceae 


. 4 


195 


Basellaceae 


. . 5: 


300 


Flagellariaceae 


. . 4: 


245 


Pontederiaceae 


. 4 


255 


Batidaceae . 


. . 5: 


414 


Geraniaceae 


. . 6: 


445 


Primulaceae . 


. 6 


173 


Betulaceae . 


. . 5: 


207 


Gnetaceae . 


. . 4: 


336 


Proteaceae . 


. 5 


: 147 


Bixaceae s.str. 


. . 4: 


239 


Gonystylaceae 


. . 4: 


349 


Punicaceae . 


. 4 


226 


Burmanniaceae 


. 4: 13, 


592 


Goodeniaceae 


. . 5: 


335 


Restionaceae . 


. 5 


416 


Burseraceae 


. . 5: 


209 


Haemodoracea 


e . . 5: 


111 


Rhizophoraceae 


. 5 


429 


Butomaceae . 


. . 5: 


118 


Hamamelidace 


le . .5: 


363 


Salicaceae . 


. 5 


• 107 


Callitrichaceae . 


. 4: 


251 


Hippocrateacea 


e . . 6: 


389 


Salvadoraceae . 


. 4 


225 


Campanulaceae 


. 6: 


107 


Hydrocaryacea 


e . . 4: 


43 


Sarcospermaceae 


. 4 


32 


Cannabinaceae 


. 4: 


223 


Hydrocharitace 


ae . . 5: 


381 


Saururaceae . 


. 4 


47 


Capparidaceae 


. 6: 


61 


Hydrophyllacej 


le . .4: 


207 


Scyphostegiaceae 


. 5 


297 


Caprifoliaceae . 


. 4: 175, 


598 


Juglandaceae 


. . 6: 


143 


Simaroubaceae 


. 6 


. 193 


Celastraceae . 


. 6: 227, 


389 


Juncaceae . 


. . 4: 


210 


Sonneratiaceae 


. 4: 280,513 


Centrolepidaceae 


. . 5: 


421 


Juncaginaceae 


. . 4: 


57 


Sparganiaceae . 


. 4: 233 


Ceratophyllaceae 


. 4: 


41 


Loganiaceae 


. . . 6: 


293 


Sphenocleaceae 


. 4: 27 


Chenopodiaceae 


. 4: 99, 


594 


Malpighiaceae 


. . 5: 


125 


Staclchousiaceae 


. 4: 35 


Cochlospermaceae 


. 4: 


61 


Martyniaceae 


. . 4: 


216 


Staphyleaceae . 


. 6: 49 


Combretaceae . 


. . 4: 


533 


Molluginaceae 


. . 4: 


267 


Stylidiaceae 


. 4: 529 


Connaraceae . 


. . 5: 


495 


Moringaceae 


. . 4: 


45 


Styracaceae 


. 4: 49 


Convolvulaceae 


4: 388, 


599 


Myoporaceae 


. . 4: 


265 


Thymelaeaceae 


4: 349, 6: 1 


Corynocarpaceae 


. 4: 


262 


Myricaceae 


. . 4: 


277 


Trigoniaceae . 


. 4: 59 


Crassulaceae . 


. . 4: 


197 


Najadaceae 


. . . 6: 


157 


Tumeraceae 


. 4: 235 


Datiscaceae 


. 4: 


382 


Nyctaginaceae . 


. . 6: 


450 


Typhaceae . 


. 4: 243 


Dichapetalaceae 


. . 5: 


305 


Nyssaceae . 


. . 4: 


29 


Umbelliferae . 


4: 113,595 


Dilleniaceae . 


. . 4: 


141 


Papaveraceae 


. . . 5: 


114 


Valerianaceae . 


. 4: 253 


Dioscoreaceae . 


. . 4: 


293 


Pedaliaceae 


. . 4: 


216 


Xyridaceae 


4: 366, 598 


Dipsacaceae . 


. . 4: 


290 


Pentaphragmat 


aceae . 4: 


517 


Zygophyllaceae 


. 4 


64 



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